longtext: 2BUC-pdb

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HEADER    COMPLEX (HYDROLASE/INHIBITOR)           09-JUN-05   2BUC
TITLE     CRYSTAL STRUCTURE OF PORCINE DIPEPTIDYL PEPDIDASE IV (CD26)
TITLE    2 IN COMPLEX WITH A TETRAHYDROISOQUINOLINE INHIBITOR
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: DIPEPTIDYL PEPTIDASE IV;
COMPND   3 SYNONYM: DPP IV, T-CELL ACTIVATION ANTIGEN CD26;
COMPND   4 CHAIN: A, B, C, D;
COMPND   5 FRAGMENT: EXTRACELLULAR DOMAIN, RESIDUES 39-766;
COMPND   6 EC: 3.4.14.5
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: SUS SCROFA;
SOURCE   3 ORGANISM_COMMON: PIG;
SOURCE   4 ORGAN: KIDNEY CORTEX
KEYWDS    DPP-IV, DIABETES MELLITUS, DRUG DESIGN, HYDROLASE, SERINE
KEYWDS   2 PROTEASE, AMINOPEPTIDASE, GLYCOPROTEIN, PROTEASE,
KEYWDS   3 SIGNAL-ANCHOR, TRANSMEMBRANE, COMPLEX
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.NORDHOFF,S.CEREZO-GALVEZ,A.FEURER,O.HILL,V.G.MATASSA,
AUTHOR   2 G.METZ,C.RUMMEY,M.THIEMANN,P.J.EDWARDS
REVDAT   1   23-JAN-06 2BUC    0
JRNL        AUTH   S.NORDHOFF,S.CEREZO-GALVEZ,A.FEURER,O.HILL,
JRNL        AUTH 2 V.G.MATASSA,G.METZ,C.RUMMEY,M.THIEMANN,P.J.EDWARDS
JRNL        TITL   THE REVERSED BINDING OF B-PHENETHYLAMINE
JRNL        TITL 2 INHIBITORS OF DPP-IV. X-RAY STRUCTURES AND
JRNL        TITL 3 PROPERTIES OF NOVEL FRAGMENT AND ELABORATED
JRNL        TITL 4 INHIBITORS.
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION. 2.5  ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.1.24
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3   REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.50
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.28
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NONE
REMARK   3   COMPLETENESS FOR RANGE        (%) : 100.00
REMARK   3   NUMBER OF REFLECTIONS             : 113230
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.22208
REMARK   3   R VALUE            (WORKING SET) : 0.22094
REMARK   3   FREE R VALUE                     : 0.27872
REMARK   3   FREE R VALUE TEST SET SIZE   (%) :  2.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 2311
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.500
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.564
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 8268
REMARK   3   BIN R VALUE           (WORKING SET) : 0.300
REMARK   3   BIN FREE R VALUE SET COUNT          : 168
REMARK   3   BIN FREE R VALUE                    : 0.324
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 23864
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 606
REMARK   3   SOLVENT ATOMS            : 951
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 54.498
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.68
REMARK   3    B22 (A**2) : -0.39
REMARK   3    B33 (A**2) : 0.08
REMARK   3    B12 (A**2) : 0.45
REMARK   3    B13 (A**2) : -0.98
REMARK   3    B23 (A**2) : 0.67
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.825
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.332
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.268
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 12.168
REMARK   3
REMARK   3  CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.937
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.891
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES    COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED           (A): 25219 ; 0.005 ; 0.021
REMARK   3   BOND LENGTHS OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED      (DEGREES): 34354 ; 0.908 ; 1.950
REMARK   3   BOND ANGLES OTHERS       (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1 (DEGREES):  2908 ; 4.985 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2 (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 3 (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 4 (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   CHIRAL-CENTER RESTRAINTS    (A**3):  3657 ; 0.061 ; 0.200
REMARK   3   GENERAL PLANES REFINED         (A): 19360 ; 0.002 ; 0.020
REMARK   3   GENERAL PLANES OTHERS          (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED    (A): 12049 ; 0.162 ; 0.200
REMARK   3   NON-BONDED CONTACTS OTHERS     (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED     (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED         (A):  1455 ; 0.105 ; 0.200
REMARK   3   SYMMETRY VDW REFINED           (A):   127 ; 0.147 ; 0.200
REMARK   3   SYMMETRY VDW OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED        (A):     7 ; 0.117 ; 0.200
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS. COUNT  RMS   WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED   (A**2): 14512 ; 0.269 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE REFINED  (A**2): 23548 ; 0.500 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED   (A**2): 10707 ; 0.483 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED  (A**2): 10806 ; 0.864 ; 4.500
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF NCS GROUPS : 1
REMARK   3
REMARK   3  NCS GROUP NUMBER               :   1
REMARK   3     CHAIN NAMES                    : A B C D
REMARK   3     NUMBER OF COMPONENTS NCS GROUP :    3
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE
REMARK   3           1     A     39       A     540      5
REMARK   3           1     B     39       B     540      5
REMARK   3           1     C     39       C     540      5
REMARK   3           1     D     39       D     540      5
REMARK   3           2     A    555       A     620      5
REMARK   3           2     B    555       B     620      5
REMARK   3           2     C    555       C     620      5
REMARK   3           2     D    555       D     620      5
REMARK   3           3     A    640       A     766      5
REMARK   3           3     B    640       B     766      5
REMARK   3           3     C    640       C     766      5
REMARK   3           3     D    640       D     766      5
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT
REMARK   3   MEDIUM POSITIONAL  1    A    (A):   3290 ;  0.18 ;  0.50
REMARK   3   MEDIUM POSITIONAL  1    B    (A):   3290 ;  0.15 ;  0.50
REMARK   3   MEDIUM POSITIONAL  1    C    (A):   3290 ;  0.18 ;  0.50
REMARK   3   MEDIUM POSITIONAL  1    D    (A):   3290 ;  0.16 ;  0.50
REMARK   3   LOOSE POSITIONAL   1    A    (A):   2417 ;  0.45 ;  5.00
REMARK   3   LOOSE POSITIONAL   1    B    (A):   2417 ;  0.41 ;  5.00
REMARK   3   LOOSE POSITIONAL   1    C    (A):   2417 ;  0.47 ;  5.00
REMARK   3   LOOSE POSITIONAL   1    D    (A):   2417 ;  0.41 ;  5.00
REMARK   3   MEDIUM THERMAL     1    A (A**2):   3290 ;  0.14 ;  2.00
REMARK   3   MEDIUM THERMAL     1    B (A**2):   3290 ;  0.15 ;  2.00
REMARK   3   MEDIUM THERMAL     1    C (A**2):   3290 ;  0.15 ;  2.00
REMARK   3   MEDIUM THERMAL     1    D (A**2):   3290 ;  0.15 ;  2.00
REMARK   3   LOOSE THERMAL      1    A (A**2):   2417 ;  0.45 ; 10.00
REMARK   3   LOOSE THERMAL      1    B (A**2):   2417 ;  0.46 ; 10.00
REMARK   3   LOOSE THERMAL      1    C (A**2):   2417 ;  0.51 ; 10.00
REMARK   3   LOOSE THERMAL      1    D (A**2):   2417 ;  0.48 ; 10.00
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : BABINET MODEL WITH MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   :1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3   RIDING POSITIONS.
REMARK   4
REMARK   4 2BUC COMPLIES WITH FORMAT V. 2.3, 09-JULY-1998
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY EBI  ON 10-JUN-2005.
REMARK 100 THE EBI ID CODE IS EBI-24440.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SLS
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97954
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : NULL
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL
REMARK 200  DATA SCALING SOFTWARE          : NULL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 115542
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.50
REMARK 200  RESOLUTION RANGE LOW       (A) : 19.28
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NONE
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.1
REMARK 200  DATA REDUNDANCY                : 1.98
REMARK 200  R MERGE                    (I) : 0.20
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: OTHER
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NONE
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): NULL
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA):NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 THIS ENTRY CONTAINS THE CRYSTALLOGRAPHIC ASYMMETRIC UNIT
REMARK 300 WHICH CONSISTS OF   4 CHAIN(S). SEE REMARK 350 FOR
REMARK 300 INFORMATION ON GENERATING THE BIOLOGICAL MOLECULE(S).
REMARK 300
REMARK 300 QUATERNARY STRUCTURE FOR THIS ENTRY: DIMERIC
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE:  1
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350 BIOMOLECULE:  2
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES: ENGH AND HUBER, 1991
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    LEU A 415   CA  -  CB  -  CG  ANGL. DEV. =   9.4 DEGREES
REMARK 500    LEU A 542   CA  -  CB  -  CG  ANGL. DEV. =   6.7 DEGREES
REMARK 500    VAL A 711   N   -  CA  -  C   ANGL. DEV. =  -6.6 DEGREES
REMARK 500    LEU B 415   CA  -  CB  -  CG  ANGL. DEV. =  12.3 DEGREES
REMARK 500    LEU B 542   CA  -  CB  -  CG  ANGL. DEV. =   6.5 DEGREES
REMARK 500    LEU C 542   CA  -  CB  -  CG  ANGL. DEV. =   6.5 DEGREES
REMARK 500    VAL C 711   N   -  CA  -  C   ANGL. DEV. =  -6.9 DEGREES
REMARK 500    LEU D 415   CA  -  CB  -  CG  ANGL. DEV. =  11.1 DEGREES
REMARK 500    VAL D 711   N   -  CA  -  C   ANGL. DEV. =  -6.4 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A  40      131.22     68.00
REMARK 500    SER A 630     -113.91     57.88
REMARK 500    ARG B  40      125.12     72.43
REMARK 500    SER B 630     -104.36     52.04
REMARK 500    ARG C  40      130.52     75.83
REMARK 500    SER C 630     -114.02     55.26
REMARK 500    ARG D  40      122.30     69.03
REMARK 500    SER D 630     -111.56     56.69
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES ARE GIVEN CHAIN IDENTIFIERS TO
REMARK 525 INDICATE THE PROTEIN CHAIN TO WHICH THEY ARE MOST CLOSELY
REMARK 525 ASSOCIATED WITH:
REMARK 525   PROTEIN CHAIN  SOLVENT CHAIN
REMARK 525     A              Z
REMARK 525     B              Y
REMARK 525     C              X
REMARK 525     D              W
REMARK 700
REMARK 700 SHEET
REMARK 700 THE SHEET STRUCTURE OF THIS MOLECULE IS BIFURCATED. IN
REMARK 700 ORDER TO REPRESENT THIS FEATURE IN THE SHEET RECORDS BELOW,
REMARK 700 TWO SHEETS ARE DEFINED.
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 SITE_DESCRIPTION: 008 BINDING SITE FOR CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 SITE_DESCRIPTION: 008 BINDING SITE FOR CHAIN B
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 SITE_DESCRIPTION: 008 BINDING SITE FOR CHAIN C
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 SITE_DESCRIPTION: 008 BINDING SITE FOR CHAIN D
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN B
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN C
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN D
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN A
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN B
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN C
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 SITE_DESCRIPTION: SO4 BINDING SITE FOR CHAIN D
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 1ORV   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF PORCINE DIPEPTIDYL
REMARK 900  PEPTIDASE IV (CD26)
REMARK 900 RELATED ID: 1ORW   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF PORCINE DIPEPTIDYL
REMARK 900  PEPTIDASE IV (CD26)IN COMPLEX WITH A
REMARK 900  PEPTIDOMIMETIC INHIBITOR
REMARK 900 RELATED ID: 2BUA   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF PORCINE DIPEPTIDYL
REMARK 900  PEPTIDASE IV (CD26) IN COMPLEX WITH A LOW
REMARK 900   MOLECULAR WEIGHT INHIBITOR.
DBREF  2BUC A   39   766  UNP    P22411   DPP4_PIG        39    766
DBREF  2BUC B   39   766  UNP    P22411   DPP4_PIG        39    766
DBREF  2BUC C   39   766  UNP    P22411   DPP4_PIG        39    766
DBREF  2BUC D   39   766  UNP    P22411   DPP4_PIG        39    766
SEQRES   1 A  728  SER ARG ARG THR TYR THR LEU THR ASP TYR LEU LYS SER
SEQRES   2 A  728  THR PHE ARG VAL LYS PHE TYR THR LEU GLN TRP ILE SER
SEQRES   3 A  728  ASP HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU
SEQRES   4 A  728  LEU PHE ASN ALA GLU TYR GLY ASN SER SER ILE PHE LEU
SEQRES   5 A  728  GLU ASN SER THR PHE ASP GLU LEU GLY TYR SER THR ASN
SEQRES   6 A  728  ASP TYR SER VAL SER PRO ASP ARG GLN PHE ILE LEU PHE
SEQRES   7 A  728  GLU TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR
SEQRES   8 A  728  ALA SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU
SEQRES   9 A  728  ILE THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP ILE
SEQRES  10 A  728  THR TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP
SEQRES  11 A  728  ASN ASN ASP ILE TYR VAL LYS ASN GLU PRO ASN LEU SER
SEQRES  12 A  728  SER GLN ARG ILE THR TRP THR GLY LYS GLU ASN VAL ILE
SEQRES  13 A  728  TYR ASN GLY VAL THR ASP TRP VAL TYR GLU GLU GLU VAL
SEQRES  14 A  728  PHE SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY
SEQRES  15 A  728  THR PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL
SEQRES  16 A  728  PRO LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU
SEQRES  17 A  728  GLN TYR PRO LYS THR VAL ARG ILE PRO TYR PRO LYS ALA
SEQRES  18 A  728  GLY ALA GLU ASN PRO THR VAL LYS PHE PHE VAL VAL ASP
SEQRES  19 A  728  THR ARG THR LEU SER PRO ASN ALA SER VAL THR SER TYR
SEQRES  20 A  728  GLN ILE VAL PRO PRO ALA SER VAL LEU ILE GLY ASP HIS
SEQRES  21 A  728  TYR LEU CYS GLY VAL THR TRP VAL THR GLU GLU ARG ILE
SEQRES  22 A  728  SER LEU GLN TRP ILE ARG ARG ALA GLN ASN TYR SER ILE
SEQRES  23 A  728  ILE ASP ILE CYS ASP TYR ASP GLU SER THR GLY ARG TRP
SEQRES  24 A  728  ILE SER SER VAL ALA ARG GLN HIS ILE GLU ILE SER THR
SEQRES  25 A  728  THR GLY TRP VAL GLY ARG PHE ARG PRO ALA GLU PRO HIS
SEQRES  26 A  728  PHE THR SER ASP GLY ASN SER PHE TYR LYS ILE ILE SER
SEQRES  27 A  728  ASN GLU GLU GLY TYR LYS HIS ILE CYS HIS PHE GLN THR
SEQRES  28 A  728  ASP LYS SER ASN CYS THR PHE ILE THR LYS GLY ALA TRP
SEQRES  29 A  728  GLU VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU
SEQRES  30 A  728  TYR TYR ILE SER ASN GLU HIS LYS GLY MET PRO GLY GLY
SEQRES  31 A  728  ARG ASN LEU TYR ARG ILE GLN LEU ASN ASP TYR THR LYS
SEQRES  32 A  728  VAL THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS
SEQRES  33 A  728  GLN TYR TYR SER ALA SER PHE SER ASN LYS ALA LYS TYR
SEQRES  34 A  728  TYR GLN LEU ARG CYS PHE GLY PRO GLY LEU PRO LEU TYR
SEQRES  35 A  728  THR LEU HIS SER SER SER SER ASP LYS GLU LEU ARG VAL
SEQRES  36 A  728  LEU GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASP
SEQRES  37 A  728  VAL GLN MET PRO SER LYS LYS LEU ASP VAL ILE ASN LEU
SEQRES  38 A  728  HIS GLY THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO
SEQRES  39 A  728  HIS PHE ASP LYS SER LYS LYS TYR PRO LEU LEU ILE GLU
SEQRES  40 A  728  VAL TYR ALA GLY PRO CYS SER GLN LYS VAL ASP THR VAL
SEQRES  41 A  728  PHE ARG LEU SER TRP ALA THR TYR LEU ALA SER THR GLU
SEQRES  42 A  728  ASN ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY
SEQRES  43 A  728  TYR GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG
SEQRES  44 A  728  LEU GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA THR
SEQRES  45 A  728  ARG GLN PHE SER LYS MET GLY PHE VAL ASP ASP LYS ARG
SEQRES  46 A  728  ILE ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR
SEQRES  47 A  728  SER MET VAL LEU GLY ALA GLY SER GLY VAL PHE LYS CYS
SEQRES  48 A  728  GLY ILE ALA VAL ALA PRO VAL SER LYS TRP GLU TYR TYR
SEQRES  49 A  728  ASP SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR
SEQRES  50 A  728  PRO GLU ASP ASN LEU ASP TYR TYR ARG ASN SER THR VAL
SEQRES  51 A  728  MET SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU
SEQRES  52 A  728  LEU ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN
SEQRES  53 A  728  GLN SER ALA GLN LEU SER LYS ALA LEU VAL ASP ALA GLY
SEQRES  54 A  728  VAL ASP PHE GLN THR MET TRP TYR THR ASP GLU ASP HIS
SEQRES  55 A  728  GLY ILE ALA SER ASN MET ALA HIS GLN HIS ILE TYR THR
SEQRES  56 A  728  HIS MET SER HIS PHE LEU LYS GLN CYS PHE SER LEU PRO
SEQRES   1 B  728  SER ARG ARG THR TYR THR LEU THR ASP TYR LEU LYS SER
SEQRES   2 B  728  THR PHE ARG VAL LYS PHE TYR THR LEU GLN TRP ILE SER
SEQRES   3 B  728  ASP HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU
SEQRES   4 B  728  LEU PHE ASN ALA GLU TYR GLY ASN SER SER ILE PHE LEU
SEQRES   5 B  728  GLU ASN SER THR PHE ASP GLU LEU GLY TYR SER THR ASN
SEQRES   6 B  728  ASP TYR SER VAL SER PRO ASP ARG GLN PHE ILE LEU PHE
SEQRES   7 B  728  GLU TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR
SEQRES   8 B  728  ALA SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU
SEQRES   9 B  728  ILE THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP ILE
SEQRES  10 B  728  THR TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP
SEQRES  11 B  728  ASN ASN ASP ILE TYR VAL LYS ASN GLU PRO ASN LEU SER
SEQRES  12 B  728  SER GLN ARG ILE THR TRP THR GLY LYS GLU ASN VAL ILE
SEQRES  13 B  728  TYR ASN GLY VAL THR ASP TRP VAL TYR GLU GLU GLU VAL
SEQRES  14 B  728  PHE SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY
SEQRES  15 B  728  THR PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL
SEQRES  16 B  728  PRO LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU
SEQRES  17 B  728  GLN TYR PRO LYS THR VAL ARG ILE PRO TYR PRO LYS ALA
SEQRES  18 B  728  GLY ALA GLU ASN PRO THR VAL LYS PHE PHE VAL VAL ASP
SEQRES  19 B  728  THR ARG THR LEU SER PRO ASN ALA SER VAL THR SER TYR
SEQRES  20 B  728  GLN ILE VAL PRO PRO ALA SER VAL LEU ILE GLY ASP HIS
SEQRES  21 B  728  TYR LEU CYS GLY VAL THR TRP VAL THR GLU GLU ARG ILE
SEQRES  22 B  728  SER LEU GLN TRP ILE ARG ARG ALA GLN ASN TYR SER ILE
SEQRES  23 B  728  ILE ASP ILE CYS ASP TYR ASP GLU SER THR GLY ARG TRP
SEQRES  24 B  728  ILE SER SER VAL ALA ARG GLN HIS ILE GLU ILE SER THR
SEQRES  25 B  728  THR GLY TRP VAL GLY ARG PHE ARG PRO ALA GLU PRO HIS
SEQRES  26 B  728  PHE THR SER ASP GLY ASN SER PHE TYR LYS ILE ILE SER
SEQRES  27 B  728  ASN GLU GLU GLY TYR LYS HIS ILE CYS HIS PHE GLN THR
SEQRES  28 B  728  ASP LYS SER ASN CYS THR PHE ILE THR LYS GLY ALA TRP
SEQRES  29 B  728  GLU VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU
SEQRES  30 B  728  TYR TYR ILE SER ASN GLU HIS LYS GLY MET PRO GLY GLY
SEQRES  31 B  728  ARG ASN LEU TYR ARG ILE GLN LEU ASN ASP TYR THR LYS
SEQRES  32 B  728  VAL THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS
SEQRES  33 B  728  GLN TYR TYR SER ALA SER PHE SER ASN LYS ALA LYS TYR
SEQRES  34 B  728  TYR GLN LEU ARG CYS PHE GLY PRO GLY LEU PRO LEU TYR
SEQRES  35 B  728  THR LEU HIS SER SER SER SER ASP LYS GLU LEU ARG VAL
SEQRES  36 B  728  LEU GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASP
SEQRES  37 B  728  VAL GLN MET PRO SER LYS LYS LEU ASP VAL ILE ASN LEU
SEQRES  38 B  728  HIS GLY THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO
SEQRES  39 B  728  HIS PHE ASP LYS SER LYS LYS TYR PRO LEU LEU ILE GLU
SEQRES  40 B  728  VAL TYR ALA GLY PRO CYS SER GLN LYS VAL ASP THR VAL
SEQRES  41 B  728  PHE ARG LEU SER TRP ALA THR TYR LEU ALA SER THR GLU
SEQRES  42 B  728  ASN ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY
SEQRES  43 B  728  TYR GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG
SEQRES  44 B  728  LEU GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA THR
SEQRES  45 B  728  ARG GLN PHE SER LYS MET GLY PHE VAL ASP ASP LYS ARG
SEQRES  46 B  728  ILE ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR
SEQRES  47 B  728  SER MET VAL LEU GLY ALA GLY SER GLY VAL PHE LYS CYS
SEQRES  48 B  728  GLY ILE ALA VAL ALA PRO VAL SER LYS TRP GLU TYR TYR
SEQRES  49 B  728  ASP SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR
SEQRES  50 B  728  PRO GLU ASP ASN LEU ASP TYR TYR ARG ASN SER THR VAL
SEQRES  51 B  728  MET SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU
SEQRES  52 B  728  LEU ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN
SEQRES  53 B  728  GLN SER ALA GLN LEU SER LYS ALA LEU VAL ASP ALA GLY
SEQRES  54 B  728  VAL ASP PHE GLN THR MET TRP TYR THR ASP GLU ASP HIS
SEQRES  55 B  728  GLY ILE ALA SER ASN MET ALA HIS GLN HIS ILE TYR THR
SEQRES  56 B  728  HIS MET SER HIS PHE LEU LYS GLN CYS PHE SER LEU PRO
SEQRES   1 C  728  SER ARG ARG THR TYR THR LEU THR ASP TYR LEU LYS SER
SEQRES   2 C  728  THR PHE ARG VAL LYS PHE TYR THR LEU GLN TRP ILE SER
SEQRES   3 C  728  ASP HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU
SEQRES   4 C  728  LEU PHE ASN ALA GLU TYR GLY ASN SER SER ILE PHE LEU
SEQRES   5 C  728  GLU ASN SER THR PHE ASP GLU LEU GLY TYR SER THR ASN
SEQRES   6 C  728  ASP TYR SER VAL SER PRO ASP ARG GLN PHE ILE LEU PHE
SEQRES   7 C  728  GLU TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR
SEQRES   8 C  728  ALA SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU
SEQRES   9 C  728  ILE THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP ILE
SEQRES  10 C  728  THR TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP
SEQRES  11 C  728  ASN ASN ASP ILE TYR VAL LYS ASN GLU PRO ASN LEU SER
SEQRES  12 C  728  SER GLN ARG ILE THR TRP THR GLY LYS GLU ASN VAL ILE
SEQRES  13 C  728  TYR ASN GLY VAL THR ASP TRP VAL TYR GLU GLU GLU VAL
SEQRES  14 C  728  PHE SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY
SEQRES  15 C  728  THR PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL
SEQRES  16 C  728  PRO LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU
SEQRES  17 C  728  GLN TYR PRO LYS THR VAL ARG ILE PRO TYR PRO LYS ALA
SEQRES  18 C  728  GLY ALA GLU ASN PRO THR VAL LYS PHE PHE VAL VAL ASP
SEQRES  19 C  728  THR ARG THR LEU SER PRO ASN ALA SER VAL THR SER TYR
SEQRES  20 C  728  GLN ILE VAL PRO PRO ALA SER VAL LEU ILE GLY ASP HIS
SEQRES  21 C  728  TYR LEU CYS GLY VAL THR TRP VAL THR GLU GLU ARG ILE
SEQRES  22 C  728  SER LEU GLN TRP ILE ARG ARG ALA GLN ASN TYR SER ILE
SEQRES  23 C  728  ILE ASP ILE CYS ASP TYR ASP GLU SER THR GLY ARG TRP
SEQRES  24 C  728  ILE SER SER VAL ALA ARG GLN HIS ILE GLU ILE SER THR
SEQRES  25 C  728  THR GLY TRP VAL GLY ARG PHE ARG PRO ALA GLU PRO HIS
SEQRES  26 C  728  PHE THR SER ASP GLY ASN SER PHE TYR LYS ILE ILE SER
SEQRES  27 C  728  ASN GLU GLU GLY TYR LYS HIS ILE CYS HIS PHE GLN THR
SEQRES  28 C  728  ASP LYS SER ASN CYS THR PHE ILE THR LYS GLY ALA TRP
SEQRES  29 C  728  GLU VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU
SEQRES  30 C  728  TYR TYR ILE SER ASN GLU HIS LYS GLY MET PRO GLY GLY
SEQRES  31 C  728  ARG ASN LEU TYR ARG ILE GLN LEU ASN ASP TYR THR LYS
SEQRES  32 C  728  VAL THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS
SEQRES  33 C  728  GLN TYR TYR SER ALA SER PHE SER ASN LYS ALA LYS TYR
SEQRES  34 C  728  TYR GLN LEU ARG CYS PHE GLY PRO GLY LEU PRO LEU TYR
SEQRES  35 C  728  THR LEU HIS SER SER SER SER ASP LYS GLU LEU ARG VAL
SEQRES  36 C  728  LEU GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASP
SEQRES  37 C  728  VAL GLN MET PRO SER LYS LYS LEU ASP VAL ILE ASN LEU
SEQRES  38 C  728  HIS GLY THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO
SEQRES  39 C  728  HIS PHE ASP LYS SER LYS LYS TYR PRO LEU LEU ILE GLU
SEQRES  40 C  728  VAL TYR ALA GLY PRO CYS SER GLN LYS VAL ASP THR VAL
SEQRES  41 C  728  PHE ARG LEU SER TRP ALA THR TYR LEU ALA SER THR GLU
SEQRES  42 C  728  ASN ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY
SEQRES  43 C  728  TYR GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG
SEQRES  44 C  728  LEU GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA THR
SEQRES  45 C  728  ARG GLN PHE SER LYS MET GLY PHE VAL ASP ASP LYS ARG
SEQRES  46 C  728  ILE ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR
SEQRES  47 C  728  SER MET VAL LEU GLY ALA GLY SER GLY VAL PHE LYS CYS
SEQRES  48 C  728  GLY ILE ALA VAL ALA PRO VAL SER LYS TRP GLU TYR TYR
SEQRES  49 C  728  ASP SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR
SEQRES  50 C  728  PRO GLU ASP ASN LEU ASP TYR TYR ARG ASN SER THR VAL
SEQRES  51 C  728  MET SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU
SEQRES  52 C  728  LEU ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN
SEQRES  53 C  728  GLN SER ALA GLN LEU SER LYS ALA LEU VAL ASP ALA GLY
SEQRES  54 C  728  VAL ASP PHE GLN THR MET TRP TYR THR ASP GLU ASP HIS
SEQRES  55 C  728  GLY ILE ALA SER ASN MET ALA HIS GLN HIS ILE TYR THR
SEQRES  56 C  728  HIS MET SER HIS PHE LEU LYS GLN CYS PHE SER LEU PRO
SEQRES   1 D  728  SER ARG ARG THR TYR THR LEU THR ASP TYR LEU LYS SER
SEQRES   2 D  728  THR PHE ARG VAL LYS PHE TYR THR LEU GLN TRP ILE SER
SEQRES   3 D  728  ASP HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN ILE LEU
SEQRES   4 D  728  LEU PHE ASN ALA GLU TYR GLY ASN SER SER ILE PHE LEU
SEQRES   5 D  728  GLU ASN SER THR PHE ASP GLU LEU GLY TYR SER THR ASN
SEQRES   6 D  728  ASP TYR SER VAL SER PRO ASP ARG GLN PHE ILE LEU PHE
SEQRES   7 D  728  GLU TYR ASN TYR VAL LYS GLN TRP ARG HIS SER TYR THR
SEQRES   8 D  728  ALA SER TYR ASP ILE TYR ASP LEU ASN LYS ARG GLN LEU
SEQRES   9 D  728  ILE THR GLU GLU ARG ILE PRO ASN ASN THR GLN TRP ILE
SEQRES  10 D  728  THR TRP SER PRO VAL GLY HIS LYS LEU ALA TYR VAL TRP
SEQRES  11 D  728  ASN ASN ASP ILE TYR VAL LYS ASN GLU PRO ASN LEU SER
SEQRES  12 D  728  SER GLN ARG ILE THR TRP THR GLY LYS GLU ASN VAL ILE
SEQRES  13 D  728  TYR ASN GLY VAL THR ASP TRP VAL TYR GLU GLU GLU VAL
SEQRES  14 D  728  PHE SER ALA TYR SER ALA LEU TRP TRP SER PRO ASN GLY
SEQRES  15 D  728  THR PHE LEU ALA TYR ALA GLN PHE ASN ASP THR GLU VAL
SEQRES  16 D  728  PRO LEU ILE GLU TYR SER PHE TYR SER ASP GLU SER LEU
SEQRES  17 D  728  GLN TYR PRO LYS THR VAL ARG ILE PRO TYR PRO LYS ALA
SEQRES  18 D  728  GLY ALA GLU ASN PRO THR VAL LYS PHE PHE VAL VAL ASP
SEQRES  19 D  728  THR ARG THR LEU SER PRO ASN ALA SER VAL THR SER TYR
SEQRES  20 D  728  GLN ILE VAL PRO PRO ALA SER VAL LEU ILE GLY ASP HIS
SEQRES  21 D  728  TYR LEU CYS GLY VAL THR TRP VAL THR GLU GLU ARG ILE
SEQRES  22 D  728  SER LEU GLN TRP ILE ARG ARG ALA GLN ASN TYR SER ILE
SEQRES  23 D  728  ILE ASP ILE CYS ASP TYR ASP GLU SER THR GLY ARG TRP
SEQRES  24 D  728  ILE SER SER VAL ALA ARG GLN HIS ILE GLU ILE SER THR
SEQRES  25 D  728  THR GLY TRP VAL GLY ARG PHE ARG PRO ALA GLU PRO HIS
SEQRES  26 D  728  PHE THR SER ASP GLY ASN SER PHE TYR LYS ILE ILE SER
SEQRES  27 D  728  ASN GLU GLU GLY TYR LYS HIS ILE CYS HIS PHE GLN THR
SEQRES  28 D  728  ASP LYS SER ASN CYS THR PHE ILE THR LYS GLY ALA TRP
SEQRES  29 D  728  GLU VAL ILE GLY ILE GLU ALA LEU THR SER ASP TYR LEU
SEQRES  30 D  728  TYR TYR ILE SER ASN GLU HIS LYS GLY MET PRO GLY GLY
SEQRES  31 D  728  ARG ASN LEU TYR ARG ILE GLN LEU ASN ASP TYR THR LYS
SEQRES  32 D  728  VAL THR CYS LEU SER CYS GLU LEU ASN PRO GLU ARG CYS
SEQRES  33 D  728  GLN TYR TYR SER ALA SER PHE SER ASN LYS ALA LYS TYR
SEQRES  34 D  728  TYR GLN LEU ARG CYS PHE GLY PRO GLY LEU PRO LEU TYR
SEQRES  35 D  728  THR LEU HIS SER SER SER SER ASP LYS GLU LEU ARG VAL
SEQRES  36 D  728  LEU GLU ASP ASN SER ALA LEU ASP LYS MET LEU GLN ASP
SEQRES  37 D  728  VAL GLN MET PRO SER LYS LYS LEU ASP VAL ILE ASN LEU
SEQRES  38 D  728  HIS GLY THR LYS PHE TRP TYR GLN MET ILE LEU PRO PRO
SEQRES  39 D  728  HIS PHE ASP LYS SER LYS LYS TYR PRO LEU LEU ILE GLU
SEQRES  40 D  728  VAL TYR ALA GLY PRO CYS SER GLN LYS VAL ASP THR VAL
SEQRES  41 D  728  PHE ARG LEU SER TRP ALA THR TYR LEU ALA SER THR GLU
SEQRES  42 D  728  ASN ILE ILE VAL ALA SER PHE ASP GLY ARG GLY SER GLY
SEQRES  43 D  728  TYR GLN GLY ASP LYS ILE MET HIS ALA ILE ASN ARG ARG
SEQRES  44 D  728  LEU GLY THR PHE GLU VAL GLU ASP GLN ILE GLU ALA THR
SEQRES  45 D  728  ARG GLN PHE SER LYS MET GLY PHE VAL ASP ASP LYS ARG
SEQRES  46 D  728  ILE ALA ILE TRP GLY TRP SER TYR GLY GLY TYR VAL THR
SEQRES  47 D  728  SER MET VAL LEU GLY ALA GLY SER GLY VAL PHE LYS CYS
SEQRES  48 D  728  GLY ILE ALA VAL ALA PRO VAL SER LYS TRP GLU TYR TYR
SEQRES  49 D  728  ASP SER VAL TYR THR GLU ARG TYR MET GLY LEU PRO THR
SEQRES  50 D  728  PRO GLU ASP ASN LEU ASP TYR TYR ARG ASN SER THR VAL
SEQRES  51 D  728  MET SER ARG ALA GLU ASN PHE LYS GLN VAL GLU TYR LEU
SEQRES  52 D  728  LEU ILE HIS GLY THR ALA ASP ASP ASN VAL HIS PHE GLN
SEQRES  53 D  728  GLN SER ALA GLN LEU SER LYS ALA LEU VAL ASP ALA GLY
SEQRES  54 D  728  VAL ASP PHE GLN THR MET TRP TYR THR ASP GLU ASP HIS
SEQRES  55 D  728  GLY ILE ALA SER ASN MET ALA HIS GLN HIS ILE TYR THR
SEQRES  56 D  728  HIS MET SER HIS PHE LEU LYS GLN CYS PHE SER LEU PRO
HET    NAG  A1085      14
HET    NAG  A1092      14
HET    NAG  A1229      14
HET    NAG  A1230      14
HET    NAG  A1279      14
HET    NAG  A1321      14
HET    NDG  A1685      14
HET    NDG  A1686      14
HET    NAG  B1085      14
HET    NAG  B1092      14
HET    NAG  B1093      14
HET    NAG  B1229      14
HET    NAG  B1279      14
HET    NAG  B1321      14
HET    NAG  B1685      14
HET    NAG  B1686      14
HET    NAG  C1085      14
HET    NDG  C1092      14
HET    NAG  C1229      14
HET    NAG  C1279      14
HET    NAG  C1321      14
HET    NAG  C1685      14
HET    NDG  C1686      14
HET    NDG  D1085      14
HET    NAG  D1092      14
HET    NAG  D1093      14
HET    NDG  D1229      14
HET    NDG  D1230      14
HET    NDG  D1279      14
HET    NAG  D1321      14
HET    NDG  D1322      14
HET    NAG  D1685      14
HET    NAG  D1686      14
HET    008  A1767      26
HET    008  B1767      26
HET    008  C1767      26
HET    008  D1767      26
HET    SO4  A1768       5
HET    SO4  B1768       5
HET    SO4  C1768       5
HET    SO4  D1768       5
HET    SO4  A1769       5
HET    SO4  B1769       5
HET    SO4  C1769       5
HET    SO4  D1769       5
HETNAM     SO4 SULFATE ION
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETSYN     NAG NAG
HETNAM     NDG 2-(ACETYLAMINO)-2-DEOXY-A-D-GLUCOPYRANOSE
HETNAM     008 (S)-2-[(R)-3-AMINO-4-(2-FLUORO-PHENYL)-
HETNAM   2 008  BUTYRYL]-1,2,3,4-TETRAHYDRO-ISOQUINOLINE-3-
HETNAM   3 008  CARBOXYLIC ACID AMIDE
FORMUL   5  008    4(C20 H22 N3 O2 F1)
FORMUL   6  SO4    8(O4 S1 2-)
FORMUL   7  NAG    24(C8 H15 N1 O6)
FORMUL   8  NDG    9(C8 H15 N1 O6)
FORMUL  10  HOH   *951(H2 O1)
HELIX    1   1 THR A   44  LYS A   50  1                                   7
HELIX    2   2 ASN A   92  GLU A   97  1                                   6
HELIX    3   3 ASP A  200  GLU A  206  1                                   7
HELIX    4   4 ARG A  274  LEU A  276  5                                   3
HELIX    5   5 PRO A  290  ILE A  295  1                                   6
HELIX    6   6 SER A  340  ALA A  342  5                                   3
HELIX    7   7 GLU A  421  MET A  425  5                                   5
HELIX    8   8 ASN A  497  ASP A  506  1                                  10
HELIX    9   9 SER A  562  THR A  570  1                                   9
HELIX   10  10 GLY A  587  HIS A  592  1                                   6
HELIX   11  11 ALA A  593  ASN A  595  5                                   3
HELIX   12  12 THR A  600  MET A  616  1                                  17
HELIX   13  13 SER A  630  GLY A  641  1                                  12
HELIX   14  14 LYS A  658  TYR A  662  5                                   5
HELIX   15  15 ASP A  663  GLY A  672  1                                  10
HELIX   16  16 ASN A  679  ASN A  685  1                                   7
HELIX   17  17 THR A  687  VAL A  698  5                                  12
HELIX   18  18 HIS A  712  GLY A  727  1                                  16
HELIX   19  19 SER A  744  PHE A  763  1                                  20
HELIX   20  20 THR B   44  LYS B   50  1                                   7
HELIX   21  21 ASN B   92  GLU B   97  1                                   6
HELIX   22  22 ASP B  200  VAL B  207  1                                   8
HELIX   23  23 PRO B  290  ILE B  295  1                                   6
HELIX   24  24 SER B  340  ALA B  342  5                                   3
HELIX   25  25 GLU B  421  MET B  425  5                                   5
HELIX   26  26 ASN B  497  GLN B  505  1                                   9
HELIX   27  27 SER B  562  THR B  570  1                                   9
HELIX   28  28 GLY B  587  HIS B  592  1                                   6
HELIX   29  29 ALA B  593  ASN B  595  5                                   3
HELIX   30  30 THR B  600  MET B  616  1                                  17
HELIX   31  31 SER B  630  GLY B  641  1                                  12
HELIX   32  32 LYS B  658  TYR B  662  5                                   5
HELIX   33  33 ASP B  663  GLY B  672  1                                  10
HELIX   34  34 ASN B  679  ASN B  685  1                                   7
HELIX   35  35 VAL B  688  GLN B  697  5                                  10
HELIX   36  36 HIS B  712  GLY B  727  1                                  16
HELIX   37  37 SER B  744  PHE B  763  1                                  20
HELIX   38  38 THR C   44  LYS C   50  1                                   7
HELIX   39  39 ASN C   92  GLU C   97  1                                   6
HELIX   40  40 ASP C  200  VAL C  207  1                                   8
HELIX   41  41 ARG C  274  LEU C  276  5                                   3
HELIX   42  42 PRO C  290  ILE C  295  1                                   6
HELIX   43  43 SER C  340  ALA C  342  5                                   3
HELIX   44  44 GLU C  421  MET C  425  5                                   5
HELIX   45  45 ASN C  497  GLN C  505  1                                   9
HELIX   46  46 SER C  562  THR C  570  1                                   9
HELIX   47  47 GLY C  587  HIS C  592  1                                   6
HELIX   48  48 ALA C  593  ASN C  595  5                                   3
HELIX   49  49 THR C  600  MET C  616  1                                  17
HELIX   50  50 SER C  630  GLY C  641  1                                  12
HELIX   51  51 ASP C  663  GLY C  672  1                                  10
HELIX   52  52 ASN C  679  ASN C  685  1                                   7
HELIX   53  53 VAL C  688  VAL C  698  5                                  11
HELIX   54  54 HIS C  712  GLY C  727  1                                  16
HELIX   55  55 SER C  744  PHE C  763  1                                  20
HELIX   56  56 THR D   44  LYS D   50  1                                   7
HELIX   57  57 ASN D   92  GLU D   97  1                                   6
HELIX   58  58 ASP D  200  VAL D  207  1                                   8
HELIX   59  59 ARG D  274  LEU D  276  5                                   3
HELIX   60  60 PRO D  290  ILE D  295  1                                   6
HELIX   61  61 SER D  340  ALA D  342  5                                   3
HELIX   62  62 GLU D  421  MET D  425  5                                   5
HELIX   63  63 ASN D  497  GLN D  505  1                                   9
HELIX   64  64 SER D  562  THR D  570  1                                   9
HELIX   65  65 GLY D  587  HIS D  592  1                                   6
HELIX   66  66 ALA D  593  ASN D  595  5                                   3
HELIX   67  67 THR D  600  LYS D  615  1                                  16
HELIX   68  68 SER D  630  GLY D  641  1                                  12
HELIX   69  69 ASP D  663  GLY D  672  1                                  10
HELIX   70  70 ASN D  679  ASN D  685  1                                   7
HELIX   71  71 VAL D  688  GLN D  697  5                                  10
HELIX   72  72 PHE D  713  ALA D  726  1                                  14
HELIX   73  73 SER D  744  PHE D  763  1                                  20
SHEET    1  AA 2 ARG A  41  THR A  42  0
SHEET    2  AA 2 VAL A 507  GLN A 508  1  N  GLN A 508   O  ARG A  41
SHEET    1  AB 4 LEU A  60  TRP A  62  0
SHEET    2  AB 4 GLU A  67  GLN A  72 -1  O  LEU A  69   N  GLN A  61
SHEET    3  AB 4 ASN A  75  ASN A  80 -1  O  ASN A  75   N  GLN A  72
SHEET    4  AB 4 SER A  86  LEU A  90 -1  O  SER A  87   N  LEU A  78
SHEET    1  AC 4 ASP A 104  VAL A 107  0
SHEET    2  AC 4 PHE A 113  LYS A 122 -1  O  LEU A 115   N  SER A 106
SHEET    3  AC 4 TYR A 128  ASP A 136 -1  O  THR A 129   N  VAL A 121
SHEET    4  AC 4 ILE A 148  THR A 152 -1  O  ILE A 148   N  TYR A 132
SHEET    1  AD 4 TRP A 154  TRP A 157  0
SHEET    2  AD 4 LEU A 164  TRP A 168 -1  O  ALA A 165   N  THR A 156
SHEET    3  AD 4 ASP A 171  LYS A 175 -1  O  ASP A 171   N  TRP A 168
SHEET    4  AD 4 GLN A 183  ARG A 184 -1  O  GLN A 183   N  VAL A 174
SHEET    1  AE 3 ILE A 194  ASN A 196  0
SHEET    2  AE 3 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3  AE 3 LEU A 214  TRP A 216 -1  O  TRP A 215   N  ALA A 224
SHEET    1  AF 4 ILE A 194  ASN A 196  0
SHEET    2  AF 4 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3  AF 4 THR A 265  ASP A 272 -1  O  THR A 265   N  ASN A 229
SHEET    4  AF 4 SER A 284  ILE A 287 -1  O  TYR A 285   N  VAL A 270
SHEET    1  AG 2 LEU A 235  PHE A 240  0
SHEET    2  AG 2 LYS A 250  PRO A 255 -1  O  LYS A 250   N  PHE A 240
SHEET    1  AH 4 HIS A 298  THR A 307  0
SHEET    2  AH 4 ARG A 310  ARG A 317 -1  O  ARG A 310   N  VAL A 306
SHEET    3  AH 4 TYR A 322  TYR A 330 -1  O  TYR A 322   N  ARG A 317
SHEET    4  AH 4 GLN A 344  ILE A 348 -1  O  HIS A 345   N  ILE A 325
SHEET    1  AI 4 HIS A 298  THR A 307  0
SHEET    2  AI 4 ARG A 310  ARG A 317 -1  O  ARG A 310   N  VAL A 306
SHEET    3  AI 4 TYR A 322  TYR A 330 -1  O  TYR A 322   N  ARG A 317
SHEET    4  AI 4 TRP A 337  ILE A 338 -1  O  ILE A 338   N  ASP A 329
SHEET    1  AJ 4 HIS A 363  PHE A 364  0
SHEET    2  AJ 4 SER A 370  SER A 376 -1  O  TYR A 372   N  HIS A 363
SHEET    3  AJ 4 LYS A 382  GLN A 388 -1  O  HIS A 383   N  ILE A 375
SHEET    4  AJ 4 THR A 395  PHE A 396 -1  O  THR A 395   N  HIS A 386
SHEET    1  AK 4 VAL A 404  LEU A 410  0
SHEET    2  AK 4 TYR A 414  SER A 419 -1  O  TYR A 416   N  GLU A 408
SHEET    3  AK 4 ASN A 430  GLN A 435 -1  O  ASN A 430   N  SER A 419
SHEET    4  AK 4 VAL A 442  CYS A 444 -1  O  THR A 443   N  ARG A 433
SHEET    1  AL 4 CYS A 454  PHE A 461  0
SHEET    2  AL 4 TYR A 467  PRO A 475 -1  O  GLN A 469   N  SER A 460
SHEET    3  AL 4 LEU A 479  SER A 484 -1  O  LEU A 479   N  CYS A 472
SHEET    4  AL 4 GLU A 490  GLU A 495 -1  N  LEU A 491   O  LEU A 482
SHEET    1  AM 8 SER A 511  LEU A 519  0
SHEET    2  AM 8 THR A 522  LEU A 530 -1  O  THR A 522   N  LEU A 519
SHEET    3  AM 8 ILE A 574  PHE A 578 -1  O  VAL A 575   N  ILE A 529
SHEET    4  AM 8 TYR A 540  VAL A 546  1  O  PRO A 541   N  ILE A 574
SHEET    5  AM 8 VAL A 619  TRP A 629  1  N  ASP A 620   O  TYR A 540
SHEET    6  AM 8 CYS A 649  VAL A 653  1  O  CYS A 649   N  ILE A 626
SHEET    7  AM 8 GLU A 699  GLY A 705  1  O  GLU A 699   N  GLY A 650
SHEET    8  AM 8 GLN A 731  TYR A 735  1  O  GLN A 731   N  LEU A 702
SHEET    1  BA 2 ARG B  41  THR B  42  0
SHEET    2  BA 2 VAL B 507  GLN B 508  1  N  GLN B 508   O  ARG B  41
SHEET    1  BB 4 LEU B  60  TRP B  62  0
SHEET    2  BB 4 GLU B  67  GLN B  72 -1  O  LEU B  69   N  GLN B  61
SHEET    3  BB 4 ASN B  75  ASN B  80 -1  O  ASN B  75   N  GLN B  72
SHEET    4  BB 4 SER B  86  LEU B  90 -1  O  SER B  87   N  LEU B  78
SHEET    1  BC 4 ASP B 104  VAL B 107  0
SHEET    2  BC 4 PHE B 113  LYS B 122 -1  O  LEU B 115   N  SER B 106
SHEET    3  BC 4 TYR B 128  ASP B 136 -1  O  THR B 129   N  VAL B 121
SHEET    4  BC 4 ILE B 148  THR B 152 -1  O  ILE B 148   N  TYR B 132
SHEET    1  BD 4 ASP B 104  VAL B 107  0
SHEET    2  BD 4 PHE B 113  LYS B 122 -1  O  LEU B 115   N  SER B 106
SHEET    3  BD 4 TYR B 128  ASP B 136 -1  O  THR B 129   N  VAL B 121
SHEET    4  BD 4 GLN B 141  LEU B 142 -1  O  GLN B 141   N  ASP B 136
SHEET    1  BE 4 TRP B 154  TRP B 157  0
SHEET    2  BE 4 LEU B 164  TRP B 168 -1  O  ALA B 165   N  THR B 156
SHEET    3  BE 4 ASP B 171  LYS B 175 -1  O  ASP B 171   N  TRP B 168
SHEET    4  BE 4 GLN B 183  ARG B 184 -1  O  GLN B 183   N  VAL B 174
SHEET    1  BF 7 ILE B 194  ASN B 196  0
SHEET    2  BF 7 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3  BF 7 LEU B 214  TRP B 216 -1  O  TRP B 215   N  ALA B 224
SHEET    4  BF 7 PHE B 222  ASN B 229 -1  O  ALA B 224   N  TRP B 215
SHEET    5  BF 7 SER B 284  ILE B 287
SHEET    6  BF 7 THR B 265  ASP B 272 -1  O  PHE B 268   N  ILE B 287
SHEET    7  BF 7 PHE B 222  ASN B 229 -1  O  LEU B 223   N  VAL B 271
SHEET    1  BG 2 LEU B 235  PHE B 240  0
SHEET    2  BG 2 LYS B 250  PRO B 255 -1  O  LYS B 250   N  PHE B 240
SHEET    1  BH 7 HIS B 298  THR B 307  0
SHEET    2  BH 7 ARG B 310  ARG B 317 -1  O  ARG B 310   N  VAL B 306
SHEET    3  BH 7 TYR B 322  TYR B 330 -1  O  TYR B 322   N  ARG B 317
SHEET    4  BH 7 TRP B 337  ILE B 338 -1  O  ILE B 338   N  ASP B 329
SHEET    5  BH 7 TYR B 322  TYR B 330 -1  O  ASP B 329   N  ILE B 338
SHEET    6  BH 7 GLN B 344  ILE B 348 -1  O  HIS B 345   N  ILE B 325
SHEET    7  BH 7 TYR B 322  TYR B 330 -1  O  SER B 323   N  GLU B 347
SHEET    1  BI 4 HIS B 363  PHE B 364  0
SHEET    2  BI 4 SER B 370  SER B 376 -1  O  TYR B 372   N  HIS B 363
SHEET    3  BI 4 LYS B 382  GLN B 388 -1  O  HIS B 383   N  ILE B 375
SHEET    4  BI 4 CYS B 394  PHE B 396 -1  O  THR B 395   N  HIS B 386
SHEET    1  BJ 4 VAL B 404  LEU B 410  0
SHEET    2  BJ 4 TYR B 414  SER B 419 -1  O  TYR B 416   N  GLU B 408
SHEET    3  BJ 4 ASN B 430  GLN B 435 -1  O  ASN B 430   N  SER B 419
SHEET    4  BJ 4 VAL B 442  CYS B 444 -1  O  THR B 443   N  ARG B 433
SHEET    1  BK 4 TYR B 457  PHE B 461  0
SHEET    2  BK 4 TYR B 467  CYS B 472 -1  O  GLN B 469   N  SER B 460
SHEET    3  BK 4 LEU B 479  SER B 484 -1  O  LEU B 479   N  CYS B 472
SHEET    4  BK 4 GLU B 490  GLU B 495 -1  N  LEU B 491   O  LEU B 482
SHEET    1  BL 8 SER B 511  LEU B 519  0
SHEET    2  BL 8 THR B 522  LEU B 530 -1  O  THR B 522   N  LEU B 519
SHEET    3  BL 8 ILE B 574  PHE B 578 -1  O  VAL B 575   N  ILE B 529
SHEET    4  BL 8 TYR B 540  VAL B 546  1  O  PRO B 541   N  ILE B 574
SHEET    5  BL 8 VAL B 619  TRP B 629  1  N  ASP B 620   O  TYR B 540
SHEET    6  BL 8 CYS B 649  VAL B 653  1  O  CYS B 649   N  ILE B 626
SHEET    7  BL 8 GLU B 699  GLY B 705  1  O  GLU B 699   N  GLY B 650
SHEET    8  BL 8 GLN B 731  TYR B 735  1  O  GLN B 731   N  LEU B 702
SHEET    1  CA 2 ARG C  41  THR C  42  0
SHEET    2  CA 2 VAL C 507  GLN C 508  1  N  GLN C 508   O  ARG C  41
SHEET    1  CB 4 LEU C  60  TRP C  62  0
SHEET    2  CB 4 GLU C  67  GLN C  72 -1  O  LEU C  69   N  GLN C  61
SHEET    3  CB 4 ASN C  75  ASN C  80 -1  O  ASN C  75   N  GLN C  72
SHEET    4  CB 4 SER C  86  LEU C  90 -1  O  SER C  87   N  LEU C  78
SHEET    1  CC 4 ASP C 104  VAL C 107  0
SHEET    2  CC 4 PHE C 113  LYS C 122 -1  O  LEU C 115   N  SER C 106
SHEET    3  CC 4 TYR C 128  ASP C 136 -1  O  THR C 129   N  VAL C 121
SHEET    4  CC 4 GLN C 141  LEU C 142 -1  O  GLN C 141   N  ASP C 136
SHEET    1  CD 4 TRP C 154  TRP C 157  0
SHEET    2  CD 4 LEU C 164  TRP C 168 -1  O  ALA C 165   N  THR C 156
SHEET    3  CD 4 ASP C 171  LYS C 175 -1  O  ASP C 171   N  TRP C 168
SHEET    4  CD 4 GLN C 183  ARG C 184 -1  O  GLN C 183   N  VAL C 174
SHEET    1  CE 7 ILE C 194  ASN C 196  0
SHEET    2  CE 7 PHE C 222  ASN C 229 -1  O  PHE C 228   N  TYR C 195
SHEET    3  CE 7 LEU C 214  TRP C 216 -1  O  TRP C 215   N  ALA C 224
SHEET    4  CE 7 PHE C 222  ASN C 229 -1  O  ALA C 224   N  TRP C 215
SHEET    5  CE 7 THR C 283  ILE C 287
SHEET    6  CE 7 THR C 265  ASP C 272 -1  O  PHE C 268   N  ILE C 287
SHEET    7  CE 7 PHE C 222  ASN C 229 -1  O  LEU C 223   N  VAL C 271
SHEET    1  CF 2 LEU C 235  PHE C 240  0
SHEET    2  CF 2 LYS C 250  PRO C 255 -1  O  LYS C 250   N  PHE C 240
SHEET    1  CG 7 HIS C 298  THR C 307  0
SHEET    2  CG 7 ARG C 310  ARG C 317 -1  O  ARG C 310   N  VAL C 306
SHEET    3  CG 7 TYR C 322  TYR C 330 -1  O  TYR C 322   N  ARG C 317
SHEET    4  CG 7 TRP C 337  ILE C 338 -1  O  ILE C 338   N  ASP C 329
SHEET    5  CG 7 TYR C 322  TYR C 330 -1  O  ASP C 329   N  ILE C 338
SHEET    6  CG 7 GLN C 344  ILE C 348 -1  O  HIS C 345   N  ILE C 325
SHEET    7  CG 7 TYR C 322  TYR C 330 -1  O  SER C 323   N  GLU C 347
SHEET    1  CH 4 HIS C 363  PHE C 364  0
SHEET    2  CH 4 SER C 370  SER C 376 -1  O  TYR C 372   N  HIS C 363
SHEET    3  CH 4 LYS C 382  GLN C 388 -1  O  HIS C 383   N  ILE C 375
SHEET    4  CH 4 CYS C 394  PHE C 396 -1  O  THR C 395   N  HIS C 386
SHEET    1  CI 4 VAL C 404  LEU C 410  0
SHEET    2  CI 4 TYR C 414  SER C 419 -1  O  TYR C 416   N  GLU C 408
SHEET    3  CI 4 ASN C 430  GLN C 435 -1  O  ASN C 430   N  SER C 419
SHEET    4  CI 4 VAL C 442  CYS C 444 -1  O  THR C 443   N  ARG C 433
SHEET    1  CJ 4 TYR C 457  PHE C 461  0
SHEET    2  CJ 4 TYR C 467  CYS C 472 -1  O  GLN C 469   N  SER C 460
SHEET    3  CJ 4 LEU C 479  SER C 484 -1  O  LEU C 479   N  CYS C 472
SHEET    4  CJ 4 GLU C 490  GLU C 495 -1  N  LEU C 491   O  LEU C 482
SHEET    1  CK 8 SER C 511  LEU C 519  0
SHEET    2  CK 8 THR C 522  LEU C 530 -1  O  THR C 522   N  LEU C 519
SHEET    3  CK 8 ILE C 574  PHE C 578 -1  O  VAL C 575   N  ILE C 529
SHEET    4  CK 8 TYR C 540  VAL C 546  1  O  PRO C 541   N  ILE C 574
SHEET    5  CK 8 VAL C 619  TRP C 629  1  N  ASP C 620   O  TYR C 540
SHEET    6  CK 8 CYS C 649  VAL C 653  1  O  CYS C 649   N  ILE C 626
SHEET    7  CK 8 GLU C 699  GLY C 705  1  O  GLU C 699   N  GLY C 650
SHEET    8  CK 8 GLN C 731  TYR C 735  1  O  GLN C 731   N  LEU C 702
SHEET    1  DA 2 ARG D  41  THR D  42  0
SHEET    2  DA 2 VAL D 507  GLN D 508  1  N  GLN D 508   O  ARG D  41
SHEET    1  DB 4 LEU D  60  TRP D  62  0
SHEET    2  DB 4 GLU D  67  GLN D  72 -1  O  LEU D  69   N  GLN D  61
SHEET    3  DB 4 ASN D  75  ASN D  80 -1  O  ASN D  75   N  GLN D  72
SHEET    4  DB 4 SER D  86  LEU D  90 -1  O  SER D  87   N  LEU D  78
SHEET    1  DC 4 ASP D 104  VAL D 107  0
SHEET    2  DC 4 PHE D 113  LYS D 122 -1  O  LEU D 115   N  SER D 106
SHEET    3  DC 4 TYR D 128  ASP D 136 -1  O  THR D 129   N  VAL D 121
SHEET    4  DC 4 GLN D 141  LEU D 142 -1  O  GLN D 141   N  ASP D 136
SHEET    1  DD 4 THR D 152  TRP D 157  0
SHEET    2  DD 4 LEU D 164  TRP D 168 -1  O  ALA D 165   N  THR D 156
SHEET    3  DD 4 ASP D 171  LYS D 175 -1  O  ASP D 171   N  TRP D 168
SHEET    4  DD 4 GLN D 183  ARG D 184 -1  O  GLN D 183   N  VAL D 174
SHEET    1  DE 7 ILE D 194  ASN D 196  0
SHEET    2  DE 7 PHE D 222  ASN D 229 -1  O  PHE D 228   N  TYR D 195
SHEET    3  DE 7 LEU D 214  TRP D 216 -1  O  TRP D 215   N  ALA D 224
SHEET    4  DE 7 PHE D 222  ASN D 229 -1  O  ALA D 224   N  TRP D 215
SHEET    5  DE 7 THR D 283  ILE D 287
SHEET    6  DE 7 THR D 265  ASP D 272 -1  O  PHE D 268   N  ILE D 287
SHEET    7  DE 7 PHE D 222  ASN D 229 -1  O  LEU D 223   N  VAL D 271
SHEET    1  DF 2 LEU D 235  PHE D 240  0
SHEET    2  DF 2 LYS D 250  PRO D 255 -1  O  LYS D 250   N  PHE D 240
SHEET    1  DG 7 HIS D 298  THR D 307  0
SHEET    2  DG 7 ARG D 310  ARG D 317 -1  O  ARG D 310   N  VAL D 306
SHEET    3  DG 7 TYR D 322  TYR D 330 -1  O  TYR D 322   N  ARG D 317
SHEET    4  DG 7 TRP D 337  ILE D 338 -1  O  ILE D 338   N  ASP D 329
SHEET    5  DG 7 TYR D 322  TYR D 330 -1  O  ASP D 329   N  ILE D 338
SHEET    6  DG 7 GLN D 344  ILE D 348 -1  O  HIS D 345   N  ILE D 325
SHEET    7  DG 7 TYR D 322  TYR D 330 -1  O  SER D 323   N  GLU D 347
SHEET    1  DH 4 HIS D 363  PHE D 364  0
SHEET    2  DH 4 SER D 370  SER D 376 -1  O  TYR D 372   N  HIS D 363
SHEET    3  DH 4 LYS D 382  GLN D 388 -1  O  HIS D 383   N  ILE D 375
SHEET    4  DH 4 CYS D 394  PHE D 396 -1  O  THR D 395   N  HIS D 386
SHEET    1  DI 4 VAL D 404  LEU D 410  0
SHEET    2  DI 4 TYR D 414  SER D 419 -1  O  TYR D 416   N  GLU D 408
SHEET    3  DI 4 ASN D 430  GLN D 435 -1  O  ASN D 430   N  SER D 419
SHEET    4  DI 4 VAL D 442  CYS D 444 -1  O  THR D 443   N  ARG D 433
SHEET    1  DJ 4 TYR D 457  PHE D 461  0
SHEET    2  DJ 4 TYR D 467  CYS D 472 -1  O  GLN D 469   N  SER D 460
SHEET    3  DJ 4 LEU D 479  SER D 484 -1  O  LEU D 479   N  CYS D 472
SHEET    4  DJ 4 GLU D 490  GLU D 495 -1  N  LEU D 491   O  LEU D 482
SHEET    1  DK 8 SER D 511  LEU D 519  0
SHEET    2  DK 8 THR D 522  LEU D 530 -1  O  THR D 522   N  LEU D 519
SHEET    3  DK 8 ILE D 574  PHE D 578 -1  O  VAL D 575   N  ILE D 529
SHEET    4  DK 8 TYR D 540  VAL D 546  1  O  PRO D 541   N  ILE D 574
SHEET    5  DK 8 VAL D 619  TRP D 629  1  N  ASP D 620   O  TYR D 540
SHEET    6  DK 8 CYS D 649  VAL D 653  1  O  CYS D 649   N  ILE D 626
SHEET    7  DK 8 GLU D 699  GLY D 705  1  O  GLU D 699   N  GLY D 650
SHEET    8  DK 8 GLN D 731  TYR D 735  1  O  GLN D 731   N  LEU D 702
SSBOND   1 CYS A  385    CYS A  394                          1555   1555
SSBOND   2 CYS A  444    CYS A  447                          1555   1555
SSBOND   3 CYS A  454    CYS A  472                          1555   1555
SSBOND   4 CYS A  649    CYS A  762                          1555   1555
SSBOND   5 CYS B  385    CYS B  394                          1555   1555
SSBOND   6 CYS B  444    CYS B  447                          1555   1555
SSBOND   7 CYS B  454    CYS B  472                          1555   1555
SSBOND   8 CYS B  649    CYS B  762                          1555   1555
SSBOND   9 CYS C  385    CYS C  394                          1555   1555
SSBOND  10 CYS C  444    CYS C  447                          1555   1555
SSBOND  11 CYS C  454    CYS C  472                          1555   1555
SSBOND  12 CYS C  649    CYS C  762                          1555   1555
SSBOND  13 CYS D  385    CYS D  394                          1555   1555
SSBOND  14 CYS D  444    CYS D  447                          1555   1555
SSBOND  15 CYS D  454    CYS D  472                          1555   1555
SSBOND  16 CYS D  649    CYS D  762                          1555   1555
LINK         ND2 ASN A  85                 C1  NAG A1085     1555   1555
LINK         ND2 ASN A  92                 C1  NAG A1092     1555   1555
LINK         ND2 ASN A 229                 C1  NAG A1229     1555   1555
LINK         ND2 ASN A 279                 C1  NAG A1279     1555   1555
LINK         ND2 ASN A 321                 C1  NAG A1321     1555   1555
LINK         ND2 ASN A 685                 C1  NDG A1685     1555   1555
LINK         O4  NAG A1229                 C1  NAG A1230     1555   1555
LINK         O4  NDG A1685                 C1  NDG A1686     1555   1555
LINK         ND2 ASN B  85                 C1  NAG B1085     1555   1555
LINK         ND2 ASN B  92                 C1  NAG B1092     1555   1555
LINK         ND2 ASN B 229                 C1  NAG B1229     1555   1555
LINK         ND2 ASN B 279                 C1  NAG B1279     1555   1555
LINK         ND2 ASN B 321                 C1  NAG B1321     1555   1555
LINK         ND2 ASN B 685                 C1  NAG B1685     1555   1555
LINK         O4  NAG B1092                 C1  NAG B1093     1555   1555
LINK         O4  NAG B1685                 C1  NAG B1686     1555   1555
LINK         ND2 ASN C  85                 C1  NAG C1085     1555   1555
LINK         ND2 ASN C  92                 C1  NDG C1092     1555   1555
LINK         ND2 ASN C 229                 C1  NAG C1229     1555   1555
LINK         ND2 ASN C 279                 C1  NAG C1279     1555   1555
LINK         ND2 ASN C 321                 C1  NAG C1321     1555   1555
LINK         ND2 ASN C 685                 C1  NAG C1685     1555   1555
LINK         O4  NAG C1685                 C1  NDG C1686     1555   1555
LINK         ND2 ASN D  85                 C1  NDG D1085     1555   1555
LINK         ND2 ASN D  92                 C1  NAG D1092     1555   1555
LINK         ND2 ASN D 229                 C1  NDG D1229     1555   1555
LINK         ND2 ASN D 279                 C1  NDG D1279     1555   1555
LINK         ND2 ASN D 321                 C1  NAG D1321     1555   1555
LINK         ND2 ASN D 685                 C1  NAG D1685     1555   1555
LINK         O4  NAG D1092                 C1  NAG D1093     1555   1555
LINK         O4  NDG D1229                 C1  NDG D1230     1555   1555
LINK         O4  NAG D1321                 C1  NDG D1322     1555   1555
LINK         O4  NAG D1685                 C1  NAG D1686     1555   1555
CISPEP   1 GLY A  474    PRO A  475          0         4.63
CISPEP   2 GLY B  474    PRO B  475          0         4.90
CISPEP   3 GLY C  474    PRO C  475          0         2.10
CISPEP   4 GLY D  474    PRO D  475          0         4.39
SITE     1 AC1 13 ARG A 125  GLU A 205  GLU A 206  SER A 209
SITE     2 AC1 13 PHE A 357  TYR A 547  SER A 630  TYR A 662
SITE     3 AC1 13 TYR A 666  ASN A 710  HIS A 740  HOH Z 163
SITE     4 AC1 13 HOH Z 243
SITE     1 AC2 12 ARG B 125  GLU B 205  GLU B 206  SER B 209
SITE     2 AC2 12 PHE B 357  ARG B 358  TYR B 547  TYR B 662
SITE     3 AC2 12 TYR B 666  ASN B 710  HIS B 740  HOH Y 187
SITE     1 AC3 11 ARG C 125  GLU C 205  GLU C 206  SER C 209
SITE     2 AC3 11 PHE C 357  TYR C 662  TYR C 666  ASN C 710
SITE     3 AC3 11 HIS C 740  HOH X 160  HOH X 227
SITE     1 AC4 10 ARG D 125  GLU D 205  GLU D 206  SER D 209
SITE     2 AC4 10 PHE D 357  TYR D 547  TYR D 662  TYR D 666
SITE     3 AC4 10 ASN D 710  HIS D 740
SITE     1 AC5  7 ALA A 360  GLU A 361  HIS A 363  ILE A 374
SITE     2 AC5  7 GLY A 406  ILE A 407  HOH Z 123
SITE     1 AC6  8 ALA B 360  GLU B 361  HIS B 363  ILE B 374
SITE     2 AC6  8 GLY B 406  ILE B 407  HOH Y 138  HOH Y 246
SITE     1 AC7  7 ALA C 360  GLU C 361  HIS C 363  ILE C 374
SITE     2 AC7  7 GLY C 406  ILE C 407  HOH X 116
SITE     1 AC8  8 ALA D 360  GLU D 361  HIS D 363  ILE D 374
SITE     2 AC8  8 GLY D 406  ILE D 407  HOH W 132  HOH W 233
SITE     1 AC9  5 ARG A 125  TYR A 547  SER A 630  HIS A 740
SITE     2 AC9  5 HOH Z 244
SITE     1 BC1  4 ARG B 125  TYR B 547  SER B 630  HIS B 740
SITE     1 BC2  5 ARG C 125  TYR C 547  SER C 630  HIS C 740
SITE     2 BC2  5 HOH X 228
SITE     1 BC3  4 ARG D 125  TYR D 547  SER D 630  HIS D 740
CRYST1   61.644  117.491  133.057 112.44  94.72  91.30 P 1           4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016222  0.000368  0.001605        0.00000
SCALE2      0.000000  0.008513  0.003551        0.00000
SCALE3      0.000000  0.000000  0.008171        0.00000
TER    5967      PRO A 766
TER   11934      PRO B 766
TER   17901      PRO C 766
TER   23868      PRO D 766
MASTER      353    0   45   73  205    0   27    625421    4  662  224
END