longtext: 1HLG-pdb

content
HEADER    HYDROLASE                               12-MAR-99   1HLG
TITLE     CRYSTAL STRUCTURE OF HUMAN GASTRIC LIPASE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE, GASTRIC;
COMPND   3 CHAIN: A, B;
COMPND   4 EC: 3.1.1.3;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGAN: STOMAC;
SOURCE   5 CELL: CHIEF CELL;
SOURCE   6 EXPRESSION_SYSTEM: INSECT CELL;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: VIRUS;
SOURCE   8 EXPRESSION_SYSTEM_VECTOR: BACULOVIRUS
KEYWDS    LIPASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    A.ROUSSEL,S.CANAAN,R.VERGER,C.CAMBILLAU
REVDAT   1   15-MAR-00 1HLG    0
JRNL        AUTH   A.ROUSSEL,S.CANAAN,M.P.EGLOFF,M.RIVIERE,L.DUPUIS,
JRNL        AUTH 2 R.VERGER,C.CAMBILLAU
JRNL        TITL   CRYSTAL STRUCTURE OF HUMAN GASTRIC LIPASE AND
JRNL        TITL 2 MODEL OF LYSOSOMAL ACID LIPASE, TWO LIPOLYTIC
JRNL        TITL 3 ENZYMES OF MEDICAL INTEREST
JRNL        REF    J.BIOL.CHEM.                  V. 274 16995 1999
JRNL        REFN   ASTM JBCHA3  US ISSN 0021-9258
REMARK   1
REMARK   2
REMARK   2 RESOLUTION. 3.00 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : X-PLOR 3843
REMARK   3   AUTHORS     : BRUNGER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 3.00
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 15.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 99.4
REMARK   3   NUMBER OF REFLECTIONS             : 47643
REMARK   3
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.210
REMARK   3   FREE R VALUE                     : 0.228
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 2381
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.005
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 3.00
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 3.19
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 96.80
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 7293
REMARK   3   BIN R VALUE           (WORKING SET) : 0.3190
REMARK   3   BIN FREE R VALUE                    : 0.3480
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 5.10
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 389
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.018
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 5960
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 140
REMARK   3   SOLVENT ATOMS            : 92
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 49.60
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.35
REMARK   3   ESD FROM SIGMAA              (A) : 0.510
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.39
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.58
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.007
REMARK   3   BOND ANGLES            (DEGREES) : 1.30
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 24.20
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.90
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3
REMARK   3  NCS MODEL : RESTRAINTS
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PARHCSDX.PRO
REMARK   3  PARAMETER FILE  2  : PARAM19X.PRO
REMARK   3  PARAMETER FILE  3  : PARAM3.CHO
REMARK   3  TOPOLOGY FILE  1   : TOPHCSDX.PRO
REMARK   3  TOPOLOGY FILE  2   : TOPH19X.PRO
REMARK   3  TOPOLOGY FILE  3   : TOPH3.CHO
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 1HLG COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 16-MAR-1999.
REMARK 100 THE RCSB ID CODE IS RCSB000645.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : SEP-1998
REMARK 200  TEMPERATURE           (KELVIN) : 100.0
REMARK 200  PH                             : 5.00
REMARK 200  NUMBER OF CRYSTALS USED        : 3
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : LURE
REMARK 200  BEAMLINE                       : W32
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.963
REMARK 200  MONOCHROMATOR                  : LURE
REMARK 200  OPTICS                         : LURE
REMARK 200
REMARK 200  DETECTOR TYPE                  : MAR 345
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 45522
REMARK 200  RESOLUTION RANGE HIGH      (A) : 3.000
REMARK 200  RESOLUTION RANGE LOW       (A) : 15.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.6
REMARK 200  DATA REDUNDANCY                : 4.000
REMARK 200  R MERGE                    (I) : 0.06800
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.9000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 3.00
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 3.10
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 4.10
REMARK 200  R MERGE FOR SHELL          (I) : 0.39900
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.000
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SIRAS
REMARK 200 SOFTWARE USED: CCP4 PACKAGE
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 81.0
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 6.35
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: I 21 3
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   1/2-X,-Y,1/2+Z
REMARK 290       3555   -X,1/2+Y,1/2-Z
REMARK 290       4555   1/2+X,1/2-Y,-Z
REMARK 290       5555   Z,X,Y
REMARK 290       6555   1/2+Z,1/2-X,-Y
REMARK 290       7555   1/2-Z,-X,1/2+Y
REMARK 290       8555   -Z,1/2+X,1/2-Y
REMARK 290       9555   Y,Z,X
REMARK 290      10555   -Y,1/2+Z,1/2-X
REMARK 290      11555   1/2+Y,1/2-Z,-X
REMARK 290      12555   1/2-Y,-Z,1/2+X
REMARK 290      13555   1/2+X,1/2+Y,1/2+Z
REMARK 290      14555   -X,1/2-Y,Z
REMARK 290      15555   1/2-X,Y,-Z
REMARK 290      16555   X,-Y,1/2-Z
REMARK 290      17555   1/2+Z,1/2+X,1/2+Y
REMARK 290      18555   Z,-X,1/2-Y
REMARK 290      19555   -Z,1/2-X,Y
REMARK 290      20555   1/2-Z,X,-Y
REMARK 290      21555   1/2+Y,1/2+Z,1/2+X
REMARK 290      22555   1/2-Y,Z,-X
REMARK 290      23555   Y,-Z,1/2-X
REMARK 290      24555   -Y,1/2-Z,X
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      122.16500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000      122.16500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      122.16500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000      122.16500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1   6  0.000000  0.000000  1.000000      122.16500
REMARK 290   SMTRY2   6 -1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY3   6  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1   7  0.000000  0.000000 -1.000000      122.16500
REMARK 290   SMTRY2   7 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  1.000000  0.000000      122.16500
REMARK 290   SMTRY1   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2   8  1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY3   8  0.000000 -1.000000  0.000000      122.16500
REMARK 290   SMTRY1   9  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   9  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3   9  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  10  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  10  0.000000  0.000000  1.000000      122.16500
REMARK 290   SMTRY3  10 -1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY1  11  0.000000  1.000000  0.000000      122.16500
REMARK 290   SMTRY2  11  0.000000  0.000000 -1.000000      122.16500
REMARK 290   SMTRY3  11 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  12  0.000000 -1.000000  0.000000      122.16500
REMARK 290   SMTRY2  12  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  12  1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY1  13  1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY2  13  0.000000  1.000000  0.000000      122.16500
REMARK 290   SMTRY3  13  0.000000  0.000000  1.000000      122.16500
REMARK 290   SMTRY1  14 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  14  0.000000 -1.000000  0.000000      122.16500
REMARK 290   SMTRY3  14  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1  15 -1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY2  15  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3  15  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1  16  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  16  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3  16  0.000000  0.000000 -1.000000      122.16500
REMARK 290   SMTRY1  17  0.000000  0.000000  1.000000      122.16500
REMARK 290   SMTRY2  17  1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY3  17  0.000000  1.000000  0.000000      122.16500
REMARK 290   SMTRY1  18  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2  18 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  18  0.000000 -1.000000  0.000000      122.16500
REMARK 290   SMTRY1  19  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2  19 -1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY3  19  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1  20  0.000000  0.000000 -1.000000      122.16500
REMARK 290   SMTRY2  20  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  20  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1  21  0.000000  1.000000  0.000000      122.16500
REMARK 290   SMTRY2  21  0.000000  0.000000  1.000000      122.16500
REMARK 290   SMTRY3  21  1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY1  22  0.000000 -1.000000  0.000000      122.16500
REMARK 290   SMTRY2  22  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3  22 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  23  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2  23  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  23 -1.000000  0.000000  0.000000      122.16500
REMARK 290   SMTRY1  24  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  24  0.000000  0.000000 -1.000000      122.16500
REMARK 290   SMTRY3  24  1.000000  0.000000  0.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT
REMARK 300 WHICH CONSISTS OF 2 CHAIN(S). SEE REMARK 350 FOR
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ASN A    54
REMARK 465     THR A    55
REMARK 465     GLY A    56
REMARK 465     ASN B    54
REMARK 465     THR B    55
REMARK 465     GLY B    56
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI
REMARK 500   O    HOH      46     O    HOH     146              2.17
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    ASN A  51   OD1   ASN A  51   CG     0.128
REMARK 500    ASN A  51   ND2   ASN A  51   CG     0.107
REMARK 500    SER A  52   OG    SER A  52   CB     0.112
REMARK 500    ARG A 229   NH1   ARG A 229   CZ     0.113
REMARK 500    MET A 297   CE    MET A 297   SD     0.128
REMARK 500    MET A 360   CE    MET A 360   SD     0.146
REMARK 500    LYS B  50   CG    LYS B  50   CB     0.188
REMARK 500    LYS B  50   CD    LYS B  50   CG     0.228
REMARK 500    LYS B  50   CE    LYS B  50   CD     0.180
REMARK 500    LYS B  50   NZ    LYS B  50   CE     0.180
REMARK 500    LYS B  50   O     LYS B  50   C      0.147
REMARK 500    ASN B  51   CG    ASN B  51   CB     0.176
REMARK 500    ASN B  51   OD1   ASN B  51   CG     0.128
REMARK 500    MET B 231   SD    MET B 231   CG     0.158
REMARK 500    MET B 231   CE    MET B 231   SD     0.154
REMARK 500    ASP B 377   OD1   ASP B 377   CG     0.165
REMARK 500    ASP B 377   OD2   ASP B 377   CG     0.161
REMARK 500    LYS B 378   CG    LYS B 378   CB     0.118
REMARK 500    LYS B 378   CD    LYS B 378   CG     0.141
REMARK 500    LYS B 378   O     LYS B 378   C      0.140
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    GLY A  37   N   -  CA  -  C   ANGL. DEV. = 12.7 DEGREES
REMARK 500    LYS A  50   N   -  CA  -  C   ANGL. DEV. =-10.8 DEGREES
REMARK 500    SER A  76   N   -  CA  -  C   ANGL. DEV. = 11.4 DEGREES
REMARK 500    TYR A 110   N   -  CA  -  C   ANGL. DEV. = 10.4 DEGREES
REMARK 500    ASP A 209   N   -  CA  -  C   ANGL. DEV. =-11.2 DEGREES
REMARK 500    ASN A 252   N   -  CA  -  C   ANGL. DEV. =-11.2 DEGREES
REMARK 500    GLN A 301   N   -  CA  -  C   ANGL. DEV. =-10.6 DEGREES
REMARK 500    GLY B  37   N   -  CA  -  C   ANGL. DEV. = 12.5 DEGREES
REMARK 500    LYS B  49   CD  -  CE  -  NZ  ANGL. DEV. = 13.9 DEGREES
REMARK 500    LYS B  50   CD  -  CE  -  NZ  ANGL. DEV. = 11.2 DEGREES
REMARK 500    LYS B  50   N   -  CA  -  C   ANGL. DEV. =-12.8 DEGREES
REMARK 500    SER B  76   N   -  CA  -  C   ANGL. DEV. = 11.7 DEGREES
REMARK 500    ASP B 209   N   -  CA  -  C   ANGL. DEV. =-10.1 DEGREES
REMARK 500    ASN B 252   N   -  CA  -  C   ANGL. DEV. =-11.7 DEGREES
REMARK 500    GLN B 301   N   -  CA  -  C   ANGL. DEV. =-11.1 DEGREES
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: SA
REMARK 800 SITE_DESCRIPTION:
REMARK 800 ACTIVE SITE CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: SB
REMARK 800 SITE_DESCRIPTION:
REMARK 800 ACTIVE SITE CHAIN B
REMARK 800
DBREF  1HLG A    9   379  SWS    P07098   LIPG_HUMAN      28    398
DBREF  1HLG B    9   379  SWS    P07098   LIPG_HUMAN      28    398
SEQRES   1 A  371  SER PRO GLU VAL THR MET ASN ILE SER GLN MET ILE THR
SEQRES   2 A  371  TYR TRP GLY TYR PRO ASN GLU GLU TYR GLU VAL VAL THR
SEQRES   3 A  371  GLU ASP GLY TYR ILE LEU GLU VAL ASN ARG ILE PRO TYR
SEQRES   4 A  371  GLY LYS LYS ASN SER GLY ASN THR GLY GLN ARG PRO VAL
SEQRES   5 A  371  VAL PHE LEU GLN HIS GLY LEU LEU ALA SER ALA THR ASN
SEQRES   6 A  371  TRP ILE SER ASN LEU PRO ASN ASN SER LEU ALA PHE ILE
SEQRES   7 A  371  LEU ALA ASP ALA GLY TYR ASP VAL TRP LEU GLY ASN SER
SEQRES   8 A  371  ARG GLY ASN THR TRP ALA ARG ARG ASN LEU TYR TYR SER
SEQRES   9 A  371  PRO ASP SER VAL GLU PHE TRP ALA PHE SER PHE ASP GLU
SEQRES  10 A  371  MET ALA LYS TYR ASP LEU PRO ALA THR ILE ASP PHE ILE
SEQRES  11 A  371  VAL LYS LYS THR GLY GLN LYS GLN LEU HIS TYR VAL GLY
SEQRES  12 A  371  HIS SER GLN GLY THR THR ILE GLY PHE ILE ALA PHE SER
SEQRES  13 A  371  THR ASN PRO SER LEU ALA LYS ARG ILE LYS THR PHE TYR
SEQRES  14 A  371  ALA LEU ALA PRO VAL ALA THR VAL LYS TYR THR LYS SER
SEQRES  15 A  371  LEU ILE ASN LYS LEU ARG PHE VAL PRO GLN SER LEU PHE
SEQRES  16 A  371  LYS PHE ILE PHE GLY ASP LYS ILE PHE TYR PRO HIS ASN
SEQRES  17 A  371  PHE PHE ASP GLN PHE LEU ALA THR GLU VAL CYS SER ARG
SEQRES  18 A  371  GLU MET LEU ASN LEU LEU CYS SER ASN ALA LEU PHE ILE
SEQRES  19 A  371  ILE CYS GLY PHE ASP SER LYS ASN PHE ASN THR SER ARG
SEQRES  20 A  371  LEU ASP VAL TYR LEU SER HIS ASN PRO ALA GLY THR SER
SEQRES  21 A  371  VAL GLN ASN MET PHE HIS TRP THR GLN ALA VAL LYS SER
SEQRES  22 A  371  GLY LYS PHE GLN ALA TYR ASP TRP GLY SER PRO VAL GLN
SEQRES  23 A  371  ASN ARG MET HIS TYR ASP GLN SER GLN PRO PRO TYR TYR
SEQRES  24 A  371  ASN VAL THR ALA MET ASN VAL PRO ILE ALA VAL TRP ASN
SEQRES  25 A  371  GLY GLY LYS ASP LEU LEU ALA ASP PRO GLN ASP VAL GLY
SEQRES  26 A  371  LEU LEU LEU PRO LYS LEU PRO ASN LEU ILE TYR HIS LYS
SEQRES  27 A  371  GLU ILE PRO PHE TYR ASN HIS LEU ASP PHE ILE TRP ALA
SEQRES  28 A  371  MET ASP ALA PRO GLN GLU VAL TYR ASN ASP ILE VAL SER
SEQRES  29 A  371  MET ILE SER GLU ASP LYS LYS
SEQRES   1 B  371  SER PRO GLU VAL THR MET ASN ILE SER GLN MET ILE THR
SEQRES   2 B  371  TYR TRP GLY TYR PRO ASN GLU GLU TYR GLU VAL VAL THR
SEQRES   3 B  371  GLU ASP GLY TYR ILE LEU GLU VAL ASN ARG ILE PRO TYR
SEQRES   4 B  371  GLY LYS LYS ASN SER GLY ASN THR GLY GLN ARG PRO VAL
SEQRES   5 B  371  VAL PHE LEU GLN HIS GLY LEU LEU ALA SER ALA THR ASN
SEQRES   6 B  371  TRP ILE SER ASN LEU PRO ASN ASN SER LEU ALA PHE ILE
SEQRES   7 B  371  LEU ALA ASP ALA GLY TYR ASP VAL TRP LEU GLY ASN SER
SEQRES   8 B  371  ARG GLY ASN THR TRP ALA ARG ARG ASN LEU TYR TYR SER
SEQRES   9 B  371  PRO ASP SER VAL GLU PHE TRP ALA PHE SER PHE ASP GLU
SEQRES  10 B  371  MET ALA LYS TYR ASP LEU PRO ALA THR ILE ASP PHE ILE
SEQRES  11 B  371  VAL LYS LYS THR GLY GLN LYS GLN LEU HIS TYR VAL GLY
SEQRES  12 B  371  HIS SER GLN GLY THR THR ILE GLY PHE ILE ALA PHE SER
SEQRES  13 B  371  THR ASN PRO SER LEU ALA LYS ARG ILE LYS THR PHE TYR
SEQRES  14 B  371  ALA LEU ALA PRO VAL ALA THR VAL LYS TYR THR LYS SER
SEQRES  15 B  371  LEU ILE ASN LYS LEU ARG PHE VAL PRO GLN SER LEU PHE
SEQRES  16 B  371  LYS PHE ILE PHE GLY ASP LYS ILE PHE TYR PRO HIS ASN
SEQRES  17 B  371  PHE PHE ASP GLN PHE LEU ALA THR GLU VAL CYS SER ARG
SEQRES  18 B  371  GLU MET LEU ASN LEU LEU CYS SER ASN ALA LEU PHE ILE
SEQRES  19 B  371  ILE CYS GLY PHE ASP SER LYS ASN PHE ASN THR SER ARG
SEQRES  20 B  371  LEU ASP VAL TYR LEU SER HIS ASN PRO ALA GLY THR SER
SEQRES  21 B  371  VAL GLN ASN MET PHE HIS TRP THR GLN ALA VAL LYS SER
SEQRES  22 B  371  GLY LYS PHE GLN ALA TYR ASP TRP GLY SER PRO VAL GLN
SEQRES  23 B  371  ASN ARG MET HIS TYR ASP GLN SER GLN PRO PRO TYR TYR
SEQRES  24 B  371  ASN VAL THR ALA MET ASN VAL PRO ILE ALA VAL TRP ASN
SEQRES  25 B  371  GLY GLY LYS ASP LEU LEU ALA ASP PRO GLN ASP VAL GLY
SEQRES  26 B  371  LEU LEU LEU PRO LYS LEU PRO ASN LEU ILE TYR HIS LYS
SEQRES  27 B  371  GLU ILE PRO PHE TYR ASN HIS LEU ASP PHE ILE TRP ALA
SEQRES  28 B  371  MET ASP ALA PRO GLN GLU VAL TYR ASN ASP ILE VAL SER
SEQRES  29 B  371  MET ILE SER GLU ASP LYS LYS
MODRES 1HLG ASN A   15  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN A   80  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN A  252  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN A  308  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN B   15  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN B   80  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN B  252  ASN  GLYCOSYLATION SITE
MODRES 1HLG ASN B  308  ASN  GLYCOSYLATION SITE
HET    NAG  C   1      14
HET    NAG  C   2      14
HET    NAG  C   3      14
HET    NAG  C   4      14
HET    NAG  C   5      14
HET    NAG  D   1      14
HET    NAG  D   2      14
HET    NAG  D   3      14
HET    NAG  D   4      14
HET    NAG  D   5      14
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETSYN     NAG NAG
FORMUL   3  NAG    10(C8 H15 N1 O6)
FORMUL  11  HOH   *92(H2 O1)
HELIX    1   1 SER A    9  MET A   14  5                                   6
HELIX    2   2 ASN A   15  TRP A   23  1                                   9
HELIX    3   3 SER A   70  SER A   76  5                                   7
HELIX    4   4 SER A   82  ALA A   90  1                                   9
HELIX    5   5 VAL A  116  ALA A  120  5                                   5
HELIX    6   6 SER A  122  TYR A  129  1                                   8
HELIX    7   7 TYR A  129  GLY A  143  1                                  15
HELIX    8   8 SER A  153  ASN A  166  1                                  14
HELIX    9   9 ASN A  166  LYS A  171  1                                   6
HELIX   10  10 SER A  190  VAL A  198  5                                   9
HELIX   11  11 PRO A  199  GLY A  208  1                                  10
HELIX   12  12 PHE A  221  MET A  231  1                                  11
HELIX   13  13 LEU A  232  SER A  237  1                                   6
HELIX   14  14 ASN A  238  GLY A  245  1                                   8
HELIX   15  15 ASN A  252  SER A  254  5                                   3
HELIX   16  16 ARG A  255  SER A  261  1                                   7
HELIX   17  17 SER A  268  GLY A  282  1                                  15
HELIX   18  18 SER A  291  ASP A  300  1                                  10
HELIX   19  19 ASN A  308  MET A  312  5                                   5
HELIX   20  20 ASP A  328  LEU A  339  1                                  12
HELIX   21  21 LEU A  354  ALA A  359  1                                   6
HELIX   22  22 ASP A  361  VAL A  366  1                                   6
HELIX   23  23 VAL A  366  SER A  375  1                                  10
HELIX   24  24 SER B    9  MET B   14  5                                   6
HELIX   25  25 ASN B   15  TRP B   23  1                                   9
HELIX   26  26 SER B   70  SER B   76  5                                   7
HELIX   27  27 SER B   82  ALA B   90  1                                   9
HELIX   28  28 VAL B  116  ALA B  120  5                                   5
HELIX   29  29 SER B  122  TYR B  129  1                                   8
HELIX   30  30 TYR B  129  GLY B  143  1                                  15
HELIX   31  31 SER B  153  SER B  164  1                                  12
HELIX   32  32 ASN B  166  LYS B  171  1                                   6
HELIX   33  33 SER B  190  VAL B  198  5                                   9
HELIX   34  34 PRO B  199  GLY B  208  1                                  10
HELIX   35  35 PHE B  221  MET B  231  1                                  11
HELIX   36  36 LEU B  232  SER B  237  1                                   6
HELIX   37  37 ASN B  238  GLY B  245  1                                   8
HELIX   38  38 ASN B  252  SER B  254  5                                   3
HELIX   39  39 ARG B  255  SER B  261  1                                   7
HELIX   40  40 SER B  268  GLY B  282  1                                  15
HELIX   41  41 SER B  291  ASP B  300  1                                  10
HELIX   42  42 ASN B  308  MET B  312  5                                   5
HELIX   43  43 ASP B  328  LEU B  339  1                                  12
HELIX   44  44 LEU B  354  ALA B  359  1                                   6
HELIX   45  45 ASP B  361  VAL B  366  1                                   6
HELIX   46  46 VAL B  366  SER B  375  1                                  10
SHEET    1   A 8 GLU A  28  VAL A  33  0
SHEET    2   A 8 ILE A  39  ILE A  45 -1  N  LEU A  40   O  VAL A  32
SHEET    3   A 8 ASP A  93  LEU A  96 -1  O  VAL A  94   N  ILE A  45
SHEET    4   A 8 VAL A  60  GLN A  64  1  O  VAL A  61   N  TRP A  95
SHEET    5   A 8 LEU A 147  HIS A 152  1  N  HIS A 148   O  VAL A  60
SHEET    6   A 8 ILE A 173  LEU A 179  1  N  LYS A 174   O  LEU A 147
SHEET    7   A 8 ILE A 316  GLY A 321  1  O  ALA A 317   N  ALA A 178
SHEET    8   A 8 LEU A 342  ILE A 348  1  N  ILE A 343   O  ILE A 316
SHEET    1   B 8 GLU B  28  VAL B  33  0
SHEET    2   B 8 TYR B  38  ILE B  45 -1  N  LEU B  40   O  VAL B  32
SHEET    3   B 8 ASP B  93  LEU B  96 -1  O  VAL B  94   N  ILE B  45
SHEET    4   B 8 VAL B  60  GLN B  64  1  N  VAL B  61   O  ASP B  93
SHEET    5   B 8 LEU B 147  HIS B 152  1  N  HIS B 148   O  VAL B  60
SHEET    6   B 8 ILE B 173  LEU B 179  1  N  LYS B 174   O  LEU B 147
SHEET    7   B 8 ILE B 316  GLY B 321  1  O  ALA B 317   N  ALA B 178
SHEET    8   B 8 LEU B 342  ILE B 348  1  N  ILE B 343   O  ILE B 316
SHEET    1   C 3 GLU B  28  VAL B  33  0
SHEET    2   C 3 TYR B  38  ILE B  45 -1  N  LEU B  40   O  VAL B  32
SHEET    3   C 3 ARG B 107  ASN B 108 -1  O  ARG B 107   N  ILE B  39
SSBOND   1 CYS A  227    CYS A  236
SSBOND   2 CYS B  227    CYS B  236
LINK         O4  NAG C   3                 C1  NAG C   4
LINK         O4  NAG D   3                 C1  NAG D   4
LINK         ND2 ASN A  15                 C1  NAG C   1
LINK         ND2 ASN A  80                 C1  NAG C   2
LINK         ND2 ASN A 252                 C1  NAG C   3
LINK         ND2 ASN A 308                 C1  NAG C   5
LINK         ND2 ASN B  15                 C1  NAG D   1
LINK         ND2 ASN B  80                 C1  NAG D   2
LINK         ND2 ASN B 252                 C1  NAG D   3
LINK         ND2 ASN B 308                 C1  NAG D   5
CISPEP   1 ASN A  263    PRO A  264          0        -0.34
CISPEP   2 ASN B  263    PRO B  264          0        -0.36
SITE     1  SA  3 SER A 153  ASP A 324  HIS A 353
SITE     2  SB  3 SER B 153  ASP B 324  HIS B 353
CRYST1  244.330  244.330  244.330  90.00  90.00  90.00 I 21 3       48
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.004093  0.000000  0.000000        0.00000
SCALE2      0.000000  0.004093  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004093        0.00000
END