longtext: 1IVY-pdb

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HEADER    CARBOXYPEPTIDASE                        12-JUN-96   1IVY
TITLE     PHYSIOLOGICAL DIMER HPP PRECURSOR
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: HUMAN PROTECTIVE PROTEIN;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: PROTECTIVE PROTEIN/CATHEPSIN A,
COMPND   5 CARBOXYPEPTIDASE L;
COMPND   6 EC: 3.4.16.5;
COMPND   7 ENGINEERED: YES;
COMPND   8 BIOLOGICAL_UNIT: HETERODIMER
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 EXPRESSION_SYSTEM: SF21 INSECT CELLS;
SOURCE   5 EXPRESSION_SYSTEM_GENE: HUPP54;
SOURCE   6 OTHER_DETAILS: BACULOVIRUS MEDIATED OVER-EXPRESSION. SEE
SOURCE   7 BONTEN ET AL. 1995, J.B.C. 270, P. 26441-26445
KEYWDS    CARBOXYPEPTIDASE, SERINE CARBOXYPEPTIDASE,
KEYWDS   2 PROTECTIVE PROTEIN, GLYCOPROTEIN, ZYMOGEN
EXPDTA    X-RAY DIFFRACTION
AUTHOR    G.RUDENKO,E.BONTEN,A.D'AZZO,W.G.J.HOL
REVDAT   1   21-APR-97 1IVY    0
JRNL        AUTH   G.RUDENKO,E.BONTEN,A.D'AZZO,W.G.HOL
JRNL        TITL   THREE-DIMENSIONAL STRUCTURE OF THE HUMAN
JRNL        TITL 2 'PROTECTIVE PROTEIN': STRUCTURE OF THE PRECURSOR
JRNL        TITL 3 FORM SUGGESTS A COMPLEX ACTIVATION MECHANISM
JRNL        REF    STRUCTURE (LONDON)            V.   3  1249 1995
JRNL        REFN   ASTM STRUE6  UK ISSN 0969-2126                 2005
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   G.RUDENKO,E.BONTEN,A.D'AZZO,W.G.J.HOL
REMARK   1  TITL   STRUCTURE DETERMINATION OF THE HUMAN PROTECTIVE
REMARK   1  TITL 2 PROTEIN: TWOFOLD AVERAGING REVEALS THE
REMARK   1  TITL 3 THREE-DIMENSIONAL STRUCTURE OF A DOMAIN WHICH WAS
REMARK   1  TITL 4 ENTIRELY ABSENT IN THE INITIAL MODEL
REMARK   1  REF    ACTA CRYSTALLOGR.,SECT.D      V.  52   923 1996
REMARK   1  REFN   ASTM ABCRE6  DK ISSN 0907-4449                 0766
REMARK   1 REFERENCE 2
REMARK   1  AUTH   N.J.GALJART,N.GILLEMANS,A.HARRIS,
REMARK   1  AUTH 2 G.T.VAN DER HORST,F.W.VERHEIJEN,H.GALJAARD,A.D'AZZO
REMARK   1  TITL   EXPRESSION OF CDNA ENCODING THE HUMAN 'PROTECTIVE
REMARK   1  TITL 2 PROTEIN' ASSOCIATED WITH LYSOSOMAL
REMARK   1  TITL 3 BETA-GALACTOSIDASE AND NEURAMINIDASE: HOMOLOGY TO
REMARK   1  TITL 4 YEAST PROTEASES
REMARK   1  REF    CELL(CAMBRIDGE,MASS.)         V.  54   755 1988
REMARK   1  REFN   ASTM CELLB5  US ISSN 0092-8674                 0998
REMARK   2
REMARK   2 RESOLUTION. 2.2  ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : X-PLOR 3.1
REMARK   3   AUTHORS     : BRUNGER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.2
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 8.
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 82.9
REMARK   3   NUMBER OF REFLECTIONS             : 57704
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL
REMARK   3   R VALUE            (WORKING SET) : 0.213
REMARK   3   FREE R VALUE                     : 0.268
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.0
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.2
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.3
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 56.9
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : 0.240
REMARK   3   BIN FREE R VALUE                    : NULL
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 7173
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 84
REMARK   3   SOLVENT ATOMS            : 296
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : 0.282
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.012
REMARK   3   BOND ANGLES            (DEGREES) : 1.72
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  PARAMETER FILE  2  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3  TOPOLOGY FILE  2   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS:
REMARK   3   ESTIMATED COORD. ERROR     0.282 ANGSTROMS
REMARK   3   MEAN B (A**2) (MAIN CHAIN): 16.6
REMARK   3   MEAN B (A**2) (SIDE CHAIN): 18.3
REMARK   4
REMARK   4 1IVY COMPLIES WITH FORMAT V. 2.2, 16-DEC-1996
REMARK   6
REMARK   6 BECAUSE OF SIGNIFICANT DIFFERENCES BETWEEN THE TWO
REMARK   6 MONOMERS DUE TO A CRYSTAL CONTACT, THE COORDINATES OF A
REMARK   6 DIMER HAVE BEEN SUBMITTED.
REMARK   7
REMARK   7 RESIDUES ASN B 216 AND LEU B 217 ARE IN AN AREA OF POOR
REMARK   7 ELECTRON DENSITY.  THE EXACT GEOMETRY OF THESE RESIDUES
REMARK   7 IS NOT CLEAR FROM THE ELECTRON DENSITY MAP, BUT THEIR
REMARK   7 GENERAL POSITION IS.  IN ADDITION, THE RESTRAINING
REMARK   7 DISULFIDE CYS 213 - CYS 218 SUPPORTS THE DEPOSITORS'
REMARK   7 MODEL IN THIS AREA.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 24-MAR-1994
REMARK 200  TEMPERATURE           (KELVIN) : 95
REMARK 200  PH                             : 8.
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRL
REMARK 200  BEAMLINE                       : 7.1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.08
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : MOSFLM 5.2
REMARK 200  DATA SCALING SOFTWARE          : ROTAVATA, AGROVATA,
REMARK 200                                   TRUNCATE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 67740
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.2
REMARK 200  RESOLUTION RANGE LOW       (A) : 32.3
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 95.7
REMARK 200  DATA REDUNDANCY                : ~6.
REMARK 200  R MERGE                    (I) : 0.051
REMARK 200  R SYM                      (I) : NULL
REMARK 200  <I/SIGMA(I)> FOR THE DATA SET  : 9.8
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.26
REMARK 200  COMPLETENESS FOR SHELL     (%) : 87.0
REMARK 200  DATA REDUNDANCY IN SHELL       : ~6.
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : 0.130
REMARK 200  <I/SIGMA(I)> FOR SHELL         : 5.7
REMARK 200
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR
REMARK 200                       REPLACEMENT AND TWO-FOLD AVERAGING
REMARK 200 SOFTWARE USED: X-PLOR, RAVE, CCCP4 SUITE
REMARK 200 STARTING MODEL: SERINE CARBOXYPEPTIDASE FROM WHEAT
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 62.
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.2
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG 8000, PH 8.0 - 8.3 AT
REMARK 280 4 - 12 DEGREES CELSIUS
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   1/2-X,1/2+Y,-Z
REMARK 290       4555   1/2+X,1/2-Y,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000       57.51754
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       74.05213
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       57.51754
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       74.05213
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLN A   8    CG   CD   OE1  NE2
REMARK 470     ARG A   9    CG   CD   NE   CZ   NH1  NH2
REMARK 470     GLU A  46    CG   CD   OE1  OE2
REMARK 470     ASN A  47    CG   OD1  ND2
REMARK 470     PHE A 113    CG   CD1  CE1  CZ   CE2  CD2
REMARK 470     GLU A 326    CG   CD   OE1  OE2
REMARK 470     VAL A 393    CG1  CG2
REMARK 470     GLU B  46    CG   CD   OE1  OE2
REMARK 470     GLN B 215    CG   CD   OE1  NE2
REMARK 470     LYS B 217    CG   CD   CE   NZ
REMARK 470     PHE B 261    CG   CD1  CE1  CZ   CE2  CD2
REMARK 470     TYR B 263    CG   CD1  CE1  CZ   CE2  CD2  OH
REMARK 470     LYS B 265    CG   CD   CE   NZ
REMARK 470     LYS B 283    CG   CD   CE   NZ
REMARK 470     LYS B 296    CG   CD   CE   NZ
REMARK 470     GLU B 326    CG   CD   OE1  OE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 ASN B  216    LYS B  217          0       124.19
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: CAT
REMARK 800 SITE_DESCRIPTION: CATALYTIC TRIAD SERINE CARBOXYPEPTIDASE.
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 1IVY  A    SWS     P10619       1 -    28 NOT IN ATOMS LIST
REMARK 999 1IVY  B    SWS     P10619       1 -    28 NOT IN ATOMS LIST
DBREF  1IVY A    1   452  SWS    P10619   PRTP_HUMAN      29    480
DBREF  1IVY B    1   258  SWS    P10619   PRTP_HUMAN      29    286
DBREF  1IVY B  261   452  SWS    P10619   PRTP_HUMAN     289    480
SEQADV 1IVY     B       SWS  P10619    SER   287 GAP IN PDB ENTRY
SEQADV 1IVY     B       SWS  P10619    HIS   288 GAP IN PDB ENTRY
SEQRES   1 A  452  ALA PRO ASP GLN ASP GLU ILE GLN ARG LEU PRO GLY LEU
SEQRES   2 A  452  ALA LYS GLN PRO SER PHE ARG GLN TYR SER GLY TYR LEU
SEQRES   3 A  452  LYS SER SER GLY SER LYS HIS LEU HIS TYR TRP PHE VAL
SEQRES   4 A  452  GLU SER GLN LYS ASP PRO GLU ASN SER PRO VAL VAL LEU
SEQRES   5 A  452  TRP LEU ASN GLY GLY PRO GLY CYS SER SER LEU ASP GLY
SEQRES   6 A  452  LEU LEU THR GLU HIS GLY PRO PHE LEU VAL GLN PRO ASP
SEQRES   7 A  452  GLY VAL THR LEU GLU TYR ASN PRO TYR SER TRP ASN LEU
SEQRES   8 A  452  ILE ALA ASN VAL LEU TYR LEU GLU SER PRO ALA GLY VAL
SEQRES   9 A  452  GLY PHE SER TYR SER ASP ASP LYS PHE TYR ALA THR ASN
SEQRES  10 A  452  ASP THR GLU VAL ALA GLN SER ASN PHE GLU ALA LEU GLN
SEQRES  11 A  452  ASP PHE PHE ARG LEU PHE PRO GLU TYR LYS ASN ASN LYS
SEQRES  12 A  452  LEU PHE LEU THR GLY GLU SER TYR ALA GLY ILE TYR ILE
SEQRES  13 A  452  PRO THR LEU ALA VAL LEU VAL MET GLN ASP PRO SER MET
SEQRES  14 A  452  ASN LEU GLN GLY LEU ALA VAL GLY ASN GLY LEU SER SER
SEQRES  15 A  452  TYR GLU GLN ASN ASP ASN SER LEU VAL TYR PHE ALA TYR
SEQRES  16 A  452  TYR HIS GLY LEU LEU GLY ASN ARG LEU TRP SER SER LEU
SEQRES  17 A  452  GLN THR HIS CYS CYS SER GLN ASN LYS CYS ASN PHE TYR
SEQRES  18 A  452  ASP ASN LYS ASP LEU GLU CYS VAL THR ASN LEU GLN GLU
SEQRES  19 A  452  VAL ALA ARG ILE VAL GLY ASN SER GLY LEU ASN ILE TYR
SEQRES  20 A  452  ASN LEU TYR ALA PRO CYS ALA GLY GLY VAL PRO SER HIS
SEQRES  21 A  452  PHE ARG TYR GLU LYS ASP THR VAL VAL VAL GLN ASP LEU
SEQRES  22 A  452  GLY ASN ILE PHE THR ARG LEU PRO LEU LYS ARG MET TRP
SEQRES  23 A  452  HIS GLN ALA LEU LEU ARG SER GLY ASP LYS VAL ARG MET
SEQRES  24 A  452  ASP PRO PRO CYS THR ASN THR THR ALA ALA SER THR TYR
SEQRES  25 A  452  LEU ASN ASN PRO TYR VAL ARG LYS ALA LEU ASN ILE PRO
SEQRES  26 A  452  GLU GLN LEU PRO GLN TRP ASP MET CYS ASN PHE LEU VAL
SEQRES  27 A  452  ASN LEU GLN TYR ARG ARG LEU TYR ARG SER MET ASN SER
SEQRES  28 A  452  GLN TYR LEU LYS LEU LEU SER SER GLN LYS TYR GLN ILE
SEQRES  29 A  452  LEU LEU TYR ASN GLY ASP VAL ASP MET ALA CYS ASN PHE
SEQRES  30 A  452  MET GLY ASP GLU TRP PHE VAL ASP SER LEU ASN GLN LYS
SEQRES  31 A  452  MET GLU VAL GLN ARG ARG PRO TRP LEU VAL LYS TYR GLY
SEQRES  32 A  452  ASP SER GLY GLU GLN ILE ALA GLY PHE VAL LYS GLU PHE
SEQRES  33 A  452  SER HIS ILE ALA PHE LEU THR ILE LYS GLY ALA GLY HIS
SEQRES  34 A  452  MET VAL PRO THR ASP LYS PRO LEU ALA ALA PHE THR MET
SEQRES  35 A  452  PHE SER ARG PHE LEU ASN LYS GLN PRO TYR
SEQRES   1 B  452  ALA PRO ASP GLN ASP GLU ILE GLN ARG LEU PRO GLY LEU
SEQRES   2 B  452  ALA LYS GLN PRO SER PHE ARG GLN TYR SER GLY TYR LEU
SEQRES   3 B  452  LYS SER SER GLY SER LYS HIS LEU HIS TYR TRP PHE VAL
SEQRES   4 B  452  GLU SER GLN LYS ASP PRO GLU ASN SER PRO VAL VAL LEU
SEQRES   5 B  452  TRP LEU ASN GLY GLY PRO GLY CYS SER SER LEU ASP GLY
SEQRES   6 B  452  LEU LEU THR GLU HIS GLY PRO PHE LEU VAL GLN PRO ASP
SEQRES   7 B  452  GLY VAL THR LEU GLU TYR ASN PRO TYR SER TRP ASN LEU
SEQRES   8 B  452  ILE ALA ASN VAL LEU TYR LEU GLU SER PRO ALA GLY VAL
SEQRES   9 B  452  GLY PHE SER TYR SER ASP ASP LYS PHE TYR ALA THR ASN
SEQRES  10 B  452  ASP THR GLU VAL ALA GLN SER ASN PHE GLU ALA LEU GLN
SEQRES  11 B  452  ASP PHE PHE ARG LEU PHE PRO GLU TYR LYS ASN ASN LYS
SEQRES  12 B  452  LEU PHE LEU THR GLY GLU SER TYR ALA GLY ILE TYR ILE
SEQRES  13 B  452  PRO THR LEU ALA VAL LEU VAL MET GLN ASP PRO SER MET
SEQRES  14 B  452  ASN LEU GLN GLY LEU ALA VAL GLY ASN GLY LEU SER SER
SEQRES  15 B  452  TYR GLU GLN ASN ASP ASN SER LEU VAL TYR PHE ALA TYR
SEQRES  16 B  452  TYR HIS GLY LEU LEU GLY ASN ARG LEU TRP SER SER LEU
SEQRES  17 B  452  GLN THR HIS CYS CYS SER GLN ASN LYS CYS ASN PHE TYR
SEQRES  18 B  452  ASP ASN LYS ASP LEU GLU CYS VAL THR ASN LEU GLN GLU
SEQRES  19 B  452  VAL ALA ARG ILE VAL GLY ASN SER GLY LEU ASN ILE TYR
SEQRES  20 B  452  ASN LEU TYR ALA PRO CYS ALA GLY GLY VAL PRO SER HIS
SEQRES  21 B  452  PHE ARG TYR GLU LYS ASP THR VAL VAL VAL GLN ASP LEU
SEQRES  22 B  452  GLY ASN ILE PHE THR ARG LEU PRO LEU LYS ARG MET TRP
SEQRES  23 B  452  HIS GLN ALA LEU LEU ARG SER GLY ASP LYS VAL ARG MET
SEQRES  24 B  452  ASP PRO PRO CYS THR ASN THR THR ALA ALA SER THR TYR
SEQRES  25 B  452  LEU ASN ASN PRO TYR VAL ARG LYS ALA LEU ASN ILE PRO
SEQRES  26 B  452  GLU GLN LEU PRO GLN TRP ASP MET CYS ASN PHE LEU VAL
SEQRES  27 B  452  ASN LEU GLN TYR ARG ARG LEU TYR ARG SER MET ASN SER
SEQRES  28 B  452  GLN TYR LEU LYS LEU LEU SER SER GLN LYS TYR GLN ILE
SEQRES  29 B  452  LEU LEU TYR ASN GLY ASP VAL ASP MET ALA CYS ASN PHE
SEQRES  30 B  452  MET GLY ASP GLU TRP PHE VAL ASP SER LEU ASN GLN LYS
SEQRES  31 B  452  MET GLU VAL GLN ARG ARG PRO TRP LEU VAL LYS TYR GLY
SEQRES  32 B  452  ASP SER GLY GLU GLN ILE ALA GLY PHE VAL LYS GLU PHE
SEQRES  33 B  452  SER HIS ILE ALA PHE LEU THR ILE LYS GLY ALA GLY HIS
SEQRES  34 B  452  MET VAL PRO THR ASP LYS PRO LEU ALA ALA PHE THR MET
SEQRES  35 B  452  PHE SER ARG PHE LEU ASN LYS GLN PRO TYR
HET    NAG  A3010      14
HET    NAG  A3011      14
HET    NAG  A3020      14
HET    NAG  B3030      14
HET    NAG  B3031      14
HET    NAG  B3040      14
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   3  NAG    6(C8 H15 N1 O6)
FORMUL   4  HOH   *296(H2 O1)
HELIX    1   1 LEU A   63  LEU A   67  1                                   5
HELIX    2   2 ASP A  118  LEU A  135  1                                  18
HELIX    3   3 ALA A  152  VAL A  163  5                                  12
HELIX    4   4 TYR A  183  TYR A  196  1                                  14
HELIX    5   5 ASN A  202  CYS A  212  1                                  11
HELIX    6   6 LEU A  226  GLY A  240  1                                  15
HELIX    7   7 LEU A  290  SER A  293  5                                   4
HELIX    8   8 THR A  307  LEU A  313  1                                   7
HELIX    9   9 PRO A  316  ALA A  321  1                                   6
HELIX   10  10 PHE A  336  GLN A  341  1                                   6
HELIX   11  11 ASN A  350  SER A  359  1                                  10
HELIX   12  12 PHE A  377  SER A  386  1                                  10
HELIX   13  13 VAL A  431  ASP A  434  1                                   4
HELIX   14  14 PRO A  436  LEU A  447  1                                  12
HELIX   15  15 LEU B   63  LEU B   67  1                                   5
HELIX   16  16 ASP B  118  LEU B  135  1                                  18
HELIX   17  17 ALA B  152  VAL B  163  5                                  12
HELIX   18  18 TYR B  183  TYR B  196  1                                  14
HELIX   19  19 ASN B  202  CYS B  212  1                                  11
HELIX   20  20 LEU B  226  GLY B  240  1                                  15
HELIX   21  21 THR B  307  LEU B  313  1                                   7
HELIX   22  22 PRO B  316  ALA B  321  1                                   6
HELIX   23  23 PHE B  336  GLN B  341  1                                   6
HELIX   24  24 ASN B  350  SER B  359  1                                  10
HELIX   25  25 PHE B  377  SER B  386  1                                  10
HELIX   26  26 VAL B  431  ASP B  434  1                                   4
HELIX   27  27 PRO B  436  LEU B  447  1                                  12
SHEET    1   A10 GLN A  21  LYS A  27  0
SHEET    2   A10 LYS A  32  VAL A  39 -1  N  PHE A  38   O  TYR A  22
SHEET    3   A10 ASN A  94  LEU A  98 -1  N  TYR A  97   O  TRP A  37
SHEET    4   A10 VAL A  50  LEU A  54  1  N  VAL A  51   O  ASN A  94
SHEET    5   A10 LEU A 144  GLU A 149  1  N  PHE A 145   O  VAL A  50
SHEET    6   A10 LEU A 171  GLY A 177  1  N  GLN A 172   O  LEU A 144
SHEET    7   A10 GLN A 363  GLY A 369  1  N  GLN A 363   O  LEU A 174
SHEET    8   A10 ILE A 419  ILE A 424  1  N  ALA A 420   O  ILE A 364
SHEET    9   A10 GLU A 407  PHE A 416 -1  N  LYS A 414   O  PHE A 421
SHEET   10   A10 MET A 391  LYS A 401 -1  N  VAL A 400   O  GLN A 408
SHEET    1   B 2 PHE A  73  VAL A  75  0
SHEET    2   B 2 LEU A  82  TYR A  84 -1  N  GLU A  83   O  LEU A  74
SHEET    1   C 3 PHE A 261  GLU A 264  0
SHEET    2   C 3 THR A 267  VAL A 270 -1  N  VAL A 269   O  ARG A 262
SHEET    3   C 3 LYS A 296  MET A 299  1  N  LYS A 296   O  VAL A 268
SHEET    1   D10 GLN B  21  LYS B  27  0
SHEET    2   D10 LYS B  32  VAL B  39 -1  N  PHE B  38   O  TYR B  22
SHEET    3   D10 ASN B  94  LEU B  98 -1  N  TYR B  97   O  TRP B  37
SHEET    4   D10 VAL B  50  LEU B  54  1  N  VAL B  51   O  ASN B  94
SHEET    5   D10 LEU B 144  GLU B 149  1  N  PHE B 145   O  VAL B  50
SHEET    6   D10 LEU B 171  GLY B 177  1  N  GLN B 172   O  LEU B 144
SHEET    7   D10 GLN B 363  GLY B 369  1  N  GLN B 363   O  LEU B 174
SHEET    8   D10 ILE B 419  ILE B 424  1  N  ALA B 420   O  ILE B 364
SHEET    9   D10 GLU B 407  PHE B 416 -1  N  PHE B 416   O  ILE B 419
SHEET   10   D10 MET B 391  LYS B 401 -1  N  VAL B 400   O  GLN B 408
SHEET    1   E 2 PHE B  73  VAL B  75  0
SHEET    2   E 2 LEU B  82  TYR B  84 -1  N  GLU B  83   O  LEU B  74
SHEET    1   F 2 THR B 267  VAL B 270  0
SHEET    2   F 2 LYS B 296  MET B 299  1  N  LYS B 296   O  VAL B 268
SSBOND   1 CYS A   60    CYS A  334
SSBOND   2 CYS A  212    CYS A  228
SSBOND   3 CYS A  213    CYS A  218
SSBOND   4 CYS A  253    CYS A  303
SSBOND   5 CYS B   60    CYS B  334
SSBOND   6 CYS B  212    CYS B  228
SSBOND   7 CYS B  213    CYS B  218
SSBOND   8 CYS B  253    CYS B  303
LINK         C1  NAG A3010                 ND2 ASN A 117
LINK         O4  NAG A3010                 C1  NAG A3011
LINK         C1  NAG A3020                 ND2 ASN A 305
LINK         C1  NAG B3030                 ND2 ASN B 117
LINK         O4  NAG B3030                 C1  NAG B3031
LINK         C1  NAG B3040                 ND2 ASN B 305
CISPEP   1 GLY A   57    PRO A   58          0         1.29
CISPEP   2 SER A  100    PRO A  101          0         0.52
CISPEP   3 GLY B   57    PRO B   58          0        -0.46
CISPEP   4 SER B  100    PRO B  101          0        -0.63
SITE     1 CAT  6 SER A 150  ASP A 372  HIS A 429  SER B 150
SITE     2 CAT  6 ASP B 372  HIS B 429
CRYST1  115.040  148.110   80.970  90.00  90.00  90.00 P 21 21 2     8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008693  0.000000  0.000000        0.00000
SCALE2      0.000000  0.006752  0.000000        0.00000
SCALE3      0.000000  0.000000  0.012350        0.00000