longtext: 1HKH-pdb

content
HEADER    HYDROLASE                               10-MAR-03   1HKH
TITLE     UNLIGATED GAMMA LACTAMASE FROM AN AUREOBACTERIUM SPECIES
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: GAMMA LACTAMASE;
COMPND   3 SYNONYM: CO-FACTOR FREE HALOPEROXIDASE (BY IDENTITY);
COMPND   4 CHAIN: A, B;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: AUREOBACTERIUM;
SOURCE   3 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   4 EXPRESSION_SYSTEM_STRAIN: DH5;
SOURCE   5 EXPRESSION_SYSTEM_PLASMID: PTRC99;
SOURCE   6 OTHER_DETAILS: CHIROTECH CULTURE COLLECTION
KEYWDS    ALPHA/BETA HYDROLASE, CO-FACTOR FREE HALOPEROXIDASE,
EXPDTA    X-RAY DIFFRACTION
AUTHOR    K.LINE,M.N.ISUPOV,J.A.LITTLECHILD
REVDAT   1   30-MAR-04 1HKH    0
JRNL        AUTH   K.LINE,M.N.ISUPOV,J.A.LITTLECHILD
JRNL        TITL   THE CRYSTAL STRUCTURE OF GAMMA LACTAMASE FROM
JRNL        TITL 2 AUREOBACTERIUM SPECIOES REVEALS A TETRAHEDRAL
JRNL        TITL 3 INTERMEDIATE IN THE ACTIVE SITE
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION. 1.73 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.73
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 11.0
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NONE
REMARK   3   COMPLETENESS FOR RANGE        (%) : 100
REMARK   3   NUMBER OF REFLECTIONS             : 119247
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : NONE
REMARK   3   R VALUE            (WORKING SET) : 0.1443
REMARK   3   FREE R VALUE                     : 0.1582
REMARK   3   FREE R VALUE TEST SET SIZE   (%) :  2.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 2400
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH           : 1.73
REMARK   3   BIN RESOLUTION RANGE LOW            : 1.774
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 8519
REMARK   3   BIN R VALUE           (WORKING SET) : 0.223
REMARK   3   BIN FREE R VALUE SET COUNT          : 174
REMARK   3   BIN FREE R VALUE                    : 0.221
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4523
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 10
REMARK   3   SOLVENT ATOMS            : 843
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 16.2
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 16.1
REMARK   3    B11 (A**2) : NONE
REMARK   3    B22 (A**2) : NONE
REMARK   3    B33 (A**2) : NONE
REMARK   3    B12 (A**2) : NONE
REMARK   3    B13 (A**2) : NONE
REMARK   3    B23 (A**2) : NONE
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.060
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.059
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.035
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 1.055
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.969
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.960
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES    COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED           (A):  4645 ; 0.013 ; 0.021
REMARK   3   BOND ANGLES REFINED      (DEGREES):  6334 ; 1.346 ; 1.938
REMARK   3   TORSION ANGLES, PERIOD 1 (DEGREES):   556 ; 5.336 ; 5.000
REMARK   3   CHIRAL-CENTER RESTRAINTS    (A**3):   691 ; 0.101 ; 0.200
REMARK   3   GENERAL PLANES REFINED         (A):  3655 ; 0.006 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED    (A):  2331 ; 0.196 ; 0.200
REMARK   3   H-BOND (X...Y) REFINED         (A):   692 ; 0.147 ; 0.200
REMARK   3   SYMMETRY VDW REFINED           (A):    92 ; 0.183 ; 0.200
REMARK   3   SYMMETRY H-BOND REFINED        (A):    57 ; 0.134 ; 0.200
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS. COUNT  RMS   WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED   (A**2):  2791 ; 0.594 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE REFINED  (A**2):  4514 ; 1.105 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED   (A**2):  1854 ; 1.907 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED  (A**2):  1820 ; 3.163 ; 4.500
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.COUNT  RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NONE
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NONE
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED :  BABINET MODEL WITH MAS
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3   RIDING POSITIONS
REMARK   4
REMARK   4 1HKH COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  ON 10-MAR-2003.
REMARK 100 THE EBI ID CODE IS EBI-12293.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 15-JUN-2001
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : EMBL/DESY, HAMBURG BEAMLINE X11
REMARK 200  BEAMLINE                       : X11
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.8482
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 119247
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.73
REMARK 200  RESOLUTION RANGE LOW       (A) : 11.0
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NONE
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100
REMARK 200  DATA REDUNDANCY                : 11.4
REMARK 200  R MERGE                    (I) : 0.108
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 24.76
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.73
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.76
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100
REMARK 200  DATA REDUNDANCY IN SHELL       : 11
REMARK 200  R MERGE FOR SHELL          (I) : 0.48
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 5.23
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: PDB ENTRY 1BRO
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 73.73
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 4.72
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: F 2 3
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   -X,Y,-Z
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   Z,X,Y
REMARK 290       6555   Z,-X,-Y
REMARK 290       7555   -Z,-X,Y
REMARK 290       8555   -Z,X,-Y
REMARK 290       9555   Y,Z,X
REMARK 290      10555   -Y,Z,-X
REMARK 290      11555   Y,-Z,-X
REMARK 290      12555   -Y,-Z,X
REMARK 290      13555   X,1/2+Y,1/2+Z
REMARK 290      14555   -X,1/2-Y,1/2+Z
REMARK 290      15555   -X,1/2+Y,1/2-Z
REMARK 290      16555   X,1/2-Y,1/2-Z
REMARK 290      17555   Z,1/2+X,1/2+Y
REMARK 290      18555   Z,1/2-X,1/2-Y
REMARK 290      19555   -Z,1/2-X,1/2+Y
REMARK 290      20555   -Z,1/2+X,1/2-Y
REMARK 290      21555   Y,1/2+Z,1/2+X
REMARK 290      22555   -Y,1/2+Z,1/2-X
REMARK 290      23555   Y,1/2-Z,1/2-X
REMARK 290      24555   -Y,1/2-Z,1/2+X
REMARK 290      25555   1/2+X,Y,1/2+Z
REMARK 290      26555   1/2-X,-Y,1/2+Z
REMARK 290      27555   1/2-X,Y,1/2-Z
REMARK 290      28555   1/2+X,-Y,1/2-Z
REMARK 290      29555   1/2+Z,X,1/2+Y
REMARK 290      30555   1/2+Z,-X,1/2-Y
REMARK 290      31555   1/2-Z,-X,1/2+Y
REMARK 290      32555   1/2-Z,X,1/2-Y
REMARK 290      33555   1/2+Y,Z,1/2+X
REMARK 290      34555   1/2-Y,Z,1/2-X
REMARK 290      35555   1/2+Y,-Z,1/2-X
REMARK 290      36555   1/2-Y,-Z,1/2+X
REMARK 290      37555   1/2+X,1/2+Y,Z
REMARK 290      38555   1/2-X,1/2-Y,Z
REMARK 290      39555   1/2-X,1/2+Y,-Z
REMARK 290      40555   1/2+X,1/2-Y,-Z
REMARK 290      41555   1/2+Z,1/2+X,Y
REMARK 290      42555   1/2+Z,1/2-X,-Y
REMARK 290      43555   1/2-Z,1/2-X,Y
REMARK 290      44555   1/2-Z,1/2+X,-Y
REMARK 290      45555   1/2+Y,1/2+Z,X
REMARK 290      46555   1/2-Y,1/2+Z,-X
REMARK 290      47555   1/2+Y,1/2-Z,-X
REMARK 290      48555   1/2-Y,1/2-Z,X
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1   6  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2   6 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2   7 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2   8  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1   9  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   9  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3   9  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  10  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  10  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3  10 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  11  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2  11  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  11 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  12  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  12  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  12  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  13  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  13  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY3  13  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY1  14 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  14  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY3  14  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY1  15 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  15  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY3  15  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY1  16  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  16  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY3  16  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY1  17  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2  17  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  17  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY1  18  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2  18 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  18  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY1  19  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2  19 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  19  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY1  20  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2  20  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  20  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY1  21  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2  21  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY3  21  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  22  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  22  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY3  22 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  23  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2  23  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY3  23 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  24  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  24  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY3  24  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  25  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  25  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3  25  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY1  26 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  26  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3  26  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY1  27 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  27  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3  27  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY1  28  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  28  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3  28  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY1  29  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY2  29  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  29  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY1  30  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY2  30 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  30  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY1  31  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY2  31 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  31  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY1  32  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY2  32  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  32  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY1  33  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY2  33  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3  33  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  34  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY2  34  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3  34 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  35  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY2  35  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  35 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  36  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY2  36  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  36  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY1  37  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  37  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY3  37  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1  38 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  38  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY3  38  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1  39 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  39  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY3  39  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1  40  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY2  40  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY3  40  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1  41  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY2  41  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  41  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1  42  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY2  42 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  42  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1  43  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY2  43 -1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  43  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1  44  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY2  44  1.000000  0.000000  0.000000      120.43700
REMARK 290   SMTRY3  44  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1  45  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY2  45  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY3  45  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  46  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY2  46  0.000000  0.000000  1.000000      120.43700
REMARK 290   SMTRY3  46 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  47  0.000000  1.000000  0.000000      120.43700
REMARK 290   SMTRY2  47  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY3  47 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  48  0.000000 -1.000000  0.000000      120.43700
REMARK 290   SMTRY2  48  0.000000  0.000000 -1.000000      120.43700
REMARK 290   SMTRY3  48  1.000000  0.000000  0.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 295
REMARK 295 NON-CRYSTALLOGRAPHIC SYMMETRY
REMARK 295 THE TRANSFORMATIONS PRESENTED ON THE MTRIX RECORDS BELOW
REMARK 295 DESCRIBE NON-CRYSTALLOGRAPHIC RELATIONSHIPS AMONG ATOMS
REMARK 295 IN THIS ENTRY.  APPLYING THE APPROPRIATE MTRIX
REMARK 295 TRANSFORMATION TO THE RESIDUES LISTED FIRST WILL YIELD
REMARK 295 APPROXIMATE COORDINATES FOR THE RESIDUES LISTED SECOND.
REMARK 295 CHAIN IDENTIFIERS GIVEN AS "?" REFER TO CHAINS FOR WHICH
REMARK 295 ATOMS ARE NOT FOUND IN THIS ENTRY.
REMARK 295
REMARK 295               APPLIED TO          TRANSFORMED TO
REMARK 295   TRANSFORM CHAIN  RESIDUES       CHAIN  RESIDUES     RMSD
REMARK 295     SSS
REMARK 295    M  1       A    1 .. 279         B    1 .. 279     0.219
REMARK 295
REMARK 295    WHERE SSS -> COLUMNS 8-10 OF MTRIX RECORDS
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT
REMARK 300 WHICH CONSISTS OF   2 CHAIN(S). SEE REMARK 350 FOR
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).
REMARK 300
REMARK 300 QUATERNARY STRUCTURE FOR THIS ENTRY: TRIMERIC
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE:  1
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT2   2 -1.000000  0.000000  0.000000      120.43700
REMARK 350   BIOMT3   2  0.000000 -1.000000  0.000000      120.43700
REMARK 350   BIOMT1   3  0.000000 -1.000000  0.000000      120.43700
REMARK 350   BIOMT2   3  0.000000  0.000000 -1.000000      120.43700
REMARK 350   BIOMT3   3  1.000000  0.000000  0.000000        0.00000
REMARK 350 BIOMOLECULE:  2
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  0.000000 -1.000000  0.000000      120.43700
REMARK 350   BIOMT2   2  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT3   2 -1.000000  0.000000  0.000000      120.43700
REMARK 350   BIOMT1   3  0.000000  0.000000 -1.000000      120.43700
REMARK 350   BIOMT2   3 -1.000000  0.000000  0.000000      120.43700
REMARK 350   BIOMT3   3  0.000000  1.000000  0.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375      HOH Y   9  LIES ON A SPECIAL POSITION.
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500
REMARK 500   O    HOH Y     7     O    HOH Y    39               2.13
REMARK 500   O    HOH Y    79     O    HOH Y   275               2.10
REMARK 500   O    HOH Y    80     O    HOH Y   276               2.20
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500
REMARK 500    SER A  98     -115.92     53.29
REMARK 500    SER B  98     -116.39     53.53
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES ARE GIVEN CHAIN IDENTIFIERS TO
REMARK 525 INDICATE THE PROTEIN CHAIN TO WHICH THEY ARE MOST CLOSELY
REMARK 525 ASSOCIATED WITH:
REMARK 525   PROTEIN CHAIN  SOLVENT CHAIN
REMARK 525     A              Z
REMARK 525     B              Y
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: CA
REMARK 800 SITE_DESCRIPTION: CATALYTIC TRIAD, CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: CB
REMARK 800 SITE_DESCRIPTION: CATALYTIC TRIAD, CHAIN B
REMARK 800
REMARK 800 SITE_IDENTIFIER: SOA
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: SOB
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN B
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 NAMED CO-FACTOR FREE HALOPEROXIDASE DUE TO HIGH SEQUENCE ID
DBREF  1HKH A    1   279  SWS    Q8GJP7   Q8GJP7           2    280
DBREF  1HKH B    1   279  SWS    Q8GJP7   Q8GJP7           2    280
SEQRES   1 A  279  GLY TYR ILE THR VAL GLY ASN GLU ASN SER THR PRO ILE
SEQRES   2 A  279  GLU LEU TYR TYR GLU ASP GLN GLY SER GLY GLN PRO VAL
SEQRES   3 A  279  VAL LEU ILE HIS GLY TYR PRO LEU ASP GLY HIS SER TRP
SEQRES   4 A  279  GLU ARG GLN THR ARG GLU LEU LEU ALA GLN GLY TYR ARG
SEQRES   5 A  279  VAL ILE THR TYR ASP ARG ARG GLY PHE GLY GLY SER SER
SEQRES   6 A  279  LYS VAL ASN THR GLY TYR ASP TYR ASP THR PHE ALA ALA
SEQRES   7 A  279  ASP LEU HIS THR VAL LEU GLU THR LEU ASP LEU ARG ASP
SEQRES   8 A  279  VAL VAL LEU VAL GLY PHE SER MET GLY THR GLY GLU LEU
SEQRES   9 A  279  ALA ARG TYR VAL ALA ARG TYR GLY HIS GLU ARG VAL ALA
SEQRES  10 A  279  LYS LEU ALA PHE LEU ALA SER LEU GLU PRO PHE LEU VAL
SEQRES  11 A  279  GLN ARG ASP ASP ASN PRO GLU GLY VAL PRO GLN GLU VAL
SEQRES  12 A  279  PHE ASP GLY ILE GLU ALA ALA ALA LYS GLY ASP ARG PHE
SEQRES  13 A  279  ALA TRP PHE THR ASP PHE TYR LYS ASN PHE TYR ASN LEU
SEQRES  14 A  279  ASP GLU ASN LEU GLY SER ARG ILE SER GLU GLN ALA VAL
SEQRES  15 A  279  THR GLY SER TRP ASN VAL ALA ILE GLY SER ALA PRO VAL
SEQRES  16 A  279  ALA ALA TYR ALA VAL VAL PRO ALA TRP ILE GLU ASP PHE
SEQRES  17 A  279  ARG SER ASP VAL GLU ALA VAL ARG ALA ALA GLY LYS PRO
SEQRES  18 A  279  THR LEU ILE LEU HIS GLY THR LYS ASP ASN ILE LEU PRO
SEQRES  19 A  279  ILE ASP ALA THR ALA ARG ARG PHE HIS GLN ALA VAL PRO
SEQRES  20 A  279  GLU ALA ASP TYR VAL GLU VAL GLU GLY ALA PRO HIS GLY
SEQRES  21 A  279  LEU LEU TRP THR HIS ALA ASP GLU VAL ASN ALA ALA LEU
SEQRES  22 A  279  LYS THR PHE LEU ALA LYS
SEQRES   1 B  279  GLY TYR ILE THR VAL GLY ASN GLU ASN SER THR PRO ILE
SEQRES   2 B  279  GLU LEU TYR TYR GLU ASP GLN GLY SER GLY GLN PRO VAL
SEQRES   3 B  279  VAL LEU ILE HIS GLY TYR PRO LEU ASP GLY HIS SER TRP
SEQRES   4 B  279  GLU ARG GLN THR ARG GLU LEU LEU ALA GLN GLY TYR ARG
SEQRES   5 B  279  VAL ILE THR TYR ASP ARG ARG GLY PHE GLY GLY SER SER
SEQRES   6 B  279  LYS VAL ASN THR GLY TYR ASP TYR ASP THR PHE ALA ALA
SEQRES   7 B  279  ASP LEU HIS THR VAL LEU GLU THR LEU ASP LEU ARG ASP
SEQRES   8 B  279  VAL VAL LEU VAL GLY PHE SER MET GLY THR GLY GLU LEU
SEQRES   9 B  279  ALA ARG TYR VAL ALA ARG TYR GLY HIS GLU ARG VAL ALA
SEQRES  10 B  279  LYS LEU ALA PHE LEU ALA SER LEU GLU PRO PHE LEU VAL
SEQRES  11 B  279  GLN ARG ASP ASP ASN PRO GLU GLY VAL PRO GLN GLU VAL
SEQRES  12 B  279  PHE ASP GLY ILE GLU ALA ALA ALA LYS GLY ASP ARG PHE
SEQRES  13 B  279  ALA TRP PHE THR ASP PHE TYR LYS ASN PHE TYR ASN LEU
SEQRES  14 B  279  ASP GLU ASN LEU GLY SER ARG ILE SER GLU GLN ALA VAL
SEQRES  15 B  279  THR GLY SER TRP ASN VAL ALA ILE GLY SER ALA PRO VAL
SEQRES  16 B  279  ALA ALA TYR ALA VAL VAL PRO ALA TRP ILE GLU ASP PHE
SEQRES  17 B  279  ARG SER ASP VAL GLU ALA VAL ARG ALA ALA GLY LYS PRO
SEQRES  18 B  279  THR LEU ILE LEU HIS GLY THR LYS ASP ASN ILE LEU PRO
SEQRES  19 B  279  ILE ASP ALA THR ALA ARG ARG PHE HIS GLN ALA VAL PRO
SEQRES  20 B  279  GLU ALA ASP TYR VAL GLU VAL GLU GLY ALA PRO HIS GLY
SEQRES  21 B  279  LEU LEU TRP THR HIS ALA ASP GLU VAL ASN ALA ALA LEU
SEQRES  22 B  279  LYS THR PHE LEU ALA LYS
HET    SO4  A1280       5
HET    SO4  B1280       5
HETNAM     SO4 SULFATE ION
FORMUL   3  SO4    2(O4 S1 2-)
FORMUL   4  HOH   *843(H2 O1)
HELIX    1   1 ASP A   35  SER A   38  5                                   4
HELIX    2   2 TRP A   39  GLN A   49  1                                  11
HELIX    3   3 ASP A   72  ASP A   88  1                                  17
HELIX    4   4 SER A   98  GLY A  112  1                                  15
HELIX    5   5 PRO A  140  ASP A  154  1                                  15
HELIX    6   6 ASP A  154  ASN A  168  1                                  15
HELIX    7   7 ASN A  168  LEU A  173  1                                   6
HELIX    8   8 SER A  178  GLY A  191  1                                  14
HELIX    9   9 VAL A  195  VAL A  200  1                                   6
HELIX   10  10 VAL A  200  ILE A  205  1                                   6
HELIX   11  11 PHE A  208  GLY A  219  1                                  12
HELIX   12  12 THR A  238  VAL A  246  1                                   9
HELIX   13  13 GLY A  260  HIS A  265  1                                   6
HELIX   14  14 HIS A  265  LYS A  279  1                                  15
HELIX   15  15 ASP B   35  SER B   38  5                                   4
HELIX   16  16 TRP B   39  GLN B   49  1                                  11
HELIX   17  17 ASP B   72  ASP B   88  1                                  17
HELIX   18  18 SER B   98  GLY B  112  1                                  15
HELIX   19  19 PRO B  140  ASP B  154  1                                  15
HELIX   20  20 ASP B  154  ASN B  168  1                                  15
HELIX   21  21 ASN B  168  LEU B  173  1                                   6
HELIX   22  22 SER B  178  GLY B  191  1                                  14
HELIX   23  23 VAL B  195  VAL B  200  1                                   6
HELIX   24  24 VAL B  200  ILE B  205  1                                   6
HELIX   25  25 PHE B  208  GLY B  219  1                                  12
HELIX   26  26 THR B  238  VAL B  246  1                                   9
HELIX   27  27 GLY B  260  HIS B  265  1                                   6
HELIX   28  28 HIS B  265  LYS B  279  1                                  15
SHEET    1  AA 8 TYR A   2  GLU A   8  0
SHEET    2  AA 8 THR A  11  GLN A  20 -1  O  THR A  11   N  GLU A   8
SHEET    3  AA 8 TYR A  51  TYR A  56 -1  O  VAL A  53   N  GLN A  20
SHEET    4  AA 8 GLN A  24  ILE A  29  1  O  GLN A  24   N  ARG A  52
SHEET    5  AA 8 VAL A  92  PHE A  97  1  O  VAL A  93   N  VAL A  27
SHEET    6  AA 8 VAL A 116  LEU A 122  1  N  ALA A 117   O  VAL A  92
SHEET    7  AA 8 THR A 222  GLY A 227  1  O  LEU A 223   N  PHE A 121
SHEET    8  AA 8 ASP A 250  VAL A 254  1  O  ASP A 250   N  ILE A 224
SHEET    1  BA 8 TYR B   2  GLU B   8  0
SHEET    2  BA 8 THR B  11  GLY B  21 -1  O  THR B  11   N  GLU B   8
SHEET    3  BA 8 TYR B  51  TYR B  56 -1  O  VAL B  53   N  GLN B  20
SHEET    4  BA 8 GLN B  24  ILE B  29  1  O  GLN B  24   N  ARG B  52
SHEET    5  BA 8 VAL B  92  PHE B  97  1  O  VAL B  93   N  VAL B  27
SHEET    6  BA 8 VAL B 116  LEU B 122  1  N  ALA B 117   O  VAL B  92
SHEET    7  BA 8 THR B 222  GLY B 227  1  O  LEU B 223   N  PHE B 121
SHEET    8  BA 8 ASP B 250  VAL B 254  1  O  ASP B 250   N  ILE B 224
CISPEP   1 TYR A   32    PRO A   33          0        -5.79
CISPEP   2 GLU A  126    PRO A  127          0         1.99
CISPEP   3 TYR B   32    PRO B   33          0        -6.38
CISPEP   4 GLU B  126    PRO B  127          0         1.47
SITE     1  CA  3 SER A  98  ASP A 230  HIS A 259
SITE     1  CB  3 SER B  98  ASP B 230  HIS B 259
SITE     1 SOA  3 ARG A 176  HOH Z 299  HOH Z 428
SITE     1 SOB  2 ARG B 176  HOH Y 286
CRYST1  240.874  240.874  240.874  90.00  90.00  90.00 F 2 3        96
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.004151  0.000000  0.000000        0.00000
SCALE2      0.000000  0.004151  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004151        0.00000
MTRIX1   1  0.327130  0.779240 -0.534570       99.61824    1
MTRIX2   1  0.539790  0.310230  0.782550     -112.95067    1
MTRIX3   1  0.775640 -0.544550 -0.319140       -8.86623    1
TER    2254      LYS A 279
TER    4525      LYS B 279
MASTER      471    0    2   28   16    0    4    9 5376    2   10   44
END