longtext: 2AXE-pdb

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HEADER    HYDROLASE                               01-SEP-98   2AXE
TITLE     IODINATED COMPLEX OF ACETYL XYLAN ESTERASE AT 1.80 ANGSTROMS
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYL XYLAN ESTERASE;
COMPND   3 CHAIN: NULL;
COMPND   4 EC: 3.1.1.6;
COMPND   5 BIOLOGICAL_UNIT: MONOMER
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PENICILLIUM PURPUROGENUM
KEYWDS    HYDROLASE, IODOTYROSINES, ESTERASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    D.GHOSH,M.ERMAN,M.W.SAWICKI,P.LALA,D.R.WEEKS,N.LI,
AUTHOR   2 W.PANGBORN,D.J.THIEL,H.JORNVALL,J.EYZAGUIRRE
REVDAT   1   18-MAY-99 2AXE    0
JRNL        AUTH   D.GHOSH,M.ERMAN,M.SAWICKI,P.LALA,D.R.WEEKS,N.LI,
JRNL        AUTH 2 W.PANGBORN,D.J.THIEL,H.JORNVALL,R.GUTIERREZ,
JRNL        AUTH 3 J.EYZAGUIRRE
JRNL        TITL   DETERMINATION OF A PROTEIN STRUCTURE BY IODINATION:
JRNL        TITL 2 THE STRUCTURE OF IODINATED ACETYLXYLAN ESTERASE
JRNL        REF    ACTA CRYSTALLOGR.,SECT.D      V.  55   779 1999
JRNL        REFN   ASTM ABCRE6  DK ISSN 0907-4449                 0766
REMARK   1
REMARK   2
REMARK   2 RESOLUTION. 1.80 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : SHELXL-97
REMARK   3   AUTHORS     : G.M.SHELDRICK
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.80
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 99.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.0
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.9
REMARK   3   CROSS-VALIDATION METHOD           : FREE R
REMARK   3   FREE R VALUE TEST SET SELECTION   : RANDOM
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (NO CUTOFF).
REMARK   3   R VALUE   (WORKING + TEST SET, NO CUTOFF) : 0.1733
REMARK   3   R VALUE          (WORKING SET, NO CUTOFF) : 0.1768
REMARK   3   FREE R VALUE                  (NO CUTOFF) : 0.2369
REMARK   3   FREE R VALUE TEST SET SIZE (%, NO CUTOFF) : 5.1
REMARK   3   FREE R VALUE TEST SET COUNT   (NO CUTOFF) : 763
REMARK   3   TOTAL NUMBER OF REFLECTIONS   (NO CUTOFF) : 15003
REMARK   3
REMARK   3  FIT/AGREEMENT OF MODEL FOR DATA WITH F>4SIG(F).
REMARK   3   R VALUE   (WORKING + TEST SET, F>4SIG(F)) : 0.1490
REMARK   3   R VALUE          (WORKING SET, F>4SIG(F)) : 0.1523
REMARK   3   FREE R VALUE                  (F>4SIG(F)) : 0.2137
REMARK   3   FREE R VALUE TEST SET SIZE (%, F>4SIG(F)) : 5.2
REMARK   3   FREE R VALUE TEST SET COUNT   (F>4SIG(F)) : 594
REMARK   3   TOTAL NUMBER OF REFLECTIONS   (F>4SIG(F)) : 11490
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS      : 1422
REMARK   3   NUCLEIC ACID ATOMS : 0
REMARK   3   HETEROGEN ATOMS    : 33
REMARK   3   SOLVENT ATOMS      : 91
REMARK   3
REMARK   3  MODEL REFINEMENT.
REMARK   3   OCCUPANCY SUM OF NON-HYDROGEN ATOMS      : 1540.25
REMARK   3   OCCUPANCY SUM OF HYDROGEN ATOMS          : 1182.00
REMARK   3   NUMBER OF DISCRETELY DISORDERED RESIDUES : 2
REMARK   3   NUMBER OF LEAST-SQUARES PARAMETERS       : 6189
REMARK   3   NUMBER OF RESTRAINTS                     : 6009
REMARK   3
REMARK   3  RMS DEVIATIONS FROM RESTRAINT TARGET VALUES.
REMARK   3   BOND LENGTHS                         (A) : 0.013
REMARK   3   ANGLE DISTANCES                      (A) : 0.0578
REMARK   3   SIMILAR DISTANCES (NO TARGET VALUES) (A) : NULL
REMARK   3   DISTANCES FROM RESTRAINT PLANES      (A) : NULL
REMARK   3   ZERO CHIRAL VOLUMES               (A**3) : NULL
REMARK   3   NON-ZERO CHIRAL VOLUMES           (A**3) : NULL
REMARK   3   ANTI-BUMPING DISTANCE RESTRAINTS     (A) : NULL
REMARK   3   RIGID-BOND ADP COMPONENTS         (A**2) : NULL
REMARK   3   SIMILAR ADP COMPONENTS            (A**2) : NULL
REMARK   3   APPROXIMATELY ISOTROPIC ADPS      (A**2) : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED: MOEWS & KRETSINGER, J.MOL.BIOL.91(1973)201-2
REMARK   3
REMARK   3  STEREOCHEMISTRY TARGET VALUES : ENGH AND HUBER
REMARK   3   SPECIAL CASE: NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: C-I BONDS WERE NOT RESTRAINED
REMARK   4
REMARK   4 2AXE COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK   6
REMARK   6 TYROSINES 33 AND 177 ARE IODOTYROSINES.  THREE C-I
REMARK   6 DISTANCES RANGE BETWEEN 2.09 AND 2.12 ANGSTROMS.  ONE C-I
REMARK   6 DISTANCE FOR TYR 33 IS 2.43 ANGSTROMS.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : DEC-1997
REMARK 200  TEMPERATURE           (KELVIN) : 287
REMARK 200  PH                             : 6.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : NULL
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU RU200
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418
REMARK 200  MONOCHROMATOR                  : GRAPHITE
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : NULL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 15046
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.80
REMARK 200  RESOLUTION RANGE LOW       (A) : 99.00
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NONE
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.8
REMARK 200  DATA REDUNDANCY                : 6.7
REMARK 200  R MERGE                    (I) : 0.11
REMARK 200  R SYM                      (I) : NONE
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.80
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.90
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: NULL
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SIRAS
REMARK 200 SOFTWARE USED: SHELX
REMARK 200 STARTING MODEL: NONE
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 22.9
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.60
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   1/2-X,-Y,1/2+Z
REMARK 290       3555   -X,1/2+Y,1/2-Z
REMARK 290       4555   1/2+X,1/2-Y,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       17.45018
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       36.26868
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       30.63913
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       36.26868
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       17.45018
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       30.63913
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 400
REMARK 400 COMPOUND
REMARK 400 TYR 177 IS A DIIODO- AND TYR 33 IS PARTIALLY A MONO- AND
REMARK 400 PARTIALLY A DIIODO-TYROSINE.  THEREFORE, THERE ARE 4 IODINE
REMARK 400 ATOMS IN THE COORDINATE FILE.
REMARK 450
REMARK 450 SOURCE
REMARK 450 THE PROTEIN WAS NON-ENZYMATICALLY IODINATED IN THE
REMARK 450 CRYSTALLINE STATE BY ADDING A SOLUTION OF MOLECULAR IODINE
REMARK 450 IN KI TO THE CRYSTAL.
REMARK 600
REMARK 600 HETEROGEN
REMARK 600 IODOTYROSINES 33 AND 177 ARE NAMED TYI
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: CAT
REMARK 800 SITE_DESCRIPTION:
REMARK 800 THESE RESIDUES FORM THE CATALYTIC TRIAD.
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 2AXE       GB      2944191      1 -    27 NOT IN ATOMS LIST
DBREF  2AXE      1   207  GB     2944191  AAC39371        28    234
SEQADV 2AXE TYI     33  GB   2944191   TYR    60 MODIFIED
SEQADV 2AXE TYI    177  GB   2944191   TYR   204 MODIFIED
SEQRES   1    207  SER CYS PRO ALA ILE HIS VAL PHE GLY ALA ARG GLU THR
SEQRES   2    207  THR ALA SER PRO GLY TYR GLY SER SER SER THR VAL VAL
SEQRES   3    207  ASN GLY VAL LEU SER ALA TYI PRO GLY SER THR ALA GLU
SEQRES   4    207  ALA ILE ASN TYR PRO ALA CYS GLY GLY GLN SER SER CYS
SEQRES   5    207  GLY GLY ALA SER TYR SER SER SER VAL ALA GLN GLY ILE
SEQRES   6    207  ALA ALA VAL ALA SER ALA VAL ASN SER PHE ASN SER GLN
SEQRES   7    207  CYS PRO SER THR LYS ILE VAL LEU VAL GLY TYR SER GLN
SEQRES   8    207  GLY GLY GLU ILE MET ASP VAL ALA LEU CYS GLY GLY GLY
SEQRES   9    207  ASP PRO ASN GLN GLY TYR THR ASN THR ALA VAL GLN LEU
SEQRES  10    207  SER SER SER ALA VAL ASN MET VAL LYS ALA ALA ILE PHE
SEQRES  11    207  MET GLY ASP PRO MET PHE ARG ALA GLY LEU SER TYR GLU
SEQRES  12    207  VAL GLY THR CYS ALA ALA GLY GLY PHE ASP GLN ARG PRO
SEQRES  13    207  ALA GLY PHE SER CYS PRO SER ALA ALA LYS ILE LYS SER
SEQRES  14    207  TYR CYS ASP ALA SER ASP PRO TYI CYS CYS ASN GLY SER
SEQRES  15    207  ASN ALA ALA THR HIS GLN GLY TYR GLY SER GLU TYR GLY
SEQRES  16    207  SER GLN ALA LEU ALA PHE VAL LYS SER LYS LEU GLY
MODRES 2AXE TYI     33  TYR  3,5-DIIODOTRYROSINE
MODRES 2AXE TYI    177  TYR  3,5-DIIODOTRYROSINE
HET    TYI     33      14
HET    TYI    177      14
HET    SO4    209       5
HETNAM     TYI 3,5-DIIODOTRYROSINE
HETNAM     SO4 SULFATE ION
FORMUL   1  TYI    2(C9 H9 N1 O3 I2)
FORMUL   2  SO4    O4 S1 2-
FORMUL   3  HOH   *91(H2 O1)
HELIX    1   1 GLY     20  SER     31  5                                  12
HELIX    2   2 SER     50  CYS     52  5                                   3
HELIX    3   3 TYR     57  GLN     78  1                                  22
HELIX    4   4 SER     90  CYS    101  5                                  12
HELIX    5   5 PRO    106  GLN    108  5                                   3
HELIX    6   6 SER    119  MET    124  1                                   6
HELIX    7   7 ALA    164  LYS    166  5                                   3
HELIX    8   8 ALA    184  GLN    188  1                                   5
HELIX    9   9 TYR    190  LYS    205  1                                  16
SHEET    1   A 5 THR    37  ALA    40  0
SHEET    2   A 5 ILE     5  ALA    10  1  N  VAL     7   O  THR    37
SHEET    3   A 5 LYS    83  TYR    89  1  N  LYS    83   O  HIS     6
SHEET    4   A 5 VAL   125  MET   131  1  N  LYS   126   O  ILE    84
SHEET    5   A 5 ILE   167  TYR   170  1  N  LYS   168   O  ALA   128
SSBOND   1 CYS      2    CYS     79
SSBOND   2 CYS     46    CYS     52
SSBOND   3 CYS    101    CYS    161
SSBOND   4 CYS    147    CYS    179
SSBOND   5 CYS    171    CYS    178
LINK         N   TYI    33                 C   ALA    32
LINK         C   TYI    33                 N   PRO    34
LINK         N   TYI   177                 C   PRO   176
LINK         C   TYI   177                 N   CYS   178
SITE     1 CAT  3 SER    90  ASP   175  HIS   187
CRYST1   34.900   61.278   72.538  90.00  90.00  90.00 P 21 21 21    4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.028653  0.000000  0.000000        0.00000
SCALE2      0.000000  0.016319  0.000000        0.00000
SCALE3      0.000000  0.000000  0.013786        0.00000
END