longtext: 1JJB-pdb

content
HEADER    HYDROLASE                               04-JUL-01   1JJB
TITLE     A NEUTRAL MOLECULE IN CATION-BINDING SITE: SPECIFIC BINDING
TITLE    2 OF PEG-SH TO ACETYLCHOLINESTERASE FROM TORPEDO CALIFORNICA
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYLCHOLINESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.1.1.7
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: TORPEDO CALIFORNICA;
SOURCE   3 STRAIN: PACIFIC ELECTRIC RAY;
SOURCE   4 OTHER_DETAILS: ELECTRIC ORGAN
KEYWDS    PEG (POLYETHYLENE), SERINE HYDROLASE, ALPHA/BETA HYDROLASE,
KEYWDS   2 NEUROTRANSMITTER CLEAVAGE, CATALYTIC TRIAD, GLYCOSYLATED
KEYWDS   3 PROTEIN
EXPDTA    X-RAY DIFFRACTION
AUTHOR    G.KOELLNER,T.STEINER,C.B.MILLARD,I.SILMAN,J.L.SUSSMANN
REVDAT   1   17-JUL-02 1JJB    0
JRNL        AUTH   G.KOELLNER,T.STEINER,C.B.MILLARD,I.SILMAN,
JRNL        AUTH 2 J.L.SUSSMANN
JRNL        TITL   A NEUTRAL MOLECULE IN A CATION-BINDING SITE:
JRNL        TITL 2 SPECIFIC BINDING OF A PEG-SH TO
JRNL        TITL 3 ACETYLCHOLINESTERASE FROM TORPEDO CALIFORNICA
JRNL        REF    J.MOL.BIOL.                   V. 320   721 2002
JRNL        REFN   ASTM JMOBAK  UK ISSN 0022-2836
REMARK   1
REMARK   2
REMARK   2 RESOLUTION. 2.30 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.0
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.30
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.89
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 1.000
REMARK   3   OUTLIER CUTOFF HIGH (RMS(ABS(F))) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 94.1
REMARK   3   NUMBER OF REFLECTIONS             : 41332
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.178
REMARK   3   FREE R VALUE                     : 0.224
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 10.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 4165
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.003
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.30
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.44
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 91.70
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 5928
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2450
REMARK   3   BIN FREE R VALUE                    : 0.2730
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 10.60
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 702
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.010
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4201
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 50
REMARK   3   SOLVENT ATOMS            : 623
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 32.80
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 34.60
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 6.31000
REMARK   3    B22 (A**2) : 6.31000
REMARK   3    B33 (A**2) : -12.62000
REMARK   3    B12 (A**2) : 4.13000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.23
REMARK   3   ESD FROM SIGMAA              (A) : 0.26
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.29
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.29
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.006
REMARK   3   BOND ANGLES            (DEGREES) : 1.30
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 22.70
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.81
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 0.660 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 1.140 ; 2.000
REMARK   3   SIDE-CHAIN BOND              (A**2) : 6.490 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 7.600 ; 2.500
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : FLAT MODEL
REMARK   3   KSOL        : 0.38
REMARK   3   BSOL        : 80.70
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 1JJB COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 19-JUL-2001.
REMARK 100 THE RCSB ID CODE IS RCSB013838.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 23-JUL-2000
REMARK 200  TEMPERATURE           (KELVIN) : 100.0
REMARK 200  PH                             : 5.80
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : ROTATING ANODE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : ENRAF-NONIUS FR571
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.54
REMARK 200  MONOCHROMATOR                  : GRAPHITE
REMARK 200  OPTICS                         : MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 41349
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.300
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 94.1
REMARK 200  DATA REDUNDANCY                : 5.900
REMARK 200  R MERGE                    (I) : 0.04300
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 12.9000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.30
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.38
REMARK 200  COMPLETENESS FOR SHELL     (%) : 93.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 7.00
REMARK 200  R MERGE FOR SHELL          (I) : 0.23000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: FOURIER SYNTHESIS
REMARK 200 SOFTWARE USED: CNS
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): NULL
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 34% PEG 200, 0.05M MES
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 31 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,1/3+Z
REMARK 290       3555   -X+Y,-X,2/3+Z
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,2/3-Z
REMARK 290       6555   -X,-X+Y,1/3-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       45.70633
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       91.41267
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       91.41267
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       45.70633
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT
REMARK 300 WHICH CONSISTS OF 1 CHAIN(S). SEE REMARK 350 FOR
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH  1281   LIES ON A SPECIAL POSITION.
REMARK 375      HOH  1496   LIES ON A SPECIAL POSITION.
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS(M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ASN A  42    CG    OD1   ND2
REMARK 470     ARG A  46    CZ    NH1   NH2
REMARK 470     LYS A  52    CG    CD    CE    NZ
REMARK 470     GLU A  89    CD    OE1   OE2
REMARK 470     GLU A 260    OE1   OE2
REMARK 470     GLU A 268    OE1   OE2
REMARK 470     LYS A 270    CE    NZ
REMARK 470     LYS A 357    CG    CD    CE    NZ
REMARK 470     LYS A 491    CD    CE    NZ
REMARK 470     LYS A 498    CG    CD    CE    NZ
REMARK 470     GLU A 499    OE1   OE2
REMARK 470     GLU A 508    CD    OE1   OE2
REMARK 470     LYS A 511    CD    CE    NZ
REMARK 470     ARG A 515    CZ    NH1   NH2
REMARK 470     GLN A 526    OE1   NE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI
REMARK 500   O    HOH    1550     O    HOH    1573              2.15
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH    1563     O    HOH    1563     4556     1.92
REMARK 500   O    HOH    1465     O    HOH    1465     5675     2.05
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    MET A 175   CE    MET A 175   SD    -0.140
REMARK 500    MET A 405   CE    MET A 405   SD     0.050
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ASP A  93   N   -  CA  -  C   ANGL. DEV. = -9.1 DEGREES
REMARK 500    VAL A 142   N   -  CA  -  C   ANGL. DEV. = -8.0 DEGREES
REMARK 500    ASN A 183   N   -  CA  -  C   ANGL. DEV. =  8.2 DEGREES
REMARK 500    ALA A 234   N   -  CA  -  C   ANGL. DEV. =  8.1 DEGREES
REMARK 500    ASP A 285   N   -  CA  -  C   ANGL. DEV. =-12.4 DEGREES
REMARK 500    SER A 291   N   -  CA  -  C   ANGL. DEV. =  8.0 DEGREES
REMARK 500    ILE A 296   N   -  CA  -  C   ANGL. DEV. = -9.1 DEGREES
REMARK 500    PHE A 301   N   -  CA  -  C   ANGL. DEV. = -8.3 DEGREES
REMARK 500    GLY A 328   N   -  CA  -  C   ANGL. DEV. =  9.9 DEGREES
REMARK 500    GLY A 335   N   -  CA  -  C   ANGL. DEV. =  9.3 DEGREES
REMARK 500    LYS A 346   N   -  CA  -  C   ANGL. DEV. = -8.0 DEGREES
REMARK 500    ASP A 377   N   -  CA  -  C   ANGL. DEV. = -8.9 DEGREES
REMARK 500    ILE A 439   N   -  CA  -  C   ANGL. DEV. =  8.7 DEGREES
REMARK 500    GLY A 449   N   -  CA  -  C   ANGL. DEV. =  9.0 DEGREES
REMARK 500    ASN A 457   N   -  CA  -  C   ANGL. DEV. =  8.5 DEGREES
REMARK 500    TRP A 492   N   -  CA  -  C   ANGL. DEV. = -9.6 DEGREES
REMARK 525
REMARK 525 SOLVENT
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH  1451        DISTANCE =  7.75 ANGSTROMS
REMARK 525    HOH  1542        DISTANCE =  6.23 ANGSTROMS
REMARK 525    HOH  1564        DISTANCE =  6.90 ANGSTROMS
REMARK 525    HOH  1611        DISTANCE =  6.99 ANGSTROMS
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 2ACE   RELATED DB: PDB
REMARK 900 NATIVE ACETYLCHOLINESTERASE (E.C. 3.1.1.7) FROM TORPEDO
REMARK 900 CALIFORNICA
REMARK 900 RELATED ID: 1VXR   RELATED DB: PDB
REMARK 900 O-ETHYLMETHYLPHOSPHONYLATED ACETYLCHOLINESTERASE OBTAINED
REMARK 900 BY REACTION WITH O-ETHYL-S-[2-[BIS(1-METHYLETHYL) AMINO]
REMARK 900 ETHYL] METHYLPHOSPHONOTHIOATE (VX)
DBREF  1JJB A    4   535  SWS    P04058   ACES_TORCA      25    556
SEQRES   1 A  532  SER GLU LEU LEU VAL ASN THR LYS SER GLY LYS VAL MET
SEQRES   2 A  532  GLY THR ARG VAL PRO VAL LEU SER SER HIS ILE SER ALA
SEQRES   3 A  532  PHE LEU GLY ILE PRO PHE ALA GLU PRO PRO VAL GLY ASN
SEQRES   4 A  532  MET ARG PHE ARG ARG PRO GLU PRO LYS LYS PRO TRP SER
SEQRES   5 A  532  GLY VAL TRP ASN ALA SER THR TYR PRO ASN ASN CYS GLN
SEQRES   6 A  532  GLN TYR VAL ASP GLU GLN PHE PRO GLY PHE SER GLY SER
SEQRES   7 A  532  GLU MET TRP ASN PRO ASN ARG GLU MET SER GLU ASP CYS
SEQRES   8 A  532  LEU TYR LEU ASN ILE TRP VAL PRO SER PRO ARG PRO LYS
SEQRES   9 A  532  SER THR THR VAL MET VAL TRP ILE TYR GLY GLY GLY PHE
SEQRES  10 A  532  TYR SER GLY SER SER THR LEU ASP VAL TYR ASN GLY LYS
SEQRES  11 A  532  TYR LEU ALA TYR THR GLU GLU VAL VAL LEU VAL SER LEU
SEQRES  12 A  532  SER TYR ARG VAL GLY ALA PHE GLY PHE LEU ALA LEU HIS
SEQRES  13 A  532  GLY SER GLN GLU ALA PRO GLY ASN VAL GLY LEU LEU ASP
SEQRES  14 A  532  GLN ARG MET ALA LEU GLN TRP VAL HIS ASP ASN ILE GLN
SEQRES  15 A  532  PHE PHE GLY GLY ASP PRO LYS THR VAL THR ILE PHE GLY
SEQRES  16 A  532  GLU SER ALA GLY GLY ALA SER VAL GLY MET HIS ILE LEU
SEQRES  17 A  532  SER PRO GLY SER ARG ASP LEU PHE ARG ARG ALA ILE LEU
SEQRES  18 A  532  GLN SER GLY SER PRO ASN CYS PRO TRP ALA SER VAL SER
SEQRES  19 A  532  VAL ALA GLU GLY ARG ARG ARG ALA VAL GLU LEU GLY ARG
SEQRES  20 A  532  ASN LEU ASN CYS ASN LEU ASN SER ASP GLU GLU LEU ILE
SEQRES  21 A  532  HIS CYS LEU ARG GLU LYS LYS PRO GLN GLU LEU ILE ASP
SEQRES  22 A  532  VAL GLU TRP ASN VAL LEU PRO PHE ASP SER ILE PHE ARG
SEQRES  23 A  532  PHE SER PHE VAL PRO VAL ILE ASP GLY GLU PHE PHE PRO
SEQRES  24 A  532  THR SER LEU GLU SER MET LEU ASN SER GLY ASN PHE LYS
SEQRES  25 A  532  LYS THR GLN ILE LEU LEU GLY VAL ASN LYS ASP GLU GLY
SEQRES  26 A  532  SER PHE PHE LEU LEU TYR GLY ALA PRO GLY PHE SER LYS
SEQRES  27 A  532  ASP SER GLU SER LYS ILE SER ARG GLU ASP PHE MET SER
SEQRES  28 A  532  GLY VAL LYS LEU SER VAL PRO HIS ALA ASN ASP LEU GLY
SEQRES  29 A  532  LEU ASP ALA VAL THR LEU GLN TYR THR ASP TRP MET ASP
SEQRES  30 A  532  ASP ASN ASN GLY ILE LYS ASN ARG ASP GLY LEU ASP ASP
SEQRES  31 A  532  ILE VAL GLY ASP HIS ASN VAL ILE CYS PRO LEU MET HIS
SEQRES  32 A  532  PHE VAL ASN LYS TYR THR LYS PHE GLY ASN GLY THR TYR
SEQRES  33 A  532  LEU TYR PHE PHE ASN HIS ARG ALA SER ASN LEU VAL TRP
SEQRES  34 A  532  PRO GLU TRP MET GLY VAL ILE HIS GLY TYR GLU ILE GLU
SEQRES  35 A  532  PHE VAL PHE GLY LEU PRO LEU VAL LYS GLU LEU ASN TYR
SEQRES  36 A  532  THR ALA GLU GLU GLU ALA LEU SER ARG ARG ILE MET HIS
SEQRES  37 A  532  TYR TRP ALA THR PHE ALA LYS THR GLY ASN PRO ASN GLU
SEQRES  38 A  532  PRO HIS SER GLN GLU SER LYS TRP PRO LEU PHE THR THR
SEQRES  39 A  532  LYS GLU GLN LYS PHE ILE ASP LEU ASN THR GLU PRO MET
SEQRES  40 A  532  LYS VAL HIS GLN ARG LEU ARG VAL GLN MET CYS VAL PHE
SEQRES  41 A  532  TRP ASN GLN PHE LEU PRO LYS LEU LEU ASN ALA THR
MODRES 1JJB ASN A   59  ASN  GLYCOSYLATION SITE
MODRES 1JJB ASN A  416  ASN  GLYCOSYLATION SITE
HET    NAG   9010      14
HET    NAG   9020      14
HET    PE7    999      22
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     PE7 1-DEOXY-1-THIO-HEPTAETHYLENE GLYCOL
HETSYN     NAG NAG
FORMUL   2  NAG    2(C8 H15 N1 O6)
FORMUL   4  PE7    C14 H30 O7 S1
FORMUL   5  HOH   *623(H2 O1)
HELIX    1   1 VAL A   40  ARG A   44  5                                   5
HELIX    2   2 PHE A   78  MET A   83  1                                   6
HELIX    3   3 LEU A  127  ASN A  131  5                                   5
HELIX    4   4 GLY A  132  GLU A  140  1                                   9
HELIX    5   5 VAL A  150  LEU A  156  1                                   7
HELIX    6   6 ASN A  167  ILE A  184  1                                  18
HELIX    7   7 GLN A  185  PHE A  187  5                                   3
HELIX    8   8 SER A  200  SER A  212  1                                  13
HELIX    9   9 SER A  212  ASP A  217  1                                   6
HELIX   10  10 VAL A  238  LEU A  252  1                                  15
HELIX   11  11 SER A  258  LYS A  269  1                                  12
HELIX   12  12 LYS A  270  GLU A  278  1                                   9
HELIX   13  13 TRP A  279  LEU A  282  5                                   4
HELIX   14  14 SER A  304  GLY A  312  1                                   9
HELIX   15  15 GLY A  328  ALA A  336  1                                   9
HELIX   16  16 SER A  348  VAL A  360  1                                  13
HELIX   17  17 ASN A  364  THR A  376  1                                  13
HELIX   18  18 ASN A  383  VAL A  400  1                                  18
HELIX   19  19 VAL A  400  LYS A  413  1                                  14
HELIX   20  20 PRO A  433  GLY A  437  5                                   5
HELIX   21  21 GLU A  443  PHE A  448  1                                   6
HELIX   22  22 GLY A  449  ASN A  457  5                                   9
HELIX   23  23 THR A  459  GLY A  480  1                                  22
HELIX   24  24 ARG A  517  GLN A  526  1                                  10
HELIX   25  25 GLN A  526  ALA A  534  1                                   9
SHEET    1   A 3 LEU A   7  THR A  10  0
SHEET    2   A 3 GLY A  13  MET A  16 -1  O  VAL A  15   N  VAL A   8
SHEET    3   A 3 VAL A  57  ASN A  59  1  O  TRP A  58   N  MET A  16
SHEET    1   B11 THR A  18  VAL A  22  0
SHEET    2   B11 SER A  25  PRO A  34 -1  O  ILE A  27   N  VAL A  20
SHEET    3   B11 TYR A  96  VAL A 101 -1  O  ILE A  99   N  PHE A  30
SHEET    4   B11 VAL A 142  SER A 145 -1  O  LEU A 143   N  TRP A 100
SHEET    5   B11 THR A 109  ILE A 115  1  N  TRP A 114   O  VAL A 144
SHEET    6   B11 GLY A 189  GLU A 199  1  O  THR A 195   N  VAL A 113
SHEET    7   B11 ARG A 221  GLN A 225  1  O  GLN A 225   N  GLY A 198
SHEET    8   B11 GLN A 318  ASN A 324  1  O  LEU A 320   N  LEU A 224
SHEET    9   B11 GLY A 417  PHE A 423  1  O  PHE A 423   N  VAL A 323
SHEET   10   B11 LYS A 501  LEU A 505  1  O  LEU A 505   N  PHE A 422
SHEET   11   B11 VAL A 512  GLN A 514 -1  O  HIS A 513   N  PHE A 502
SHEET    1   C 2 VAL A 236  SER A 237  0
SHEET    2   C 2 VAL A 295  ILE A 296  1  O  ILE A 296   N  VAL A 236
SSBOND   1 CYS A   67    CYS A   94
SSBOND   2 CYS A  254    CYS A  265
SSBOND   3 CYS A  402    CYS A  521
LINK         ND2 ASN A  59                 C1  NAG  9010
LINK         ND2 ASN A 416                 C1  NAG  9020
CISPEP   1 SER A  103    PRO A  104          0        -0.09
CRYST1  111.200  111.200  137.119  90.00  90.00 120.00 P 31 2 1      6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008993  0.005192  0.000000        0.00000
SCALE2      0.000000  0.010384  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007293        0.00000
END