longtext: 1EK2-pdb

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HEADER    HYDROLASE                               06-MAR-00   1EK2
TITLE     CRYSTAL STRUCTURE OF MURINE SOLUBLE EPOXIDE HYDROLASE
TITLE    2 COMPLEXED WITH CDU INHIBITOR
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: EPOXIDE HYDROLASE;
COMPND   3 CHAIN: A, B;
COMPND   4 EC: 3.3.2.3;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;
SOURCE   3 ORGANISM_COMMON: MOUSE;
SOURCE   4 EXPRESSION_SYSTEM_COMMON: INSECT;
SOURCE   5 EXPRESSION_SYSTEM_VECTOR_TYPE: VIRUS;
SOURCE   6 EXPRESSION_SYSTEM_VECTOR: BACCULOVIRUS
KEYWDS    HOMODIMER, ALPHA/BETA HYDROLASE FOLD, DISUBSTITUTED UREA
KEYWDS   2 INHIBITOR
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.A.ARGIRIADI,C.MORISSEAU,M.H.GOODROW,D.L.DOWDY,B.D.HAMMOCK,
AUTHOR   2 D.W.CHRISTIANSON
REVDAT   1   31-MAY-00 1EK2    0
JRNL        AUTH   M.A.ARGIRIADI,C.MORISSEAU,M.H.GOODROW,D.L.DOWDY,
JRNL        AUTH 2 B.D.HAMMOCK,D.W.CHRISTIANSON
JRNL        TITL   BINDING OF ALKYLUREA INHIBITORS TO EPOXIDE
JRNL        TITL 2 HYDROLASE IMPLICATES ACTIVE SITE TYROSINES IN
JRNL        TITL 3 SUBSTRATE ACTIVATION
JRNL        REF    J.BIOL.CHEM.                  V. 275 15265 2000
JRNL        REFN   ASTM JBCHA3  US ISSN 0021-9258
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   M.A.ARGIRIADI,C.MORISSEAU,B.D.HAMMOCK,
REMARK   1  AUTH 2 D.W.CHRISTIANSON
REMARK   1  TITL   DETOXIFICATION OF ENVIRONMENTAL MUTAGENS AND
REMARK   1  TITL 2 CARCINOGENS: STRUCTURE, MECHANISM, AND EVOLUTION
REMARK   1  TITL 3 OF LIVER EPOXIDE HYDROLASE
REMARK   1  REF    PROC.NAT.ACAD.SCI.USA         V.  96 10637 1999
REMARK   1  REFN   ASTM PNASA6  US ISSN 0027-8424
REMARK   2
REMARK   2 RESOLUTION. 3.00 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : X-PLOR 3.851
REMARK   3   AUTHORS     : BRUNGER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 3.00
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 2.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL
REMARK   3   NUMBER OF REFLECTIONS             : 18983
REMARK   3
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.211
REMARK   3   FREE R VALUE                     : 0.290
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : 924
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : NULL
REMARK   3   BIN FREE R VALUE                    : NULL
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 8178
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 40
REMARK   3   SOLVENT ATOMS            : 19
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 63.60
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.007
REMARK   3   BOND ANGLES            (DEGREES) : 1.20
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 1EK2 COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 21-MAR-2000.
REMARK 100 THE RCSB ID CODE IS RCSB010660.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 30-AUG-1998
REMARK 200  TEMPERATURE           (KELVIN) : 200.0
REMARK 200  PH                             : 6.00
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : CHESS
REMARK 200  BEAMLINE                       : A1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.89000
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : OTHER
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 46217
REMARK 200  RESOLUTION RANGE HIGH      (A) : 3.000
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 79.4
REMARK 200  DATA REDUNDANCY                : 2.340
REMARK 200  R MERGE                    (I) : 0.06900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.5000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 3.00
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 20.00
REMARK 200  COMPLETENESS FOR SHELL     (%) : 58.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.26
REMARK 200  R MERGE FOR SHELL          (I) : 0.28200
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: NULL
REMARK 200 SOFTWARE USED: X-PLOR
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): NULL
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: AMMONIUM SULFATE, MES,
REMARK 280  ETHANOL, DITHIOTHREITOL, CDU (N-CYCLOHEXYL-N'-DECYLUREA)
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   1/2-X,1/2+Y,-Z
REMARK 290       4555   1/2+X,1/2-Y,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000       75.95000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       71.50000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       75.95000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       71.50000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ALA A     2
REMARK 465     LEU A     3
REMARK 465     ALA A    20
REMARK 465     GLY A    21
REMARK 465     ALA A    22
REMARK 465     PHE A    23
REMARK 465     ARG A    24
REMARK 465     ARG A    25
REMARK 465     SER A    26
REMARK 465     GLU A    27
REMARK 465     GLU A    28
REMARK 465     ALA A    29
REMARK 465     LEU A    30
REMARK 465     ALA A    31
REMARK 465     LEU A    32
REMARK 465     PRO A    33
REMARK 465     ARG A    34
REMARK 465     ASP A    35
REMARK 465     PHE A    36
REMARK 465     LEU A    37
REMARK 465     LEU A    38
REMARK 465     GLY A    39
REMARK 465     ALA A    40
REMARK 465     TYR A    41
REMARK 465     GLN A    42
REMARK 465     THR A    43
REMARK 465     GLU A    44
REMARK 465     PHE A    45
REMARK 465     PRO A    46
REMARK 465     GLU A    47
REMARK 465     VAL A    64
REMARK 465     PRO A    65
REMARK 465     LEU A    66
REMARK 465     MET A    67
REMARK 465     ASP A    68
REMARK 465     GLU A    69
REMARK 465     SER A    70
REMARK 465     TYR A    71
REMARK 465     ARG A    72
REMARK 465     LYS A    73
REMARK 465     SER A    74
REMARK 465     SER A    75
REMARK 465     LYS A    76
REMARK 465     ALA A    77
REMARK 465     CYS A    78
REMARK 465     GLY A    79
REMARK 465     ALA A    80
REMARK 465     ASN A    81
REMARK 465     LEU A    82
REMARK 465     PRO A    83
REMARK 465     GLU A    84
REMARK 465     ASN A    85
REMARK 465     PHE A    86
REMARK 465     SER A    87
REMARK 465     ILE A    88
REMARK 465     SER A    89
REMARK 465     VAL A   545
REMARK 465     GLN A   546
REMARK 465     ASN A   547
REMARK 465     PRO A   548
REMARK 465     SER A   549
REMARK 465     VAL A   550
REMARK 465     THR A   551
REMARK 465     SER A   552
REMARK 465     LYS A   553
REMARK 465     ILE A   554
REMARK 465     MET B     1
REMARK 465     ALA B     2
REMARK 465     LEU B     3
REMARK 465     VAL B   545
REMARK 465     GLN B   546
REMARK 465     ASN B   547
REMARK 465     PRO B   548
REMARK 465     SER B   549
REMARK 465     VAL B   550
REMARK 465     THR B   551
REMARK 465     SER B   552
REMARK 465     LYS B   553
REMARK 465     ILE B   554
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    MET A  54   CE    MET A  54   SD     0.059
REMARK 500    GLN A  62   CG    GLN A  62   CB     0.051
REMARK 500    MET A 139   CE    MET A 139   SD     0.054
REMARK 500    MET A 140   CE    MET A 140   SD     0.050
REMARK 500    PRO A 161   CG    PRO A 161   CB     0.076
REMARK 500    PRO A 232   CG    PRO A 232   CB     0.102
REMARK 500    MET A 253   CE    MET A 253   SD     0.051
REMARK 500    PRO A 266   CG    PRO A 266   CB     0.086
REMARK 500    PRO B  83   CG    PRO B  83   CB     0.055
REMARK 500    MET B 105   CE    MET B 105   SD     0.053
REMARK 500    MET B 158   CE    MET B 158   SD     0.050
REMARK 500    PRO B 161   CG    PRO B 161   CB     0.064
REMARK 500    PRO B 229   CG    PRO B 229   CB     0.054
REMARK 500    MET B 253   CE    MET B 253   SD     0.075
REMARK 500    PRO B 266   CG    PRO B 266   CB     0.073
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    GLU A  51   N   -  CA  -  C   ANGL. DEV. = -8.4 DEGREES
REMARK 500    GLY A  56   N   -  CA  -  C   ANGL. DEV. =  7.9 DEGREES
REMARK 500    ARG A 103   N   -  CA  -  C   ANGL. DEV. =  8.2 DEGREES
REMARK 500    ASN A 233   N   -  CA  -  C   ANGL. DEV. = -9.0 DEGREES
REMARK 500    LYS A 243   N   -  CA  -  C   ANGL. DEV. = -8.5 DEGREES
REMARK 500    ASP A 290   N   -  CA  -  C   ANGL. DEV. = -9.1 DEGREES
REMARK 500    GLY A 295   N   -  CA  -  C   ANGL. DEV. =  8.3 DEGREES
REMARK 500    VAL A 328   N   -  CA  -  C   ANGL. DEV. = -8.6 DEGREES
REMARK 500    GLY A 414   N   -  CA  -  C   ANGL. DEV. =  9.0 DEGREES
REMARK 500    THR A 422   N   -  CA  -  C   ANGL. DEV. = -9.3 DEGREES
REMARK 500    ASN A 430   N   -  CA  -  C   ANGL. DEV. = -9.7 DEGREES
REMARK 500    ARG A 466   N   -  CA  -  C   ANGL. DEV. =  8.8 DEGREES
REMARK 500    THR A 468   N   -  CA  -  C   ANGL. DEV. =  8.9 DEGREES
REMARK 500    VAL A 497   N   -  CA  -  C   ANGL. DEV. = -9.5 DEGREES
REMARK 500    LYS A 504   N   -  CA  -  C   ANGL. DEV. =  9.2 DEGREES
REMARK 500    GLY A 516   N   -  CA  -  C   ANGL. DEV. = -8.5 DEGREES
REMARK 500    VAL B   5   N   -  CA  -  C   ANGL. DEV. =  9.8 DEGREES
REMARK 500    ALA B 224   N   -  CA  -  C   ANGL. DEV. = -8.0 DEGREES
REMARK 500    LYS B 243   N   -  CA  -  C   ANGL. DEV. = -8.6 DEGREES
REMARK 500    ASP B 290   N   -  CA  -  C   ANGL. DEV. = -8.9 DEGREES
REMARK 500    GLY B 295   N   -  CA  -  C   ANGL. DEV. =  8.2 DEGREES
REMARK 500    VAL B 328   N   -  CA  -  C   ANGL. DEV. = -9.2 DEGREES
REMARK 500    GLY B 414   N   -  CA  -  C   ANGL. DEV. =  8.8 DEGREES
REMARK 500    ARG B 466   N   -  CA  -  C   ANGL. DEV. =  8.6 DEGREES
REMARK 500    THR B 468   N   -  CA  -  C   ANGL. DEV. =  8.9 DEGREES
REMARK 500    VAL B 497   N   -  CA  -  C   ANGL. DEV. = -9.3 DEGREES
REMARK 500    LYS B 504   N   -  CA  -  C   ANGL. DEV. =  9.1 DEGREES
REMARK 500    GLY B 516   N   -  CA  -  C   ANGL. DEV. = -8.4 DEGREES
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    HIS A 204       64.50    -57.44
REMARK 500    ALA A 207      124.37   -128.65
REMARK 500    SER A 208       51.34    -79.05
REMARK 500    ASN A 357       65.92    -63.94
REMARK 500    GLU A 423      130.60    -67.79
REMARK 500    THR A 431       49.94    146.47
REMARK 500    VAL B   5       57.81    150.09
REMARK 500    ASN B 205       71.49    -96.61
REMARK 500    ASN B 357       65.77    -63.48
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 1CQZ   RELATED DB: PDB
REMARK 900 RELATED ID: 1CR6   RELATED DB: PDB
REMARK 900 RELATED ID: 1EK1   RELATED DB: PDB
DBREF  1EK2 A    1   554  SWS    P34914   HYES_MOUSE       1    554
DBREF  1EK2 B    1   554  SWS    P34914   HYES_MOUSE       1    554
SEQRES   1 A  554  MET ALA LEU ARG VAL ALA ALA PHE ASP LEU ASP GLY VAL
SEQRES   2 A  554  LEU ALA LEU PRO SER ILE ALA GLY ALA PHE ARG ARG SER
SEQRES   3 A  554  GLU GLU ALA LEU ALA LEU PRO ARG ASP PHE LEU LEU GLY
SEQRES   4 A  554  ALA TYR GLN THR GLU PHE PRO GLU GLY PRO THR GLU GLN
SEQRES   5 A  554  LEU MET LYS GLY LYS ILE THR PHE SER GLN TRP VAL PRO
SEQRES   6 A  554  LEU MET ASP GLU SER TYR ARG LYS SER SER LYS ALA CYS
SEQRES   7 A  554  GLY ALA ASN LEU PRO GLU ASN PHE SER ILE SER GLN ILE
SEQRES   8 A  554  PHE SER GLN ALA MET ALA ALA ARG SER ILE ASN ARG PRO
SEQRES   9 A  554  MET LEU GLN ALA ALA ILE ALA LEU LYS LYS LYS GLY PHE
SEQRES  10 A  554  THR THR CYS ILE VAL THR ASN ASN TRP LEU ASP ASP GLY
SEQRES  11 A  554  ASP LYS ARG ASP SER LEU ALA GLN MET MET CYS GLU LEU
SEQRES  12 A  554  SER GLN HIS PHE ASP PHE LEU ILE GLU SER CYS GLN VAL
SEQRES  13 A  554  GLY MET ILE LYS PRO GLU PRO GLN ILE TYR ASN PHE LEU
SEQRES  14 A  554  LEU ASP THR LEU LYS ALA LYS PRO ASN GLU VAL VAL PHE
SEQRES  15 A  554  LEU ASP ASP PHE GLY SER ASN LEU LYS PRO ALA ARG ASP
SEQRES  16 A  554  MET GLY MET VAL THR ILE LEU VAL HIS ASN THR ALA SER
SEQRES  17 A  554  ALA LEU ARG GLU LEU GLU LYS VAL THR GLY THR GLN PHE
SEQRES  18 A  554  PRO GLU ALA PRO LEU PRO VAL PRO CYS ASN PRO ASN ASP
SEQRES  19 A  554  VAL SER HIS GLY TYR VAL THR VAL LYS PRO GLY ILE ARG
SEQRES  20 A  554  LEU HIS PHE VAL GLU MET GLY SER GLY PRO ALA LEU CYS
SEQRES  21 A  554  LEU CYS HIS GLY PHE PRO GLU SER TRP PHE SER TRP ARG
SEQRES  22 A  554  TYR GLN ILE PRO ALA LEU ALA GLN ALA GLY PHE ARG VAL
SEQRES  23 A  554  LEU ALA ILE ASP MET LYS GLY TYR GLY ASP SER SER SER
SEQRES  24 A  554  PRO PRO GLU ILE GLU GLU TYR ALA MET GLU LEU LEU CYS
SEQRES  25 A  554  LYS GLU MET VAL THR PHE LEU ASP LYS LEU GLY ILE PRO
SEQRES  26 A  554  GLN ALA VAL PHE ILE GLY HIS ASP TRP ALA GLY VAL MET
SEQRES  27 A  554  VAL TRP ASN MET ALA LEU PHE TYR PRO GLU ARG VAL ARG
SEQRES  28 A  554  ALA VAL ALA SER LEU ASN THR PRO PHE MET PRO PRO ASP
SEQRES  29 A  554  PRO ASP VAL SER PRO MET LYS VAL ILE ARG SER ILE PRO
SEQRES  30 A  554  VAL PHE ASN TYR GLN LEU TYR PHE GLN GLU PRO GLY VAL
SEQRES  31 A  554  ALA GLU ALA GLU LEU GLU LYS ASN MET SER ARG THR PHE
SEQRES  32 A  554  LYS SER PHE PHE ARG ALA SER ASP GLU THR GLY PHE ILE
SEQRES  33 A  554  ALA VAL HIS LYS ALA THR GLU ILE GLY GLY ILE LEU VAL
SEQRES  34 A  554  ASN THR PRO GLU ASP PRO ASN LEU SER LYS ILE THR THR
SEQRES  35 A  554  GLU GLU GLU ILE GLU PHE TYR ILE GLN GLN PHE LYS LYS
SEQRES  36 A  554  THR GLY PHE ARG GLY PRO LEU ASN TRP TYR ARG ASN THR
SEQRES  37 A  554  GLU ARG ASN TRP LYS TRP SER CYS LYS GLY LEU GLY ARG
SEQRES  38 A  554  LYS ILE LEU VAL PRO ALA LEU MET VAL THR ALA GLU LYS
SEQRES  39 A  554  ASP ILE VAL LEU ARG PRO GLU MET SER LYS ASN MET GLU
SEQRES  40 A  554  LYS TRP ILE PRO PHE LEU LYS ARG GLY HIS ILE GLU ASP
SEQRES  41 A  554  CYS GLY HIS TRP THR GLN ILE GLU LYS PRO THR GLU VAL
SEQRES  42 A  554  ASN GLN ILE LEU ILE LYS TRP LEU GLN THR GLU VAL GLN
SEQRES  43 A  554  ASN PRO SER VAL THR SER LYS ILE
SEQRES   1 B  554  MET ALA LEU ARG VAL ALA ALA PHE ASP LEU ASP GLY VAL
SEQRES   2 B  554  LEU ALA LEU PRO SER ILE ALA GLY ALA PHE ARG ARG SER
SEQRES   3 B  554  GLU GLU ALA LEU ALA LEU PRO ARG ASP PHE LEU LEU GLY
SEQRES   4 B  554  ALA TYR GLN THR GLU PHE PRO GLU GLY PRO THR GLU GLN
SEQRES   5 B  554  LEU MET LYS GLY LYS ILE THR PHE SER GLN TRP VAL PRO
SEQRES   6 B  554  LEU MET ASP GLU SER TYR ARG LYS SER SER LYS ALA CYS
SEQRES   7 B  554  GLY ALA ASN LEU PRO GLU ASN PHE SER ILE SER GLN ILE
SEQRES   8 B  554  PHE SER GLN ALA MET ALA ALA ARG SER ILE ASN ARG PRO
SEQRES   9 B  554  MET LEU GLN ALA ALA ILE ALA LEU LYS LYS LYS GLY PHE
SEQRES  10 B  554  THR THR CYS ILE VAL THR ASN ASN TRP LEU ASP ASP GLY
SEQRES  11 B  554  ASP LYS ARG ASP SER LEU ALA GLN MET MET CYS GLU LEU
SEQRES  12 B  554  SER GLN HIS PHE ASP PHE LEU ILE GLU SER CYS GLN VAL
SEQRES  13 B  554  GLY MET ILE LYS PRO GLU PRO GLN ILE TYR ASN PHE LEU
SEQRES  14 B  554  LEU ASP THR LEU LYS ALA LYS PRO ASN GLU VAL VAL PHE
SEQRES  15 B  554  LEU ASP ASP PHE GLY SER ASN LEU LYS PRO ALA ARG ASP
SEQRES  16 B  554  MET GLY MET VAL THR ILE LEU VAL HIS ASN THR ALA SER
SEQRES  17 B  554  ALA LEU ARG GLU LEU GLU LYS VAL THR GLY THR GLN PHE
SEQRES  18 B  554  PRO GLU ALA PRO LEU PRO VAL PRO CYS ASN PRO ASN ASP
SEQRES  19 B  554  VAL SER HIS GLY TYR VAL THR VAL LYS PRO GLY ILE ARG
SEQRES  20 B  554  LEU HIS PHE VAL GLU MET GLY SER GLY PRO ALA LEU CYS
SEQRES  21 B  554  LEU CYS HIS GLY PHE PRO GLU SER TRP PHE SER TRP ARG
SEQRES  22 B  554  TYR GLN ILE PRO ALA LEU ALA GLN ALA GLY PHE ARG VAL
SEQRES  23 B  554  LEU ALA ILE ASP MET LYS GLY TYR GLY ASP SER SER SER
SEQRES  24 B  554  PRO PRO GLU ILE GLU GLU TYR ALA MET GLU LEU LEU CYS
SEQRES  25 B  554  LYS GLU MET VAL THR PHE LEU ASP LYS LEU GLY ILE PRO
SEQRES  26 B  554  GLN ALA VAL PHE ILE GLY HIS ASP TRP ALA GLY VAL MET
SEQRES  27 B  554  VAL TRP ASN MET ALA LEU PHE TYR PRO GLU ARG VAL ARG
SEQRES  28 B  554  ALA VAL ALA SER LEU ASN THR PRO PHE MET PRO PRO ASP
SEQRES  29 B  554  PRO ASP VAL SER PRO MET LYS VAL ILE ARG SER ILE PRO
SEQRES  30 B  554  VAL PHE ASN TYR GLN LEU TYR PHE GLN GLU PRO GLY VAL
SEQRES  31 B  554  ALA GLU ALA GLU LEU GLU LYS ASN MET SER ARG THR PHE
SEQRES  32 B  554  LYS SER PHE PHE ARG ALA SER ASP GLU THR GLY PHE ILE
SEQRES  33 B  554  ALA VAL HIS LYS ALA THR GLU ILE GLY GLY ILE LEU VAL
SEQRES  34 B  554  ASN THR PRO GLU ASP PRO ASN LEU SER LYS ILE THR THR
SEQRES  35 B  554  GLU GLU GLU ILE GLU PHE TYR ILE GLN GLN PHE LYS LYS
SEQRES  36 B  554  THR GLY PHE ARG GLY PRO LEU ASN TRP TYR ARG ASN THR
SEQRES  37 B  554  GLU ARG ASN TRP LYS TRP SER CYS LYS GLY LEU GLY ARG
SEQRES  38 B  554  LYS ILE LEU VAL PRO ALA LEU MET VAL THR ALA GLU LYS
SEQRES  39 B  554  ASP ILE VAL LEU ARG PRO GLU MET SER LYS ASN MET GLU
SEQRES  40 B  554  LYS TRP ILE PRO PHE LEU LYS ARG GLY HIS ILE GLU ASP
SEQRES  41 B  554  CYS GLY HIS TRP THR GLN ILE GLU LYS PRO THR GLU VAL
SEQRES  42 B  554  ASN GLN ILE LEU ILE LYS TRP LEU GLN THR GLU VAL GLN
SEQRES  43 B  554  ASN PRO SER VAL THR SER LYS ILE
HET    CDU   1100      20
HET    CDU   1200      20
HETNAM     CDU N-CYCLOHEXYL-N'-DECYLUREA
FORMUL   3  CDU    2(C17 H34 N2 O1)
FORMUL   5  HOH   *19(H2 O1)
HELIX    1   1 THR A   50  GLY A   56  1                                   7
HELIX    2   2 SER A   93  ARG A   99  1                                   7
HELIX    3   3 ASN A  102  GLY A  116  1                                  15
HELIX    4   4 ARG A  133  SER A  144  1                                  12
HELIX    5   5 GLN A  145  PHE A  147  5                                   3
HELIX    6   6 PRO A  163  LYS A  174  1                                  12
HELIX    7   7 LYS A  176  ASN A  178  5                                   3
HELIX    8   8 ASN A  189  GLY A  197  1                                   9
HELIX    9   9 SER A  208  VAL A  216  1                                   9
HELIX   10  10 SER A  268  ARG A  273  5                                   6
HELIX   11  11 TYR A  274  ALA A  282  1                                   9
HELIX   12  12 ILE A  303  TYR A  306  5                                   4
HELIX   13  13 ALA A  307  LEU A  322  1                                  16
HELIX   14  14 ASP A  333  TYR A  346  1                                  14
HELIX   15  15 MET A  370  ILE A  376  1                                   7
HELIX   16  16 PHE A  379  GLU A  387  1                                   9
HELIX   17  17 GLY A  389  LYS A  397  1                                   9
HELIX   18  18 ASN A  398  PHE A  407  1                                  10
HELIX   19  19 LYS A  420  GLY A  425  1                                   6
HELIX   20  20 THR A  442  GLY A  457  1                                  16
HELIX   21  21 PHE A  458  ASN A  463  1                                   6
HELIX   22  22 ASN A  467  CYS A  476  1                                  10
HELIX   23  23 LYS A  477  GLY A  480  5                                   4
HELIX   24  24 ARG A  499  SER A  503  5                                   5
HELIX   25  25 ASN A  505  TRP A  509  5                                   5
HELIX   26  26 TRP A  524  LYS A  529  1                                   6
HELIX   27  27 LYS A  529  THR A  543  1                                  15
HELIX   28  28 SER B   18  LEU B   30  1                                  13
HELIX   29  29 PHE B   36  TYR B   41  5                                   6
HELIX   30  30 PRO B   49  MET B   54  1                                   6
HELIX   31  31 THR B   59  ALA B   77  1                                  19
HELIX   32  32 ILE B   88  ALA B   98  1                                  11
HELIX   33  33 ASN B  102  GLY B  116  1                                  15
HELIX   34  34 ARG B  133  SER B  144  1                                  12
HELIX   35  35 GLN B  145  PHE B  147  5                                   3
HELIX   36  36 PRO B  163  LYS B  174  1                                  12
HELIX   37  37 LEU B  190  GLY B  197  1                                   8
HELIX   38  38 SER B  208  VAL B  216  1                                   9
HELIX   39  39 SER B  268  ARG B  273  5                                   6
HELIX   40  40 TYR B  274  ALA B  282  1                                   9
HELIX   41  41 ILE B  303  TYR B  306  5                                   4
HELIX   42  42 ALA B  307  LEU B  322  1                                  16
HELIX   43  43 ASP B  333  TYR B  346  1                                  14
HELIX   44  44 MET B  370  ILE B  376  1                                   7
HELIX   45  45 PHE B  379  GLU B  387  1                                   9
HELIX   46  46 GLY B  389  LYS B  397  1                                   9
HELIX   47  47 ASN B  398  PHE B  407  1                                  10
HELIX   48  48 THR B  442  GLY B  457  1                                  16
HELIX   49  49 PHE B  458  ASN B  463  1                                   6
HELIX   50  50 ASN B  467  CYS B  476  1                                  10
HELIX   51  51 LYS B  477  GLY B  480  5                                   4
HELIX   52  52 ARG B  499  SER B  503  5                                   5
HELIX   53  53 ASN B  505  TRP B  509  5                                   5
HELIX   54  54 TRP B  524  LYS B  529  1                                   6
HELIX   55  55 LYS B  529  THR B  543  1                                  15
SHEET    1   A 5 PHE A 149  LEU A 150  0
SHEET    2   A 5 THR A 118  ILE A 121  1  O  THR A 119   N  PHE A 149
SHEET    3   A 5 VAL A   5  PHE A   8  1  N  ALA A   6   O  THR A 118
SHEET    4   A 5 VAL A 180  LEU A 183  1  O  VAL A 181   N  ALA A   7
SHEET    5   A 5 VAL A 199  ILE A 201  1  N  VAL A 199   O  VAL A 180
SHEET    1   B 2 ALA A  15  LEU A  16  0
SHEET    2   B 2 SER A 100  ILE A 101 -1  N  SER A 100   O  LEU A  16
SHEET    1   C 2 TYR A 239  THR A 241  0
SHEET    2   C 2 ARG A 247  HIS A 249 -1  N  LEU A 248   O  VAL A 240
SHEET    1   D 6 ARG A 285  ILE A 289  0
SHEET    2   D 6 ALA A 258  CYS A 262  1  N  LEU A 259   O  ARG A 285
SHEET    3   D 6 ALA A 327  HIS A 332  1  N  VAL A 328   O  ALA A 258
SHEET    4   D 6 VAL A 350  LEU A 356  1  N  ARG A 351   O  ALA A 327
SHEET    5   D 6 ALA A 487  ALA A 492  1  O  LEU A 488   N  SER A 355
SHEET    6   D 6 LYS A 514  ILE A 518  1  N  LYS A 514   O  ALA A 487
SHEET    1   E 4 PHE B 149  GLU B 152  0
SHEET    2   E 4 THR B 119  THR B 123  1  O  THR B 119   N  PHE B 149
SHEET    3   E 4 ALA B   6  PHE B   8  1  O  ALA B   6   N  CYS B 120
SHEET    4   E 4 VAL B 181  LEU B 183  1  O  VAL B 181   N  ALA B   7
SHEET    1   F 2 ALA B  15  LEU B  16  0
SHEET    2   F 2 SER B 100  ILE B 101 -1  N  SER B 100   O  LEU B  16
SHEET    1   G 2 TYR B 239  THR B 241  0
SHEET    2   G 2 ARG B 247  HIS B 249 -1  N  LEU B 248   O  VAL B 240
SHEET    1   H 6 ARG B 285  ILE B 289  0
SHEET    2   H 6 ALA B 258  CYS B 262  1  N  LEU B 259   O  ARG B 285
SHEET    3   H 6 ALA B 327  HIS B 332  1  N  VAL B 328   O  ALA B 258
SHEET    4   H 6 VAL B 350  LEU B 356  1  N  ARG B 351   O  ALA B 327
SHEET    5   H 6 ALA B 487  ALA B 492  1  O  LEU B 488   N  SER B 355
SHEET    6   H 6 LYS B 514  ILE B 518  1  N  LYS B 514   O  ALA B 487
CISPEP   1 LEU A   16    PRO A   17          0         0.41
CISPEP   2 LYS A  160    PRO A  161          0        -0.06
CISPEP   3 ASN A  231    PRO A  232          0        -0.74
CISPEP   4 GLY A  256    PRO A  257          0        -0.02
CISPEP   5 PHE A  265    PRO A  266          0        -0.61
CISPEP   6 LEU B   16    PRO B   17          0         0.30
CISPEP   7 LYS B  160    PRO B  161          0        -0.05
CISPEP   8 ASN B  231    PRO B  232          0        -0.01
CISPEP   9 GLY B  256    PRO B  257          0         0.07
CISPEP  10 PHE B  265    PRO B  266          0        -0.74
CRYST1  151.900  143.000   60.000  90.00  90.00  90.00 P 21 21 2     8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.006583  0.000000  0.000000        0.00000
SCALE2      0.000000  0.006993  0.000000        0.00000
SCALE3      0.000000  0.000000  0.016667        0.00000
END