longtext: 1QO9-pdb

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HEADER    HYDROLASE                               07-NOV-99   1QO9
TITLE     NATIVE ACETYLCHOLINESTERASE FROM DROSOPHILA MELANOGASTER
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYLCHOLINESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.1.1.7;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: DROSOPHILA MELANOGASTER;
SOURCE   3 ORGANISM_COMMON: FRUIT FLY;
SOURCE   4 EXPRESSION_SYSTEM: DROSOPHILA MELANOGASTER;
SOURCE   5 EXPRESSION_SYSTEM_STRAIN: SCHNEIDER LINE 2;
SOURCE   6 EXPRESSION_SYSTEM_CELLULAR_LOCATION: SECRETED
SOURCE   7 (EXTRACELLULAR);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID DNA;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: S2-SEC 1/3;
SOURCE  10 EXPRESSION_SYSTEM_GENE: DMACHE
KEYWDS    HYDROLASE (SERINE ESTERASE), HYDROLASE, SERINE ESTERASE,
KEYWDS   2 SYNAPSE, MEMBRANE, NERVE, MUSCLE, SIGNAL, NEUROTRANSMITTER
KEYWDS   3 DEGRADATION, GLYCOPROTEIN, GPI-ANCHOR, ALTERNATIVE SPLICING
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.HAREL,I.SILMAN,J.L.SUSSMAN
REVDAT   1   20-JUL-00 1QO9    0
JRNL        AUTH   M.HAREL,G.KRYGER,T.L.ROSENBERRY,W.D.MALLENDER,
JRNL        AUTH 2 T.LEWIS,R.J.FLETCHER,J.M.GUSS,I.SILMAN,J.L.SUSSMAN
JRNL        TITL   THREE-DIMENSIONAL STRUCTURES OF DROSOPHILA
JRNL        TITL 2 MELANOGASTER ACETYLCHOLINESTERASE AND OF ITS
JRNL        TITL 3 COMPLEXES WITH TWO POTENT INHIBITORS
JRNL        REF    PROTEIN SCI.                  V.   9  1063 2000
JRNL        REFN   ASTM PRCIEI  US ISSN 0961-8368
REMARK   2
REMARK   2 RESOLUTION. 2.7  ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS    3.8
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,
REMARK   3                 GROSSE-KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,
REMARK   3                 PANNU,READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.7
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 30.
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.0
REMARK   3   OUTLIER CUTOFF HIGH (RMS(ABS(F))) : 0.0
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 95.7
REMARK   3   NUMBER OF REFLECTIONS             : 19343
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.258
REMARK   3   FREE R VALUE                     : 0.301
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 32
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.7
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.713
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 41
REMARK   3   BIN R VALUE           (WORKING SET) : 0.44
REMARK   3   BIN FREE R VALUE                    : 0.458
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4273
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 75
REMARK   3   SOLVENT ATOMS            : 144
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 53.8
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.009
REMARK   3   BOND ANGLES            (DEGREES) : 1.67
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : NULL
REMARK   3   KSOL        : NULL
REMARK   3   BSOL        : NULL
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 1QO9 COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  ON  8-NOV-1999.
REMARK 100 THE EBI ID CODE IS EBI-4331.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 4.6
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ELETTRA BEAMLINE 5.2R
REMARK 200  BEAMLINE                       : 5.2R
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : NULL
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 20200
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.7
REMARK 200  RESOLUTION RANGE LOW       (A) : 30.0
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NONE
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.1
REMARK 200  DATA REDUNDANCY                : 1.7
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : 0.05
REMARK 200   FOR THE DATA SET  : 33.5
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.7
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.8
REMARK 200  COMPLETENESS FOR SHELL     (%) : 98.8
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : 0.16
REMARK 200   FOR SHELL         : 5.8
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SIRAS
REMARK 200 SOFTWARE USED: CCP4
REMARK 200 STARTING MODEL: 2ACE
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): NULL
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP:  P 43 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,1/2+Z
REMARK 290       3555   1/2-Y,1/2+X,3/4+Z
REMARK 290       4555   1/2+Y,1/2-X,1/4+Z
REMARK 290       5555   1/2-X,1/2+Y,3/4-Z
REMARK 290       6555   1/2+X,1/2-Y,1/4-Z
REMARK 290       7555   Y,X,-Z
REMARK 290       8555   -Y,-X,1/2-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       79.50000
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       47.17000
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       47.17000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000      119.25000
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       47.17000
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       47.17000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000       39.75000
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       47.17000
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       47.17000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000      119.25000
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       47.17000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       47.17000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       39.75000
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000       79.50000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     VAL A     1
REMARK 465     ILE A     2
REMARK 465     ALA A   104
REMARK 465     ARG A   105
REMARK 465     LEU A   106
REMARK 465     ARG A   107
REMARK 465     HIS A   108
REMARK 465     GLY A   109
REMARK 465     ARG A   110
REMARK 465     GLY A   111
REMARK 465     ALA A   112
REMARK 465     ASN A   113
REMARK 465     GLY A   114
REMARK 465     GLY A   115
REMARK 465     GLU A   116
REMARK 465     HIS A   117
REMARK 465     PRO A   118
REMARK 465     ASN A   119
REMARK 465     GLY A   120
REMARK 465     LYS A   121
REMARK 465     GLN A   122
REMARK 465     ALA A   123
REMARK 465     ASP A   124
REMARK 465     THR A   125
REMARK 465     ASP A   126
REMARK 465     HIS A   127
REMARK 465     LEU A   128
REMARK 465     ILE A   129
REMARK 465     HIS A   130
REMARK 465     ASN A   131
REMARK 465     GLY A   132
REMARK 465     ASN A   133
REMARK 465     PRO A   134
REMARK 465     GLN A   135
REMARK 465     GLY A   575
REMARK 465     THR A   576
REMARK 465     CYS A   577
REMARK 465     ASP A   578
REMARK 465     GLY A   579
REMARK 465     ASP A   580
REMARK 465     SER A   581
REMARK 465     GLY A   582
REMARK 465     SER A   583
REMARK 465     LEU A   584
REMARK 465     ALA A   585
REMARK 465     SER A   586
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ARG A   4    NE   CZ   NH1  NH2
REMARK 470     ARG A  15    CZ   NH1  NH2
REMARK 470     GLU A  51    CD   OE1  OE2
REMARK 470     ARG A 216    NE   CZ   NH1  NH2
REMARK 470     GLU A 230    CD   OE1  OE2
REMARK 470     ASN A 291    CG   OD1  ND2
REMARK 470     ASN A 300    CG   OD1  ND2
REMARK 470     ASN A 422    CG   OD1  ND2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    GLU A  69   N   -  CA  -  C   ANGL. DEV. =  12.5 DEGREES
REMARK 500    GLU A  80   N   -  CA  -  C   ANGL. DEV. = -11.7 DEGREES
REMARK 500    ASN A 136   CA  -  C   -  N   ANGL. DEV. =  23.3 DEGREES
REMARK 500    ASN A 136   O   -  C   -  N   ANGL. DEV. = -28.4 DEGREES
REMARK 500    THR A 137   C   -  N   -  CA  ANGL. DEV. =  23.0 DEGREES
REMARK 500    GLY A 140   N   -  CA  -  C   ANGL. DEV. = -12.6 DEGREES
REMARK 500    ASP A 378   N   -  CA  -  C   ANGL. DEV. = -12.8 DEGREES
REMARK 500    TYR A 379   C   -  N   -  CA  ANGL. DEV. =  10.6 DEGREES
REMARK 500    THR A 386   N   -  CA  -  C   ANGL. DEV. = -10.6 DEGREES
REMARK 500    GLY A 421   N   -  CA  -  C   ANGL. DEV. = -11.1 DEGREES
REMARK 500    THR A 544   CB  -  CA  -  C   ANGL. DEV. =  14.7 DEGREES
REMARK 500    THR A 544   N   -  CA  -  CB  ANGL. DEV. = -12.2 DEGREES
REMARK 500    ASP A 545   N   -  CA  -  C   ANGL. DEV. = -11.5 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500
REMARK 500  O     SER A   306    O     ARG A   309               2.14
REMARK 500  O     ARG A   309    O     SER A   306               2.14
REMARK 500  O     ARG A   309    OG    SER A   310               2.01
REMARK 500  OG    SER A   310    O     ARG A   309               2.01
REMARK 500  O     TYR A   374    N  AA PHE A   376               2.16
REMARK 500  N  AA PHE A   376    O     TYR A   374               2.16
REMARK 500  O     ILE A   377    N     TYR A   379               1.88
REMARK 500  N     TYR A   379    O     ILE A   377               1.88
REMARK 500  OE1   GLU A   420    O     HOH      93               1.20
REMARK 500  OE2   GLU A   420    O     HOH      93               1.26
REMARK 500  OE1BB GLU A   453    O     HOH      97               2.19
REMARK 500  N     THR A   464    OG    SER A   543               2.09
REMARK 500  NH2   ARG A   508    OG1   THR A   544               1.95
REMARK 500  OE1   GLU A   527    O     HOH     111               2.19
REMARK 500  OG    SER A   543    N     THR A   464               2.09
REMARK 500  OG1   THR A   544    NH2   ARG A   508               1.95
REMARK 500  O     HOH      93    OE1   GLU A   420               1.20
REMARK 500  O     HOH      93    OE2   GLU A   420               1.26
REMARK 500  O     HOH      97    OE1BB GLU A   453               2.19
REMARK 500  O     HOH     111    OE1   GLU A   527               2.19
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500
REMARK 500    GLU A  81      108.96    -49.30
REMARK 500    SER A 310      129.73    -63.10
REMARK 500    ASP A 378       52.18    -48.32
REMARK 500    GLN A 524      121.33    139.28
REMARK 500    SER A 543       18.54    123.14
REMARK 500    ASP A 546       81.26    147.19
REMARK 500    ALA A 552       81.96    150.36
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 ASN A 136    THR A 137                  258.10
REMARK 525
REMARK 525 SOLVENT
REMARK 525 SOME ATOMS COULD NOT BE ASSIGNED TO THE OLIGOMER, BY ANY
REMARK 525 SYMMETRY OPERATION TO WITHIN  5.00 ANGSTROM
REMARK 525
REMARK 525  M RES CSSEQI     X       Y       Z   SYMM-TRANS CLOSEST
REMARK 525                                                   DIST.
REMARK 525  1 HOH   733     4.817  94.897  -6.675    0          5.5
REMARK 525  1 HOH   741    26.859  88.462  27.885    0          6.2
REMARK 525  1 HOH   739    17.281  95.976   3.712    0          5.1
REMARK 525  1 HOH   710    44.481  35.047  24.664  001  555     6.2
REMARK 525  1 HOH   670    17.476  85.651  15.802  001  555     5.0
REMARK 525  1 HOH   671    23.425  63.080 -24.190  001  555     5.3
REMARK 525  1 HOH   615    46.937  42.601  24.808  001  555     6.3
REMARK 525  1 HOH   725     2.654  51.656   1.138  001  555     5.1
REMARK 525  1 HOH   675    13.189  37.374 -13.398  001  555     6.4
REMARK 525  1 HOH   649    47.719  39.713  24.924  001  555     7.6
REMARK 525
REMARK 525 THE FOLLOWING SOLVENT MOLECULES LIE FARTHER THAN EXPECTED
REMARK 525 FROM THE PROTEIN OR NUCLEIC ACID MOLECULE AND MAY BE
REMARK 525 ASSOCIATED WITH A SYMMETRY RELATED MOLECULE (M=MODEL
REMARK 525 NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525 THESE MOLECULES CAN BE PLACED WITHIN  5.00 ANGSTROM OF THE
REMARK 525 OBSERVED OLIGOMER BY APPLYING THE SYMMETRY TRANSFORMATION
REMARK 525 INDICATED.
REMARK 525
REMARK 525  M RES CSSEQI     ORIGINAL COORDINATES   SYMMETRY TRANS.    DIST.
REMARK 525                    X       Y       Z
REMARK 525  1 HOH   666    29.257  48.719  33.598    003      554      2.88
REMARK 525  1 HOH   717    36.384  39.923 -19.916    007      555      4.35
REMARK 525  1 HOH   657    24.810  40.102 -26.978    005      554      4.62
REMARK 525  1 HOH   737    26.527  50.696  38.181    008      665      1.20
REMARK 525  1 HOH   677    34.764  38.434 -16.159    007      555      4.33
REMARK 600
REMARK 600 HETEROGEN
REMARK 600    ALL BUT ONE OF THE SUGAR RESIDUES FORM A SINGLE GLYCOSYLATION
REMARK 600    UNIT:
REMARK 600
REMARK 600  ASN 493
REMARK 600     (1,N)
REMARK 600  N-ACETYL-D-GLUCOSAMINE            [NAG 991]
REMARK 600        (1,4)
REMARK 600     N-ACETYL-D-GLUCOSAMINE         [NAG 993]
REMARK 600           (1,4)
REMARK 600        BETA-D-MANNOSE              [BMA 994]
REMARK 600              (1,3)
REMARK 600           BETA-D-MANNOSE           [BMA 996]
REMARK 600                  (1,3)
REMARK 600               BETA-D-MANNOSE       [BMA 997]
REMARK 600
DBREF  1QO9 A    1   586  SWS    P07140   ACES_DROME      39    612
SEQRES   1 A  586  VAL ILE ASP ARG LEU VAL VAL GLN THR SER SER GLY PRO
SEQRES   2 A  586  VAL ARG GLY ARG SER VAL THR VAL GLN GLY ARG GLU VAL
SEQRES   3 A  586  HIS VAL TYR THR GLY ILE PRO TYR ALA LYS PRO PRO VAL
SEQRES   4 A  586  GLU ASP LEU ARG PHE ARG LYS PRO VAL PRO ALA GLU PRO
SEQRES   5 A  586  TRP HIS GLY VAL LEU ASP ALA THR GLY LEU SER ALA THR
SEQRES   6 A  586  CYS VAL GLN GLU ARG TYR GLU TYR PHE PRO GLY PHE SER
SEQRES   7 A  586  GLY GLU GLU ILE TRP ASN PRO ASN THR ASN VAL SER GLU
SEQRES   8 A  586  ASP CYS LEU TYR ILE ASN VAL TRP ALA PRO ALA LYS ALA
SEQRES   9 A  586  ARG LEU ARG HIS GLY ARG GLY ALA ASN GLY GLY GLU HIS
SEQRES  10 A  586  PRO ASN GLY LYS GLN ALA ASP THR ASP HIS LEU ILE HIS
SEQRES  11 A  586  ASN GLY ASN PRO GLN ASN THR THR ASN GLY LEU PRO ILE
SEQRES  12 A  586  LEU ILE TRP ILE TYR GLY GLY GLY PHE MET THR GLY SER
SEQRES  13 A  586  ALA THR LEU ASP ILE TYR ASN ALA ASP ILE MET ALA ALA
SEQRES  14 A  586  VAL GLY ASN VAL ILE VAL ALA SER PHE GLN TYR ARG VAL
SEQRES  15 A  586  GLY ALA PHE GLY PHE LEU HIS LEU ALA PRO GLU MET PRO
SEQRES  16 A  586  SER GLU PHE ALA GLU GLU ALA PRO GLY ASN VAL GLY LEU
SEQRES  17 A  586  TRP ASP GLN ALA LEU ALA ILE ARG TRP LEU LYS ASP ASN
SEQRES  18 A  586  ALA HIS ALA PHE GLY GLY ASN PRO GLU TRP MET THR LEU
SEQRES  19 A  586  PHE GLY GLU SER ALA GLY SER SER SER VAL ASN ALA GLN
SEQRES  20 A  586  LEU MET SER PRO VAL THR ARG GLY LEU VAL LYS ARG GLY
SEQRES  21 A  586  MET MET GLN SER GLY THR MET ASN ALA PRO TRP SER HIS
SEQRES  22 A  586  MET THR SER GLU LYS ALA VAL GLU ILE GLY LYS ALA LEU
SEQRES  23 A  586  ILE ASN ASP CYS ASN CYS ASN ALA SER MET LEU LYS THR
SEQRES  24 A  586  ASN PRO ALA HIS VAL MET SER CYS MET ARG SER VAL ASP
SEQRES  25 A  586  ALA LYS THR ILE SER VAL GLN GLN TRP ASN SER TYR SER
SEQRES  26 A  586  GLY ILE LEU SER PHE PRO SER ALA PRO THR ILE ASP GLY
SEQRES  27 A  586  ALA PHE LEU PRO ALA ASP PRO MET THR LEU MET LYS THR
SEQRES  28 A  586  ALA ASP LEU LYS ASP TYR ASP ILE LEU MET GLY ASN VAL
SEQRES  29 A  586  ARG ASP GLU GLY THR TYR PHE LEU LEU TYR ASP PHE ILE
SEQRES  30 A  586  ASP TYR PHE ASP LYS ASP ASP ALA THR ALA LEU PRO ARG
SEQRES  31 A  586  ASP LYS TYR LEU GLU ILE MET ASN ASN ILE PHE GLY LYS
SEQRES  32 A  586  ALA THR GLN ALA GLU ARG GLU ALA ILE ILE PHE GLN TYR
SEQRES  33 A  586  THR SER TRP GLU GLY ASN PRO GLY TYR GLN ASN GLN GLN
SEQRES  34 A  586  GLN ILE GLY ARG ALA VAL GLY ASP HIS PHE PHE THR CYS
SEQRES  35 A  586  PRO THR ASN GLU TYR ALA GLN ALA LEU ALA GLU ARG GLY
SEQRES  36 A  586  ALA SER VAL HIS TYR TYR TYR PHE THR HIS ARG THR SER
SEQRES  37 A  586  THR SER LEU TRP GLY GLU TRP MET GLY VAL LEU HIS GLY
SEQRES  38 A  586  ASP GLU ILE GLU TYR PHE PHE GLY GLN PRO LEU ASN ASN
SEQRES  39 A  586  SER LEU GLN TYR ARG PRO VAL GLU ARG GLU LEU GLY LYS
SEQRES  40 A  586  ARG MET LEU SER ALA VAL ILE GLU PHE ALA LYS THR GLY
SEQRES  41 A  586  ASN PRO ALA GLN ASP GLY GLU GLU TRP PRO ASN PHE SER
SEQRES  42 A  586  LYS GLU ASP PRO VAL TYR TYR ILE PHE SER THR ASP ASP
SEQRES  43 A  586  LYS ILE GLU LYS LEU ALA ARG GLY PRO LEU ALA ALA ARG
SEQRES  44 A  586  CYS SER PHE TRP ASN ASP TYR LEU PRO LYS VAL ARG SER
SEQRES  45 A  586  TRP ALA GLY THR CYS ASP GLY ASP SER GLY SER LEU ALA
SEQRES  46 A  586  SER
HET    NAG  A 991      14
HET    NAG  A 992      14
HET    NAG  A 993      14
HET    BMA  A 994      11
HET    BMA  A 996      11
HET    BMA  A 997      11
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     BMA BETA-D-MANNOSE
FORMUL   2  NAG    3(C8 H15 N1 O6)
FORMUL   3  BMA    3(C6 H11 O5)
FORMUL   4  HOH   *144(H2 O1)
HELIX    1   1 VAL A   39  ARG A   43  5                                   5
HELIX    2   2 LEU A  159  ASN A  163  5                                   5
HELIX    3   3 ALA A  164  ASN A  172  1                                   9
HELIX    4   4 GLY A  183  LEU A  188  1                                   6
HELIX    5   5 PRO A  195  ALA A  199  5                                   5
HELIX    6   6 ASN A  205  ALA A  222  1                                  18
HELIX    7   7 HIS A  223  PHE A  225  5                                   3
HELIX    8   8 SER A  238  SER A  250  1                                  13
HELIX    9   9 THR A  275  ASN A  288  1                                  14
HELIX   10  10 ASN A  300  ARG A  309  1                                  10
HELIX   11  11 ASP A  312  TRP A  321  1                                  10
HELIX   12  12 ASP A  344  THR A  351  1                                   8
HELIX   13  13 GLY A  368  LEU A  373  1                                   6
HELIX   14  14 PRO A  389  PHE A  401  1                                  13
HELIX   15  15 THR A  405  TYR A  416  1                                  12
HELIX   16  16 GLY A  424  PHE A  440  1                                  17
HELIX   17  17 PHE A  440  ARG A  454  1                                  15
HELIX   18  18 GLY A  473  GLY A  477  5                                   5
HELIX   19  19 GLY A  481  PHE A  488  1                                   8
HELIX   20  20 GLY A  489  ASN A  493  5                                   5
HELIX   21  21 VAL A  501  GLY A  520  1                                  20
HELIX   22  22 ARG A  553  ASP A  565  1                                  13
HELIX   23  23 TYR A  566  SER A  572  1                                   7
SHEET    1   A 2 VAL A   6  THR A   9  0
SHEET    2   A 2 GLY A  12  ARG A  15 -1  N  VAL A  14   O  VAL A   7
SHEET    1   B 9 ARG A  17  VAL A  21  0
SHEET    2   B 9 ARG A  24  TYR A  29 -1  N  VAL A  28   O  ARG A  17
SHEET    3   B 9 ILE A  96  PRO A 101 -1  N  ALA A 100   O  HIS A  27
SHEET    4   B 9 ILE A 174  PHE A 178 -1  N  SER A 177   O  ASN A  97
SHEET    5   B 9 ILE A 143  ILE A 147  1  N  LEU A 144   O  ILE A 174
SHEET    6   B 9 MET A 232  GLU A 237  1  N  THR A 233   O  ILE A 143
SHEET    7   B 9 ARG A 259  GLN A 263  1  N  MET A 261   O  LEU A 234
SHEET    8   B 9 ASP A 358  VAL A 364  1  N  ASP A 358   O  GLY A 260
SHEET    9   B 9 SER A 457  PHE A 463  1  N  SER A 457   O  ILE A 359
SSBOND   1 CYS A   66    CYS A   93
SSBOND   2 CYS A  292    CYS A  307
SSBOND   3 CYS A  442    CYS A  560
LINK         ND2 ASN A  88                 C1  NAG A 992
LINK         ND2 ASN A 493                 C1  NAG A 991
LINK         O4  NAG A 991                 C1  NAG A 993
LINK         O4  NAG A 993                 C1  BMA A 994
LINK         O3  BMA A 994                 C1  BMA A 996
LINK         O3  BMA A 996                 C1  BMA A 997
CRYST1   94.340   94.340  159.000  90.00  90.00  90.00 P 43 21 2     6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.010600  0.000000  0.000000        0.00000
SCALE2      0.000000  0.010600  0.000000        0.00000
SCALE3      0.000000  0.000000  0.006289        0.00000
END