longtext: 1EB9-pdb

content
HEADER    LYASE                                   24-JUL-01   1EB9
TITLE     STRUCTURE DETERMINANTS OF SUBSTRATE SPECIFICITY OF
TITLE    2 HYDROXYNITRILE LYASE FROM MANIHOT ESCULENTA
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: HYDROXYNITRILE LYASE;
COMPND   3 SYNONYM: (S)-HYDROXYNITRILE LYASE, (S)-HYDROXYNITRILASE,
COMPND   4  OXYNITRILASE, HYDROXYNITRILE LYASE;
COMPND   5 CHAIN: A, B;
COMPND   6 EC: 4.2.1.37;
COMPND   7 ENGINEERED: YES;
COMPND   8 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MANIHOT ESCULENTA;
SOURCE   3 ORGANISM_COMMON: CASSAVA;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI
KEYWDS    HYDROXYNITRILE LYASE, SUBSTRATE SPECIFICITY, ACTIVE-SITE
KEYWDS   2 TUNNEL MUTANT
EXPDTA    X-RAY DIFFRACTION
AUTHOR    H.LAUBLE,B.MIEHLICH,S.FOERSTER,C.KOBLER,H.WAJANT,
AUTHOR   2 F.EFFENBERGER
REVDAT   1   01-JAN-02 1EB9    0
JRNL        AUTH   H.LAUBLE,B.MIEHLICH,S.FOERSTER,C.KOBLER,H.WAJANT,
JRNL        AUTH 2 F.EFFENBERGER
JRNL        TITL   STRUCTURE DETERMINANTS OF SUBSTRATE SPECIFICITY OF
JRNL        TITL 2 HYDROXYNITRILE LYASE FROM MANIHOT ESCULENTA.
JRNL        REF    PROTEIN SCI.                  V.  11    65 2002
JRNL        REFN   ASTM PRCIEI  US ISSN 0961-8368
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   H.LAUBLE,S.FOERSTER,B.MIEHLICH,H.WAJANT,
REMARK   1  AUTH 2 F.EFFENBERGER
REMARK   1  TITL   STRUCTURE OF HYDROXYNITRILE LYASE FROM MANIHOT
REMARK   1  TITL 2 ESCULENTA IN COMPLEX WITH SUBSTRATES ACETONE AND
REMARK   1  TITL 3 CHLOROACETONE: IMPLICATIONS FOR THE MECHANISM OF
REMARK   1  TITL 4 CYANOGENESIS.
REMARK   1  REF    ACTA CRYSTALLOGR.,SECT.D      V.  57   194 2001
REMARK   1  REFN   ASTM ABCRE6  DK ISSN 0907-4449
REMARK   2
REMARK   2 RESOLUTION. 2.1  ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : X-PLOR 3.851
REMARK   3   AUTHORS     : BRUNGER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.10
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 8.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 2.0
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 10000000.00
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.001000
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 97.1
REMARK   3   NUMBER OF REFLECTIONS             : 60760
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.197
REMARK   3   FREE R VALUE                     : 0.244
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 10.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 6079
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.003
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.10
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.23
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 94.1
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 8705
REMARK   3   BIN R VALUE           (WORKING SET) : 0.252
REMARK   3   BIN FREE R VALUE                    : 0.274
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 10.1
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 976
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.009
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4160
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 36
REMARK   3   SOLVENT ATOMS            : 271
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 21.6
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 28.7
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.00
REMARK   3    B22 (A**2) : 0.00
REMARK   3    B33 (A**2) : 0.00
REMARK   3    B12 (A**2) : 0.00
REMARK   3    B13 (A**2) : 0.00
REMARK   3    B23 (A**2) : 0.00
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.23
REMARK   3   ESD FROM SIGMAA              (A) : 0.19
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 8.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.27
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.16
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.015
REMARK   3   BOND ANGLES            (DEGREES) : 2.9
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 23.9
REMARK   3   IMPROPER ANGLES        (DEGREES) : 1.12
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.83  ; 1.50
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 2.77  ; 2.00
REMARK   3   SIDE-CHAIN BOND              (A**2) : 3.92  ; 2.00
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 6.04  ; 2.50
REMARK   3
REMARK   3  NCS MODEL : NONE
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PARAM19X.PRO
REMARK   3  PARAMETER FILE  2  : PARHBA.PRO
REMARK   3  TOPOLOGY FILE  1   : TOPH19X.PRO
REMARK   3  TOPOLOGY FILE  2   : TOPHBA.PRO
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 1EB9 COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  ON 25-JUL-2001.
REMARK 100 THE EBI ID CODE IS EBI-8384.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.4
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : EMBL/DESY, HAMBURG BEAMLINE X11
REMARK 200  BEAMLINE                       : X11
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.905
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 63477
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.1
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.0
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NONE
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.4
REMARK 200  DATA REDUNDANCY                : 6.5
REMARK 200  R MERGE                    (I) : 0.052
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 19.2
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.17
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.1
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.4
REMARK 200  R MERGE FOR SHELL          (I) : 0.154
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 5.1
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: AMORE
REMARK 200 STARTING MODEL: PDB ENTRY 1DWP
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS  (%): 71
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 4.3
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M NACITRAT,
REMARK 280  PH 5.4, 5% PEG8000, 28% MPD
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP:  P 41 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,1/2+Z
REMARK 290       3555   1/2-Y,1/2+X,1/4+Z
REMARK 290       4555   1/2+Y,1/2-X,3/4+Z
REMARK 290       5555   1/2-X,1/2+Y,1/4-Z
REMARK 290       6555   1/2+X,1/2-Y,3/4-Z
REMARK 290       7555   Y,X,-Z
REMARK 290       8555   -Y,-X,1/2-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       94.42000
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       53.17000
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       53.17000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       47.21000
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       53.17000
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       53.17000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      141.63000
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       53.17000
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       53.17000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       47.21000
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       53.17000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       53.17000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000      141.63000
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000       94.42000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT
REMARK 300 WHICH CONSISTS OF   2 CHAIN(S). SEE REMARK 350 FOR
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).
REMARK 300
REMARK 300 QUATERNARY STRUCTURE FOR THIS ENTRY: TETRAMERIC
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE:  1
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT2   2  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      188.84000
REMARK 400
REMARK 400 COMPOUND
REMARK 400  CHAIN A, B ENGINEERED MUTATION TRP127ALA
REMARK 400  CHAIN A, B RESIDUES -4 TO 1 ARE CLONING ARTEFACTS
REMARK 400  KNOWN TO BE INVOLVED IN THE RELEASE OF HCN FROM INJURED
REMARK 400  TISSUES (CYANOGENESIS)
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     PRO B    -4
REMARK 465     ILE B    -3
REMARK 465     SER B    -2
REMARK 465     LYS B    -1
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500
REMARK 500    SER A  80       49.75   -109.59
REMARK 500    ARG A 129       59.46   -118.09
REMARK 500    SER B  80       48.06   -103.83
REMARK 500    ARG B 129       55.53   -110.09
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES ARE GIVEN CHAIN IDENTIFIERS TO
REMARK 525 INDICATE THE PROTEIN CHAIN TO WHICH THEY ARE MOST CLOSELY
REMARK 525 ASSOCIATED WITH:
REMARK 525   PROTEIN CHAIN  SOLVENT CHAIN
REMARK 525     A              X
REMARK 525     B              Y
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 SITE_DESCRIPTION: HBA BINDING SITE FOR CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 SITE_DESCRIPTION: HBA BINDING SITE FOR CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 SITE_DESCRIPTION: HBA BINDING SITE FOR CHAIN B
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 SITE_DESCRIPTION: HBA BINDING SITE FOR CHAIN B
REMARK 800
REMARK 800 SITE_IDENTIFIER: ASA
REMARK 800 SITE_DESCRIPTION: RESIDUES INVOLVED IN CATALYSIS SITE A
REMARK 800
REMARK 800 SITE_IDENTIFIER: ASB
REMARK 800 SITE_DESCRIPTION: RESIDUES INVOLVED IN CATALYSIS SITE B
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 1DWO   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF HYDROXYNITRILE LYASE FROM
REMARK 900  MANIHOT ESCULENTA INCOMPLEX WITH SUBSTRATES
REMARK 900  ACETONE AND CHLOROACETONE:IMPLICATIONS FOR
REMARK 900  THE MECHANISM OF CYANOGENESIS
REMARK 900 RELATED ID: 1DWP   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF HYDROXYNITRILE LYASE FROM
REMARK 900  MANIHOT ESCULENTA AT 2.2 ANGSTROM RESOLUTION
REMARK 900 RELATED ID: 1DWQ   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF HYDROXYNITRILE LYASE FROM
REMARK 900  MANIHOT ESCULENTA INCOMPLEX WITH SUBSTRATES
REMARK 900  ACETONE AND CHLOROACETONE:IMPLICATIONS FOR
REMARK 900  THEMECHANISM OF CYANOGENESIS
REMARK 900 RELATED ID: 1E89   RELATED DB: PDB
REMARK 900  ON THE MECHANISM OF CYANOGENESIS CATALYZED
REMARK 900  BY HYDROXYNITRILE LYASE FROM MANIHOT ESCULENTA
REMARK 900  CRYSTAL STRUCTURE OF ACTIVE SITE MUTANT
REMARK 900  SER80ALA IN COMPLEX WITH ACETONE CYANOHYDRIN
REMARK 900 RELATED ID: 1E8D   RELATED DB: PDB
REMARK 900  ON THE MECHANISM OF CYANOGENESIS CATALYZED
REMARK 900  BY HYDROXYNITRILE LYASE FROM MANIHOT ESCULENTA
REMARK 900  CRYSTAL STRUCTURE OF ACTIVE SITE MUTANT
REMARK 900  SER80ALA IN COMPLEX WITH ACETONE CYANOHYDRIN
REMARK 900 RELATED ID: 1EB8   RELATED DB: PDB
REMARK 900  STRUCTURE DETERMINANTS OF SUBSTRATE SPECIFICITY
REMARK 900  OF HYDROXYNITRILE LYASE FROM MANIHOT ESCULENTA
DBREF  1EB9 A   -4     1  PDB    1EB9     1EB9            -4      1
DBREF  1EB9 A    2   258  SWS    P52705   HNL_MANES        1    257
DBREF  1EB9 B   -4     1  PDB    1EB9     1EB9            -4      1
DBREF  1EB9 B    2   258  SWS    P52705   HNL_MANES        1    257
SEQADV 1EB9 ALA A  128  SWS  P52705    TRP   127 ENGINEERED MUTATION
SEQADV 1EB9 ALA B  128  SWS  P52705    TRP   127 ENGINEERED MUTATION
SEQRES   1 A  262  PRO ILE SER LYS MET VAL THR ALA HIS PHE VAL LEU ILE
SEQRES   2 A  262  HIS THR ILE CYS HIS GLY ALA TRP ILE TRP HIS LYS LEU
SEQRES   3 A  262  LYS PRO ALA LEU GLU ARG ALA GLY HIS LYS VAL THR ALA
SEQRES   4 A  262  LEU ASP MET ALA ALA SER GLY ILE ASP PRO ARG GLN ILE
SEQRES   5 A  262  GLU GLN ILE ASN SER PHE ASP GLU TYR SER GLU PRO LEU
SEQRES   6 A  262  LEU THR PHE LEU GLU LYS LEU PRO GLN GLY GLU LYS VAL
SEQRES   7 A  262  ILE ILE VAL GLY GLU SER CYS ALA GLY LEU ASN ILE ALA
SEQRES   8 A  262  ILE ALA ALA ASP ARG TYR VAL ASP LYS ILE ALA ALA GLY
SEQRES   9 A  262  VAL PHE HIS ASN SER LEU LEU PRO ASP THR VAL HIS SER
SEQRES  10 A  262  PRO SER TYR THR VAL GLU LYS LEU LEU GLU SER PHE PRO
SEQRES  11 A  262  ASP ALA ARG ASP THR GLU TYR PHE THR PHE THR ASN ILE
SEQRES  12 A  262  THR GLY GLU THR ILE THR THR MET LYS LEU GLY PHE VAL
SEQRES  13 A  262  LEU LEU ARG GLU ASN LEU PHE THR LYS CYS THR ASP GLY
SEQRES  14 A  262  GLU TYR GLU LEU ALA LYS MET VAL MET ARG LYS GLY SER
SEQRES  15 A  262  LEU PHE GLN ASN VAL LEU ALA GLN ARG PRO LYS PHE THR
SEQRES  16 A  262  GLU LYS GLY TYR GLY SER ILE LYS LYS VAL TYR ILE TRP
SEQRES  17 A  262  THR ASP GLN ASP LYS ILE PHE LEU PRO ASP PHE GLN ARG
SEQRES  18 A  262  TRP GLN ILE ALA ASN TYR LYS PRO ASP LYS VAL TYR GLN
SEQRES  19 A  262  VAL GLN GLY GLY ASP HIS LYS LEU GLN LEU THR LYS THR
SEQRES  20 A  262  GLU GLU VAL ALA HIS ILE LEU GLN GLU VAL ALA ASP ALA
SEQRES  21 A  262  TYR ALA
SEQRES   1 B  262  PRO ILE SER LYS MET VAL THR ALA HIS PHE VAL LEU ILE
SEQRES   2 B  262  HIS THR ILE CYS HIS GLY ALA TRP ILE TRP HIS LYS LEU
SEQRES   3 B  262  LYS PRO ALA LEU GLU ARG ALA GLY HIS LYS VAL THR ALA
SEQRES   4 B  262  LEU ASP MET ALA ALA SER GLY ILE ASP PRO ARG GLN ILE
SEQRES   5 B  262  GLU GLN ILE ASN SER PHE ASP GLU TYR SER GLU PRO LEU
SEQRES   6 B  262  LEU THR PHE LEU GLU LYS LEU PRO GLN GLY GLU LYS VAL
SEQRES   7 B  262  ILE ILE VAL GLY GLU SER CYS ALA GLY LEU ASN ILE ALA
SEQRES   8 B  262  ILE ALA ALA ASP ARG TYR VAL ASP LYS ILE ALA ALA GLY
SEQRES   9 B  262  VAL PHE HIS ASN SER LEU LEU PRO ASP THR VAL HIS SER
SEQRES  10 B  262  PRO SER TYR THR VAL GLU LYS LEU LEU GLU SER PHE PRO
SEQRES  11 B  262  ASP ALA ARG ASP THR GLU TYR PHE THR PHE THR ASN ILE
SEQRES  12 B  262  THR GLY GLU THR ILE THR THR MET LYS LEU GLY PHE VAL
SEQRES  13 B  262  LEU LEU ARG GLU ASN LEU PHE THR LYS CYS THR ASP GLY
SEQRES  14 B  262  GLU TYR GLU LEU ALA LYS MET VAL MET ARG LYS GLY SER
SEQRES  15 B  262  LEU PHE GLN ASN VAL LEU ALA GLN ARG PRO LYS PHE THR
SEQRES  16 B  262  GLU LYS GLY TYR GLY SER ILE LYS LYS VAL TYR ILE TRP
SEQRES  17 B  262  THR ASP GLN ASP LYS ILE PHE LEU PRO ASP PHE GLN ARG
SEQRES  18 B  262  TRP GLN ILE ALA ASN TYR LYS PRO ASP LYS VAL TYR GLN
SEQRES  19 B  262  VAL GLN GLY GLY ASP HIS LYS LEU GLN LEU THR LYS THR
SEQRES  20 B  262  GLU GLU VAL ALA HIS ILE LEU GLN GLU VAL ALA ASP ALA
SEQRES  21 B  262  TYR ALA
HET    HBA  A1259       9
HET    HBA  A1260       9
HET    HBA  B1259       9
HET    HBA  B1260       9
HETNAM     HBA P-HYDROXYBENZALDEHYDE
FORMUL   3  HBA    4(C7 H6 O2)
FORMUL   4  HOH   *271(H2 O1)
HELIX    1   1 GLY A   15  HIS A   20  5                                   6
HELIX    2   2 LYS A   21  ALA A   29  1                                   9
HELIX    3   3 GLN A   47  ILE A   51  5                                   5
HELIX    4   4 SER A   53  SER A   58  1                                   6
HELIX    5   5 SER A   58  LEU A   68  1                                  11
HELIX    6   6 ALA A   82  VAL A   94  1                                  13
HELIX    7   7 SER A  115  PHE A  125  1                                  11
HELIX    8   8 GLY A  150  ASN A  157  1                                   8
HELIX    9   9 THR A  163  MET A  174  1                                  12
HELIX   10  10 PHE A  180  GLN A  186  1                                   7
HELIX   11  11 GLY A  194  ILE A  198  5                                   5
HELIX   12  12 LEU A  212  TYR A  223  1                                  12
HELIX   13  13 LYS A  237  LYS A  242  1                                   6
HELIX   14  14 LYS A  242  ALA A  258  1                                  17
HELIX   15  15 GLY B   15  HIS B   20  5                                   6
HELIX   16  16 LYS B   21  ALA B   29  1                                   9
HELIX   17  17 GLN B   47  ILE B   51  5                                   5
HELIX   18  18 SER B   53  SER B   58  1                                   6
HELIX   19  19 SER B   58  LEU B   68  1                                  11
HELIX   20  20 ALA B   82  VAL B   94  1                                  13
HELIX   21  21 SER B  115  PHE B  125  1                                  11
HELIX   22  22 GLY B  150  ASN B  157  1                                   8
HELIX   23  23 THR B  163  MET B  174  1                                  12
HELIX   24  24 PHE B  180  ARG B  187  1                                   8
HELIX   25  25 GLY B  194  ILE B  198  5                                   5
HELIX   26  26 LEU B  212  TYR B  223  1                                  12
HELIX   27  27 LYS B  237  LYS B  242  1                                   6
HELIX   28  28 LYS B  242  ALA B  258  1                                  17
SHEET    1  AA 6 LYS A  32  LEU A  36  0
SHEET    2  AA 6 HIS A   5  ILE A   9  1  O  PHE A   6   N  THR A  34
SHEET    3  AA 6 VAL A  74  GLU A  79  1  O  ILE A  75   N  VAL A   7
SHEET    4  AA 6 ILE A  97  HIS A 103  1  N  ALA A  98   O  VAL A  74
SHEET    5  AA 6 LYS A 200  TRP A 204  1  O  VAL A 201   N  PHE A 102
SHEET    6  AA 6 LYS A 227  GLN A 230  1  O  LYS A 227   N  TYR A 202
SHEET    1  AB 2 GLU A 132  THR A 137  0
SHEET    2  AB 2 THR A 143  LYS A 148 -1  O  ILE A 144   N  PHE A 136
SHEET    1  BA 6 LYS B  32  LEU B  36  0
SHEET    2  BA 6 HIS B   5  ILE B   9  1  O  PHE B   6   N  THR B  34
SHEET    3  BA 6 VAL B  74  GLU B  79  1  O  ILE B  75   N  VAL B   7
SHEET    4  BA 6 ILE B  97  HIS B 103  1  N  ALA B  98   O  VAL B  74
SHEET    5  BA 6 LYS B 200  TRP B 204  1  O  VAL B 201   N  PHE B 102
SHEET    6  BA 6 LYS B 227  GLN B 230  1  O  LYS B 227   N  TYR B 202
SHEET    1  BB 2 GLU B 132  THR B 137  0
SHEET    2  BB 2 THR B 143  LYS B 148 -1  O  ILE B 144   N  PHE B 136
SITE     1 AC1  8 THR A  11  ILE A  12  SER A  80  CYS A  81
SITE     2 AC1  8 MET A 147  LEU A 179  HOH X 122  HOH X 133
SITE     1 AC2  3 PHE A 125  ILE A 210  HOH X 122
SITE     1 AC3  7 THR B  11  ILE B  12  SER B  80  CYS B  81
SITE     2 AC3  7 MET B 147  HOH Y 137  HOH Y 138
SITE     1 AC4  5 LEU B 122  PHE B 125  ALA B 128  ILE B 210
SITE     2 AC4  5 HOH Y 137
SITE     1 ASA  4 THR A  11  SER A  80  ASP A 208  HIS A 236
SITE     1 ASB  4 THR B  11  SER B  80  ASP B 208  HIS B 236
CRYST1  106.340  106.340  188.840  90.00  90.00  90.00 P 41 21 2    16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.009404  0.000000  0.000000        0.00000
SCALE2      0.000000  0.009404  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005295        0.00000
END