longtext: 1CUC-pdb

content
HEADER    HYDROLASE (SERINE ESTERASE)             16-NOV-95   1CUC
TITLE     CUTINASE, N172K, R196D MUTANT, ORTHORHOMBIC CRYSTAL FORM
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: CUTINASE;
COMPND   3 CHAIN: NULL;
COMPND   4 EC: 3.1.1.3;
COMPND   5 ENGINEERED: YES;
COMPND   6 MUTATION: N172K, R196E;
COMPND   7 OTHER_DETAILS: FORM II CRYSTAL (ORTHORHOMBIC)
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: FUSARIUM SOLANI PISI;
SOURCE   3 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   4 EXPRESSION_SYSTEM_PLASMID: PUC19
KEYWDS    HYDROLASE, SERINE ESTERASE, GLYCOPROTEIN
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.LONGHI,C.CAMBILLAU
REVDAT   1   11-JUL-96 1CUC    0
JRNL        AUTH   S.LONGHI,A.NICOLAS,L.CREVELD,M.EGMOND,C.T.VERRIPS,
JRNL        AUTH 2 J.DE VLIEG,C.MARTINEZ,C.CAMBILLAU
JRNL        TITL   DYNAMICS OF FUSARIUM SOLANI CUTINASE INVESTIGATED
JRNL        TITL 2 THROUGH STRUCTURAL COMPARISON AMONG DIFFERENT
JRNL        TITL 3 CRYSTAL FORMS OF ITS VARIANTS
JRNL        REF    TO BE PUBLISHED   REF NOW ASSIGNED AS
JRNL        REFN                                                  0353
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   A.NICOLAS,M.EGMOND,C.T.VERRIPS,J.DE VLIEG,S.LONGHI,
REMARK   1  AUTH 2 C.CAMBILLAU,C.MARTINEZ
REMARK   1  TITL   CONTRIBUTION OF CUTINASE SER 42 SIDE-CHAIN TO THE
REMARK   1  TITL 2 STABILIZATION OF THE OXYANION TRANSITION STATE
REMARK   1  REF    TO BE PUBLISHED   REF NOW ASSIGNED AS
REMARK   1  REFN                                                  0353
REMARK   1 REFERENCE 2
REMARK   1  AUTH   C.MARTINEZ,A.NICOLAS,H.VAN TILBEURGH,M.P.EGLOFF,
REMARK   1  AUTH 2 C.CUDREY,R.VERGER,C.CAMBILLAU
REMARK   1  TITL   CUTINASE, A LIPOLYTIC ENZYME WITH A PREFORMED
REMARK   1  TITL 2 OXYANION HOLE
REMARK   1  REF    BIOCHEMISTRY                  V.  33    83 1994
REMARK   1  REFN   ASTM BICHAW  US ISSN 0006-2960                 0033
REMARK   1 REFERENCE 3
REMARK   1  AUTH   C.MARTINEZ,P.DE GEUS,P.STANSSENS,M.LAUWEREYS,
REMARK   1  AUTH 2 C.CAMBILLAU
REMARK   1  TITL   ENGINEERING CYSTEINE MUTANTS TO OBTAIN
REMARK   1  TITL 2 CRYSTALLOGRAPHIC PHASES WITH A CUTINASE FROM
REMARK   1  TITL 3 FUSARIUM SOLANI PISI
REMARK   1  REF    PROTEIN ENG.                  V.   6   157 1993
REMARK   1  REFN   ASTM PRENE9  UK ISSN 0269-2139                 0859
REMARK   1 REFERENCE 4
REMARK   1  AUTH   C.MARTINEZ,P.DE GEUS,M.LAUWEREYS,G.MATTHYSSENS,
REMARK   1  AUTH 2 C.CAMBILLAU
REMARK   1  TITL   FUSARIUM SOLANI CUTINASE IS A LIPOLYTIC ENZYME
REMARK   1  TITL 2 WITH A CATALYTIC SERINE ACCESSIBLE TO SOLVENT
REMARK   1  REF    NATURE                        V. 356   615 1992
REMARK   1  REFN   ASTM NATUAS  UK ISSN 0028-0836                 0006
REMARK   2
REMARK   2 RESOLUTION. 1.75 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : X-PLOR 3.1
REMARK   3   AUTHORS     : BRUNGER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.75
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 6.0
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 1.
REMARK   3   COMPLETENESS FOR RANGE        (%) : NULL
REMARK   3   NUMBER OF REFLECTIONS             : 15768
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL
REMARK   3   R VALUE            (WORKING SET) : 0.157
REMARK   3   FREE R VALUE                     : NULL
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL
REMARK   3   COMPLETENESS IN THIS BIN        (%) : NULL
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : NULL
REMARK   3   BIN FREE R VALUE                    : NULL
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 1774
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 0
REMARK   3   SOLVENT ATOMS            : 561
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 33.9
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT     (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.008
REMARK   3   BOND ANGLES            (DEGREES) : 1.52
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  NCS MODEL         : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  0  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  0  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  PARAMETER FILE  2  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3  TOPOLOGY FILE  2   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 1CUC COMPLIES WITH FORMAT V. 2.0, 16-FEB-1996
REMARK   6
REMARK   6 SER 120:
REMARK   6 THE "EPSILON" CONFORMATION OF THE CATALYTIC SERINE IS A
REMARK   6 TYPICAL FEATURE OF THE ALPHA/BETA HYDROLASE FAMILY, WHICH
REMARK   6 CUTINASE BELONGS TO.
REMARK   7
REMARK   7 THE FIRST 16 N-TERMINAL RESIDUES AND LAST C-TERMINAL
REMARK   7 RESIDUE ARE NOT OBSERVED IN THE ELECTRON DENSITY.  THEY
REMARK   7 ARE NOT INCLUDED IN THIS ENTRY.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 293
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : NULL
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU RU200 ROTATING
REMARK 200                                   ANODE
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418
REMARK 200  MONOCHROMATOR                  : GRAPHITE CRYSTAL
REMARK 200  OPTICS                         : COLLIMATOR
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE (18CM)
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : MAR XDS
REMARK 200  DATA SCALING SOFTWARE          : NULL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 15984
REMARK 200  RESOLUTION RANGE HIGH      (A) : NULL
REMARK 200  RESOLUTION RANGE LOW       (A) : NULL
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 90.0
REMARK 200  DATA REDUNDANCY                : 8.5 %
REMARK 200  R MERGE                    (I) : 0.060
REMARK 200  R SYM                      (I) : NULL
REMARK 200  <I/SIGMA(I)> FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200  <I/SIGMA(I)> FOR SHELL         : NULL
REMARK 200
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 36.
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS:
REMARK 280   CRYSTAL FORMS OF THIS MUTANT.  THE THREE FORMS ARE
REMARK 280   REFERRED TO AS N172K,R196E_I, N172K,R196E_II,
REMARK 280   N172K,R196E_III BY THE AUTHORS.
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   1/2-X,-Y,1/2+Z
REMARK 290       3555   -X,1/2+Y,1/2-Z
REMARK 290       4555   1/2+X,1/2-Y,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       18.50002
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       35.69899
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       32.10067
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       35.69899
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       18.50002
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       32.10067
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  SOME OF THESE MAY BE ATOMS
REMARK 500 LOCATED ON SPECIAL POSITIONS IN THE CELL.
REMARK 500
REMARK 500 DISTANCE CUTOFF: 2.2 ANGSTROMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500  1HD2  ASN      27     H    VAL     169     1455     1.27
REMARK 500   H    VAL     169    1HD2  ASN      27     1655     1.27
REMARK 500  2HH2  ARG      96    1H    HOH     608     3655     1.58
REMARK 500  1H    HOH     608    2HH2  ARG      96     3645     1.58
REMARK 500  2HH2  ARG      96    2H    HOH     585     3655     1.59
REMARK 500  2H    HOH     585    2HH2  ARG      96     3645     1.59
REMARK 500  1H    HOH     585    2HH2  ARG      96     3645     1.60
REMARK 500  2HH2  ARG      96    1H    HOH     585     3655     1.60
REMARK 500  1H    HOH     585    2HH1  ARG      96     3645     1.63
REMARK 500  2HH1  ARG      96    1H    HOH     585     3655     1.63
REMARK 500   O    HOH     917    1H    HOH     997     1655     1.70
REMARK 500  1H    HOH     997     O    HOH     917     1455     1.70
REMARK 500  1H    HOH     605    2HH2  ARG     211     2554     1.72
REMARK 500  1H    HOH     816     O    HOH     610     1455     1.72
REMARK 500   O    HOH     610    1H    HOH     816     1655     1.72
REMARK 500  2HH2  ARG     211    1H    HOH     605     2555     1.72
REMARK 500  1H    HOH     979     O    HOH     941     3645     1.73
REMARK 500   O    HOH     941    1H    HOH     979     3655     1.73
REMARK 500   O    HOH     979    1H    HOH     993     3645     1.74
REMARK 500  1H    HOH     993     O    HOH     979     3655     1.74
REMARK 500   O    HOH     911    1H    HOH     961     4456     1.75
REMARK 500  2HH1  ARG      96     O    HOH     585     3655     1.75
REMARK 500   O    HOH     585    2HH1  ARG      96     3645     1.75
REMARK 500  1H    HOH     961     O    HOH     911     4556     1.75
REMARK 500   HG1  THR     113     O    HOH     732     4456     1.76
REMARK 500   O    HOH     732     HG1  THR     113     4556     1.76
REMARK 500   O    HOH     816    1H    HOH     560     4456     1.77
REMARK 500  1H    HOH     560     O    HOH     816     4556     1.77
REMARK 500   O    HOH     993    2H    HOH     978     3655     1.78
REMARK 500  2H    HOH     978     O    HOH     993     3645     1.78
REMARK 500  1H    HOH     521     O    HOH     986     1455     1.79
REMARK 500   O    HOH     990    2H    HOH     573     3655     1.79
REMARK 500  2H    HOH     573     O    HOH     990     3645     1.79
REMARK 500   O    HOH     986    1H    HOH     521     1655     1.79
REMARK 500  2H    HOH     910     O    HOH     963     4456     1.81
REMARK 500   O    HOH     963    2H    HOH     910     4556     1.81
REMARK 500   OG1  THR     173    2H    HOH     518     1655     1.82
REMARK 500  2H    HOH     518     OG1  THR     173     1455     1.82
REMARK 500   H    GLY     174     O    HOH     521     1655     1.83
REMARK 500   O    HOH     521     H    GLY     174     1455     1.83
REMARK 500   O    HOH     598    2H    HOH     595     2654     1.84
REMARK 500  2H    HOH     595     O    HOH     598     2655     1.84
REMARK 500  2H    HOH     976     O    HOH     988     3645     1.85
REMARK 500   O    PRO     193    1H    HOH     550     3645     1.85
REMARK 500  2HH2  ARG     211    2H    HOH     605     2555     1.85
REMARK 500  2H    HOH     605    2HH2  ARG     211     2554     1.85
REMARK 500  2HH1  ARG     158     OD1  ASP     134     4556     1.85
REMARK 500   O    HOH     649    1H    HOH     724     4556     1.85
REMARK 500   OD1  ASP     134    2HH1  ARG     158     4456     1.85
REMARK 500  1H    HOH     724     O    HOH     649     4456     1.85
REMARK 500  1H    HOH     550     O    PRO     193     3655     1.85
REMARK 500   O    HOH     988    2H    HOH     976     3655     1.85
REMARK 500   O    HOH     558     H    GLY      46     2555     1.86
REMARK 500   H    GLY      46     O    HOH     558     2554     1.86
REMARK 500  2HZ   LYS     140    1H    HOH     994     4456     1.86
REMARK 500   O    HOH     644    2H    HOH     732     4456     1.86
REMARK 500  2HH2  ARG     158     OD2  ASP     134     4556     1.86
REMARK 500  2H    HOH     732     O    HOH     644     4556     1.86
REMARK 500   OD2  ASP     134    2HH2  ARG     158     4456     1.86
REMARK 500  1H    HOH     994    2HZ   LYS     140     4556     1.86
REMARK 500  2HD2  ASN     106     O    HOH     727     4456     1.87
REMARK 500   H    SER      92     OE1  GLU     196     3655     1.87
REMARK 500  1H    HOH     526     O    HOH     605     2555     1.87
REMARK 500   OE1  GLU     196     H    SER      92     3645     1.87
REMARK 500   O    HOH     605    1H    HOH     526     2554     1.87
REMARK 500   O    HOH     727    2HD2  ASN     106     4556     1.87
REMARK 500   O    HOH     961    2H    HOH     814     4456     1.88
REMARK 500   O    ALA     209    1H    HOH     598     2655     1.88
REMARK 500  2H    HOH     814     O    HOH     961     4556     1.88
REMARK 500   O    HOH     960    1H    HOH     649     4456     1.88
REMARK 500  1H    HOH     649     O    HOH     960     4556     1.88
REMARK 500  1H    HOH     598     O    ALA     209     2654     1.88
REMARK 500   O    ASN      25    2H    HOH     566     1455     1.89
REMARK 500   O    VAL     184    2H    HOH     537     2654     1.89
REMARK 500  2H    HOH     566     O    ASN      25     1655     1.89
REMARK 500   O    ALA     190    1H    HOH     553     3645     1.89
REMARK 500  1H    HOH     553     O    ALA     190     3655     1.89
REMARK 500  2H    HOH     537     O    VAL     184     2655     1.89
REMARK 500  2H    HOH     709     O    HOH     722     4556     1.90
REMARK 500  2HH2  ARG      96     O    HOH     585     3655     1.90
REMARK 500   O    HOH     722    2H    HOH     709     4456     1.90
REMARK 500   O    HOH     585    2HH2  ARG      96     3645     1.90
REMARK 500  1H    HOH     709     O    ALA     136     4556     1.92
REMARK 500  1H    HOH     960    1HH1  ARG     158     4456     1.92
REMARK 500   HE   ARG     211     O    HOH     719     2555     1.92
REMARK 500  2H    HOH     680     O    HOH     987     2554     1.92
REMARK 500   O    HOH     719     HE   ARG     211     2554     1.92
REMARK 500  1HH1  ARG     158    1H    HOH     960     4556     1.92
REMARK 500   O    ALA     136    1H    HOH     709     4456     1.92
REMARK 500   O    HOH     987    2H    HOH     680     2555     1.92
REMARK 500   O    GLU     131    2H    HOH     979     3655     1.93
REMARK 500  1H    HOH     988     O    HOH     607     3655     1.93
REMARK 500  1HD2  ASN     152     O    HOH     522     1655     1.93
REMARK 500   O    HOH     607    1H    HOH     988     3645     1.93
REMARK 500  1H    HOH     559     O    HOH     978     3655     1.93
REMARK 500   O    GLU     196    2H    HOH     970     3645     1.93
REMARK 500  2H    HOH     979     O    GLU     131     3645     1.93
REMARK 500  2H    HOH     970     O    GLU     196     3655     1.93
REMARK 500   O    HOH     978    1H    HOH     559     3645     1.93
REMARK 500   O    HOH     522    1HD2  ASN     152     1455     1.93
REMARK 500   O    GLY     157    2H    HOH     524     1655     1.94
REMARK 500  2H    HOH     524     O    GLY     157     1455     1.94
REMARK 500   H    VAL     169     ND2  ASN      27     1655     1.95
REMARK 500   ND2  ASN      27     H    VAL     169     1455     1.95
REMARK 500  1HD2  ASN      27     N    VAL     169     1455     1.96
REMARK 500   N    VAL     169    1HD2  ASN      27     1655     1.96
REMARK 500  2HH2  ARG     211     O    HOH     605     2555     1.97
REMARK 500   O    HOH     605    2HH2  ARG     211     2554     1.97
REMARK 500   O    THR     107    2H    HOH     707     4456     1.98
REMARK 500  2H    HOH     707     O    THR     107     4556     1.98
REMARK 500   O    HOH     682    2H    HOH     558     2554     1.99
REMARK 500   O    HOH     723    1H    HOH     962     4456     1.99
REMARK 500  1H    HOH     962     O    HOH     723     4556     1.99
REMARK 500  2H    HOH     558     O    HOH     682     2555     1.99
REMARK 500   O    HOH     703    2H    HOH     726     4456     2.00
REMARK 500  2H    HOH     726     O    HOH     703     4556     2.00
REMARK 500   O    SER     213    2H    HOH     520     2555     2.01
REMARK 500  2H    HOH     520     O    SER     213     2554     2.01
REMARK 500   H    GLY      46    2H    HOH     558     2554     2.03
REMARK 500  2H    HOH     558     H    GLY      46     2555     2.03
REMARK 500   OG1  THR     107    2H    HOH     727     4456     2.04
REMARK 500  2H    HOH     727     OG1  THR     107     4556     2.04
REMARK 500   OD1  ASN      58    1H    HOH     970     3645     2.05
REMARK 500  1H    HOH     925     O    HOH     595     2654     2.05
REMARK 500  1H    HOH     990    1HH2  ARG      17     3655     2.05
REMARK 500   O    HOH     595    1H    HOH     925     2655     2.05
REMARK 500  1H    HOH     970     OD1  ASN      58     3655     2.05
REMARK 500  1HH2  ARG      17    1H    HOH     990     3645     2.05
REMARK 500  2H    HOH     980    2H    HOH     816     2554     2.07
REMARK 500  2H    HOH     816    2H    HOH     980     2555     2.07
REMARK 500  2H    HOH     986    1H    HOH     521     1655     2.07
REMARK 500  1H    HOH     521    2H    HOH     986     1455     2.07
REMARK 500  1H    HOH     582    2H    HOH     951     3645     2.08
REMARK 500  2H    HOH     951    1H    HOH     582     3655     2.08
REMARK 500   O    HOH     608    2HH2  ARG      96     3645     2.08
REMARK 500  1H    HOH     605     O    ASP     111     2554     2.08
REMARK 500  2HH2  ARG      96     O    HOH     608     3655     2.08
REMARK 500   O    ASP     111    1H    HOH     605     2555     2.08
REMARK 500  2H    HOH     800     O    LYS     172     1455     2.09
REMARK 500  2H    HOH     978    2H    HOH     993     3645     2.09
REMARK 500  2H    HOH     993    2H    HOH     978     3655     2.09
REMARK 500   O    LYS     172    2H    HOH     800     1655     2.09
REMARK 500   O    HOH     910    2H    HOH     728     4456     2.10
REMARK 500  1H    HOH     574     OXT  SER     213     2554     2.10
REMARK 500  2H    HOH     728     O    HOH     910     4556     2.10
REMARK 500   OXT  SER     213    1H    HOH     574     2555     2.10
REMARK 500  2H    HOH     638     O    HOH     548     3645     2.11
REMARK 500   O    HOH     548    2H    HOH     638     3655     2.11
REMARK 500  1HD2  ASN      58    1H    HOH     970     3645     2.11
REMARK 500  1H    HOH     970    1HD2  ASN      58     3655     2.11
REMARK 500  2H    HOH     983     O    HOH     980     2555     2.12
REMARK 500   O    HOH     980    2H    HOH     983     2554     2.12
REMARK 500  1H    HOH     983     O    HOH     624     4456     2.13
REMARK 500   O    HOH     624    1H    HOH     983     4556     2.13
REMARK 500  2HD2  ASN      27     O    HOH     802     1455     2.14
REMARK 500  2HH1  ARG     138     O    HOH     703     4556     2.14
REMARK 500   O    HOH     703    2HH1  ARG     138     4456     2.14
REMARK 500  2H    HOH     598    2H    HOH     595     2654     2.14
REMARK 500  2H    HOH     595    2H    HOH     598     2655     2.14
REMARK 500   H    GLY     174    1H    HOH     521     1655     2.14
REMARK 500  1H    HOH     610    1H    HOH     816     1655     2.14
REMARK 500   O    HOH     802    2HD2  ASN      27     1655     2.14
REMARK 500  1H    HOH     521     H    GLY     174     1455     2.14
REMARK 500  1H    HOH     816    1H    HOH     610     1455     2.14
REMARK 500   O    HOH     537    1H    HOH     580     2655     2.15
REMARK 500  1H    HOH     580     O    HOH     537     2654     2.15
REMARK 500  1H    HOH     960    1H    HOH     649     4456     2.15
REMARK 500  1H    HOH     649    1H    HOH     960     4556     2.15
REMARK 500   O    HOH     976    2H    HOH     554     3645     2.16
REMARK 500  1H    HOH     608     NH2  ARG      96     3645     2.16
REMARK 500  2H    HOH     554     O    HOH     976     3655     2.16
REMARK 500   NH2  ARG      96    1H    HOH     608     3655     2.16
REMARK 500  2HH1  ARG     138    1H    HOH     703     4556     2.16
REMARK 500   O    HOH     993    2H    HOH     977     3655     2.16
REMARK 500  1H    HOH     703    2HH1  ARG     138     4456     2.16
REMARK 500  2H    HOH     977     O    HOH     993     3645     2.16
REMARK 500   O    SER     213    1H    HOH     574     2555     2.17
REMARK 500  1H    HOH     574     O    SER     213     2554     2.17
REMARK 500  2H    HOH     979    1H    HOH     993     3645     2.18
REMARK 500  1H    HOH     993    2H    HOH     979     3655     2.18
REMARK 500  1HH2  ARG      96    1H    HOH     608     3655     2.18
REMARK 500  1H    HOH     608    1HH2  ARG      96     3645     2.18
REMARK 500  1H    HOH     917    2H    HOH     998     1655     2.19
REMARK 500  1H    HOH     727    2HD2  ASN     106     4556     2.19
REMARK 500   HG   SER      28     O    HOH     960     4456     2.19
REMARK 500   O    HOH     960     HG   SER      28     4556     2.19
REMARK 500  2HD2  ASN     106    1H    HOH     727     4456     2.19
REMARK 500  2H    HOH     998    1H    HOH     917     1455     2.19
REMARK 500   O    HOH     987    2H    HOH     520     2555     2.20
REMARK 500  2H    HOH     520     O    HOH     987     2554     2.20
REMARK 500  1H    HOH     983    1H    HOH     624     4456     2.20
REMARK 500  1H    HOH     624    1H    HOH     983     4556     2.20
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: CAT
REMARK 800 SITE_DESCRIPTION: CATALYTIC TRIAD
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 1CUC       SWS     P00590       1 -    32 NOT IN ATOMS LIST
REMARK 999 1CUC       SWS     P00590     230 -   230 NOT IN ATOMS LIST
DBREF  1CUC     17   213  SWS    P00590   CUTI_FUSSO      33    229
SEQADV 1CUC ALA     32  SWS  P00590    ARG    48 CONFLICT
SEQADV 1CUC LYS    172  SWS  P00590    ASN   188 ENGINEERED
SEQADV 1CUC GLU    196  SWS  P00590    ARG   212 ENGINEERED
SEQRES   1    214  LEU PRO THR SER ASN PRO ALA GLN GLU LEU GLU ALA ARG
SEQRES   2    214  GLN LEU GLY ARG THR THR ARG ASP ASP LEU ILE ASN GLY
SEQRES   3    214  ASN SER ALA SER CYS ALA ASP VAL ILE PHE ILE TYR ALA
SEQRES   4    214  ARG GLY SER THR GLU THR GLY ASN LEU GLY THR LEU GLY
SEQRES   5    214  PRO SER ILE ALA SER ASN LEU GLU SER ALA PHE GLY LYS
SEQRES   6    214  ASP GLY VAL TRP ILE GLN GLY VAL GLY GLY ALA TYR ARG
SEQRES   7    214  ALA THR LEU GLY ASP ASN ALA LEU PRO ARG GLY THR SER
SEQRES   8    214  SER ALA ALA ILE ARG GLU MET LEU GLY LEU PHE GLN GLN
SEQRES   9    214  ALA ASN THR LYS CYS PRO ASP ALA THR LEU ILE ALA GLY
SEQRES  10    214  GLY TYR SER GLN GLY ALA ALA LEU ALA ALA ALA SER ILE
SEQRES  11    214  GLU ASP LEU ASP SER ALA ILE ARG ASP LYS ILE ALA GLY
SEQRES  12    214  THR VAL LEU PHE GLY TYR THR LYS ASN LEU GLN ASN ARG
SEQRES  13    214  GLY ARG ILE PRO ASN TYR PRO ALA ASP ARG THR LYS VAL
SEQRES  14    214  PHE CYS LYS THR GLY ASP LEU VAL CYS THR GLY SER LEU
SEQRES  15    214  ILE VAL ALA ALA PRO HIS LEU ALA TYR GLY PRO ASP ALA
SEQRES  16    214  GLU GLY PRO ALA PRO GLU PHE LEU ILE GLU LYS VAL ARG
SEQRES  17    214  ALA VAL ARG GLY SER ALA
FORMUL   2  HOH   *187(H2 O1)
HELIX    1   1 ASP     22  ASN     25  1                                   4
HELIX    2   2 SER     28  SER     30  5                                   3
HELIX    3   3 GLY     49  PHE     63  1                                  15
HELIX    4   4 LEU     81  ALA     85  5                                   5
HELIX    5   5 SER     92  LYS    108  1                                  17
HELIX    6   6 SER    120  ASP    132  5                                  13
HELIX    7   7 SER    135  LYS    140  1                                   6
HELIX    8   8 ALA    164  ARG    166  5                                   3
HELIX    9   9 LEU    176  THR    179  5                                   4
HELIX   10  10 ALA    186  LEU    189  5                                   4
HELIX   11  11 GLY    192  GLU    196  1                                   5
HELIX   12  12 PRO    198  ARG    211  1                                  14
SHEET    1   A 5 VAL    68  GLY    72  0
SHEET    2   A 5 VAL    34  ALA    39  1  N  VAL    34   O  TRP    69
SHEET    3   A 5 THR   113  TYR   119  1  N  THR   113   O  ILE    35
SHEET    4   A 5 ILE   141  PHE   147  1  N  ALA   142   O  LEU   114
SHEET    5   A 5 THR   167  PHE   170  1  N  LYS   168   O  THR   144
SSBOND   1 CYS     31    CYS    109
SSBOND   2 CYS    171    CYS    178
SITE     1 CAT  3 SER   120  HIS   188  ASP   175
CRYST1   37.000   64.200   71.400  90.00  90.00  90.00 P 21 21 21    4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.027027  0.000000  0.000000        0.00000
SCALE2      0.000000  0.015576  0.000000        0.00000
SCALE3      0.000000  0.000000  0.014006        0.00000