longtext: 2PUJ-pdb

content
HEADER    HYDROLASE                               09-MAY-07   2PUJ
TITLE     CRYSTAL STRUCTURE OF THE S112A/H265A DOUBLE MUTANT OF A C-C
TITLE    2 HYDROLASE, BPHD FROM BURKHOLDERIA XENOVORANS LB400, IN
TITLE    3 COMPLEX WITH ITS SUBSTRATE HOPDA
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: 2-HYDROXY-6-OXO-6-PHENYLHEXA-2,4-DIENOATE
COMPND   3 HYDROLASE;
COMPND   4 CHAIN: A;
COMPND   5 EC: 3.7.1.-;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: BURKHOLDERIA XENOVORANS LB400;
SOURCE   3 ORGANISM_TAXID: 266265;
SOURCE   4 STRAIN: LB400;
SOURCE   5 GENE: BPHD;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    C-C BOND HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.BHOWMIK,J.T.BOLIN
REVDAT   3   24-FEB-09 2PUJ    1       VERSN
REVDAT   2   17-JUL-07 2PUJ    1       JRNL
REVDAT   1   22-MAY-07 2PUJ    0
JRNL        AUTH   G.P.HORSMAN,S.BHOWMIK,S.Y.SEAH,P.KUMAR,J.T.BOLIN,
JRNL        AUTH 2 L.D.ELTIS
JRNL        TITL   THE TAUTOMERIC HALF-REACTION OF BPHD, A C-C BOND
JRNL        TITL 2 HYDROLASE: KINETIC AND STRUCTURAL EVIDENCE
JRNL        TITL 3 SUPPORTING A KEY ROLE FOR HISTIDINE 265 OF THE
JRNL        TITL 4 CATALYTIC TRIAD.
JRNL        REF    J.BIOL.CHEM.                  V. 282 19894 2007
JRNL        REFN                   ISSN 0021-9258
JRNL        PMID   17442675
JRNL        DOI    10.1074/JBC.M702237200
REMARK   1
REMARK   2
REMARK   2 RESOLUTION.    1.57 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.2.0019
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.57
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 82.48
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.1
REMARK   3   NUMBER OF REFLECTIONS             : 39930
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.176
REMARK   3   R VALUE            (WORKING SET) : 0.175
REMARK   3   FREE R VALUE                     : 0.202
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 2123
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.57
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.61
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2891
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 98.67
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2520
REMARK   3   BIN FREE R VALUE SET COUNT          : 160
REMARK   3   BIN FREE R VALUE                    : 0.3010
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2259
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 24
REMARK   3   SOLVENT ATOMS            : 153
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 24.29
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.44000
REMARK   3    B22 (A**2) : 0.44000
REMARK   3    B33 (A**2) : -0.89000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.080
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.080
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.053
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 1.486
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.970
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.962
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  2343 ; 0.010 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  3177 ; 1.210 ; 1.960
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   294 ; 5.274 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   111 ;37.500 ;24.054
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   392 ;13.542 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    14 ;16.069 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   333 ; 0.077 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  1829 ; 0.005 ; 0.020
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  1123 ; 0.202 ; 0.200
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  1598 ; 0.306 ; 0.200
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):   125 ; 0.103 ; 0.200
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):     3 ; 0.163 ; 0.200
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):    54 ; 0.192 ; 0.200
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):    21 ; 0.182 ; 0.200
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  1417 ; 3.519 ; 2.000
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  2267 ; 5.109 ; 3.000
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):   968 ; 9.400 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):   904 ;12.483 ; 3.000
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : BABINET MODEL WITH MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3  RIDING POSITIONS
REMARK   4
REMARK   4 2PUJ COMPLIES WITH FORMAT V. 3.15, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 14-MAY-07.
REMARK 100 THE RCSB ID CODE IS RCSB042785.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 28-OCT-05
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 21-ID-D
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.00
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARMOSAIC 300 MM CCD
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 42069
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.570
REMARK 200  RESOLUTION RANGE LOW       (A) : 82.500
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.1
REMARK 200  DATA REDUNDANCY                : 9.300
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : 8.80000
REMARK 200   FOR THE DATA SET  : 21.2000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.57
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.61
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.1
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.90
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : 45.80000
REMARK 200   FOR SHELL         : 3.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: PDB ENTRY 2OG1
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 47.27
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.33
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 1.9 M SODIUM MALONATE, PH 6.5,
REMARK 280  VAPOR DIFFUSION, SITTING DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: I 41 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       3555   -Y,X+1/2,Z+1/4
REMARK 290       4555   Y+1/2,-X,Z+3/4
REMARK 290       5555   -X+1/2,Y,-Z+3/4
REMARK 290       6555   X,-Y+1/2,-Z+1/4
REMARK 290       7555   Y+1/2,X+1/2,-Z+1/2
REMARK 290       8555   -Y,-X,-Z
REMARK 290       9555   X+1/2,Y+1/2,Z+1/2
REMARK 290      10555   -X,-Y,Z
REMARK 290      11555   -Y+1/2,X,Z+3/4
REMARK 290      12555   Y,-X+1/2,Z+1/4
REMARK 290      13555   -X,Y+1/2,-Z+1/4
REMARK 290      14555   X+1/2,-Y,-Z+3/4
REMARK 290      15555   Y,X,-Z
REMARK 290      16555   -Y+1/2,-X+1/2,-Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000       58.40250
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       43.76400
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       21.88200
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       58.40250
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000       65.64600
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       65.64600
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       58.40250
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       21.88200
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000       58.40250
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       43.76400
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   9  1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY2   9  0.000000  1.000000  0.000000       58.40250
REMARK 290   SMTRY3   9  0.000000  0.000000  1.000000       43.76400
REMARK 290   SMTRY1  10 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  10  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3  10  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1  11  0.000000 -1.000000  0.000000       58.40250
REMARK 290   SMTRY2  11  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  11  0.000000  0.000000  1.000000       65.64600
REMARK 290   SMTRY1  12  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2  12 -1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY3  12  0.000000  0.000000  1.000000       21.88200
REMARK 290   SMTRY1  13 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  13  0.000000  1.000000  0.000000       58.40250
REMARK 290   SMTRY3  13  0.000000  0.000000 -1.000000       21.88200
REMARK 290   SMTRY1  14  1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY2  14  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3  14  0.000000  0.000000 -1.000000       65.64600
REMARK 290   SMTRY1  15  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2  15  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3  15  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1  16  0.000000 -1.000000  0.000000       58.40250
REMARK 290   SMTRY2  16 -1.000000  0.000000  0.000000       58.40250
REMARK 290   SMTRY3  16  0.000000  0.000000 -1.000000       43.76400
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 300 REMARK: THE BIOLOGICAL ASSEMBLY IS A TETRAMER GENERATED FROM THE
REMARK 300 MONOMER IN THE ASYMMETRIC UNIT BY THE OPERATIONS: X, Y, Z; -X+1/
REMARK 300 2, -Y+1/2, Z+1/2; -Y, X+1/2, Z+1/4; Y+1/2, -X, Z+3/4;
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TETRAMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  0.000000 -1.000000  0.000000        0.00000
REMARK 350   BIOMT2   2 -1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 350   BIOMT1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   3  0.000000 -1.000000  0.000000        0.00000
REMARK 350   BIOMT3   3  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   4  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT2   4  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     THR A     2
REMARK 465     ALA A     3
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    LYS A  18     -110.39     45.27
REMARK 500    ASN A  75     -123.92     45.29
REMARK 500    ALA A 112     -120.69     60.17
REMARK 500    LEU A 140       34.65    -85.12
REMARK 500    PHE A 145      -37.00   -130.10
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620  (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620  SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA A 287  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH A 429   O
REMARK 620 2 MLI A 288   O7  166.6
REMARK 620 3 MLI A 288   O9  101.6  81.7
REMARK 620 4 HOH A 340   O    89.3 100.2 120.4
REMARK 620 5 ARG A 105   O    90.6  76.1 107.2 131.4
REMARK 620 N                    1     2     3     4
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA A 287
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MLI A 288
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HPZ A 289
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 2OG1   RELATED DB: PDB
REMARK 900 RELATED ID: 1J1I   RELATED DB: PDB
REMARK 900 RELATED ID: 2D0D   RELATED DB: PDB
REMARK 900 RELATED ID: 1C4X   RELATED DB: PDB
REMARK 900 RELATED ID: 1U2E   RELATED DB: PDB
REMARK 900 RELATED ID: 2PU5   RELATED DB: PDB
REMARK 900 RELATED ID: 2PU6   RELATED DB: PDB
REMARK 900 RELATED ID: 2PU7   RELATED DB: PDB
REMARK 900 RELATED ID: 2PUH   RELATED DB: PDB
DBREF  2PUJ A    1   286  UNP    P47229   BPHD_BURXL       1    286
SEQADV 2PUJ ALA A  112  UNP  P47229    SER   112 ENGINEERED
SEQADV 2PUJ ALA A  265  UNP  P47229    HIS   265 ENGINEERED
SEQRES   1 A  286  MET THR ALA LEU THR GLU SER SER THR SER LYS PHE VAL
SEQRES   2 A  286  LYS ILE ASN GLU LYS GLY PHE SER ASP PHE ASN ILE HIS
SEQRES   3 A  286  TYR ASN GLU ALA GLY ASN GLY GLU THR VAL ILE MET LEU
SEQRES   4 A  286  HIS GLY GLY GLY PRO GLY ALA GLY GLY TRP SER ASN TYR
SEQRES   5 A  286  TYR ARG ASN VAL GLY PRO PHE VAL ASP ALA GLY TYR ARG
SEQRES   6 A  286  VAL ILE LEU LYS ASP SER PRO GLY PHE ASN LYS SER ASP
SEQRES   7 A  286  ALA VAL VAL MET ASP GLU GLN ARG GLY LEU VAL ASN ALA
SEQRES   8 A  286  ARG ALA VAL LYS GLY LEU MET ASP ALA LEU ASP ILE ASP
SEQRES   9 A  286  ARG ALA HIS LEU VAL GLY ASN ALA MET GLY GLY ALA THR
SEQRES  10 A  286  ALA LEU ASN PHE ALA LEU GLU TYR PRO ASP ARG ILE GLY
SEQRES  11 A  286  LYS LEU ILE LEU MET GLY PRO GLY GLY LEU GLY PRO SER
SEQRES  12 A  286  MET PHE ALA PRO MET PRO MET GLU GLY ILE LYS LEU LEU
SEQRES  13 A  286  PHE LYS LEU TYR ALA GLU PRO SER TYR GLU THR LEU LYS
SEQRES  14 A  286  GLN MET LEU GLN VAL PHE LEU TYR ASP GLN SER LEU ILE
SEQRES  15 A  286  THR GLU GLU LEU LEU GLN GLY ARG TRP GLU ALA ILE GLN
SEQRES  16 A  286  ARG GLN PRO GLU HIS LEU LYS ASN PHE LEU ILE SER ALA
SEQRES  17 A  286  GLN LYS ALA PRO LEU SER THR TRP ASP VAL THR ALA ARG
SEQRES  18 A  286  LEU GLY GLU ILE LYS ALA LYS THR PHE ILE THR TRP GLY
SEQRES  19 A  286  ARG ASP ASP ARG PHE VAL PRO LEU ASP HIS GLY LEU LYS
SEQRES  20 A  286  LEU LEU TRP ASN ILE ASP ASP ALA ARG LEU HIS VAL PHE
SEQRES  21 A  286  SER LYS CYS GLY ALA TRP ALA GLN TRP GLU HIS ALA ASP
SEQRES  22 A  286  GLU PHE ASN ARG LEU VAL ILE ASP PHE LEU ARG HIS ALA
HET     NA  A 287       1
HET    MLI  A 288       7
HET    HPZ  A 289      16
HETNAM      NA SODIUM ION
HETNAM     MLI MALONATE ION
HETNAM     HPZ (2E,4E)-2-HYDROXY-6-OXO-6-PHENYLHEXA-2,4-DIENOIC ACID
FORMUL   2   NA    NA 1+
FORMUL   3  MLI    C3 H2 O4 2-
FORMUL   4  HPZ    C12 H10 O4
FORMUL   5  HOH   *153(H2 O)
HELIX    1   1 THR A    5  THR A    9  1                                   5
HELIX    2   2 GLY A   47  TYR A   53  1                                   7
HELIX    3   3 ASN A   55  ALA A   62  1                                   8
HELIX    4   4 GLN A   85  LEU A  101  1                                  17
HELIX    5   5 ALA A  112  TYR A  125  1                                  14
HELIX    6   6 MET A  150  GLU A  162  1                                  13
HELIX    7   7 SER A  164  LEU A  176  1                                  13
HELIX    8   8 ASP A  178  ILE A  182  5                                   5
HELIX    9   9 THR A  183  GLN A  197  1                                  15
HELIX   10  10 GLN A  197  ALA A  211  1                                  15
HELIX   11  11 PRO A  212  ASP A  217  5                                   6
HELIX   12  12 VAL A  218  ILE A  225  5                                   8
HELIX   13  13 LEU A  242  ILE A  252  1                                  11
HELIX   14  14 TRP A  266  HIS A  271  1                                   6
HELIX   15  15 HIS A  271  ALA A  286  1                                  16
SHEET    1   A 8 SER A  10  ILE A  15  0
SHEET    2   A 8 PHE A  23  ALA A  30 -1  O  TYR A  27   N  LYS A  11
SHEET    3   A 8 ARG A  65  LYS A  69 -1  O  LEU A  68   N  ASN A  28
SHEET    4   A 8 THR A  35  LEU A  39  1  N  VAL A  36   O  ARG A  65
SHEET    5   A 8 ALA A 106  ASN A 111  1  O  VAL A 109   N  LEU A  39
SHEET    6   A 8 ILE A 129  MET A 135  1  O  ILE A 133   N  LEU A 108
SHEET    7   A 8 THR A 229  GLY A 234  1  O  THR A 232   N  LEU A 134
SHEET    8   A 8 ALA A 255  PHE A 260  1  O  ARG A 256   N  ILE A 231
LINK        NA    NA A 287                 O   HOH A 429     1555   1555  2.29
LINK        NA    NA A 287                 O7  MLI A 288     1555   1555  2.55
LINK        NA    NA A 287                 O9  MLI A 288     1555   1555  2.32
LINK        NA    NA A 287                 O   HOH A 340     1555   1555  2.17
LINK        NA    NA A 287                 O   ARG A 105     1555   4454  2.43
CISPEP   1 MET A  148    PRO A  149          0         5.40
SITE     1 AC1  4 ARG A 105  MLI A 288  HOH A 340  HOH A 429
SITE     1 AC2 10 GLU A  34  THR A  35  ARG A  65  ILE A 103
SITE     2 AC2 10 ASP A 104  ARG A 105  SER A 180  ILE A 182
SITE     3 AC2 10  NA A 287  HOH A 311
SITE     1 AC3 14 GLY A  41  GLY A  42  GLY A  43  ALA A  46
SITE     2 AC3 14 ALA A 112  MET A 113  GLY A 138  ILE A 153
SITE     3 AC3 14 LEU A 156  MET A 171  PHE A 175  ARG A 190
SITE     4 AC3 14 VAL A 240  HOH A 406
CRYST1  116.805  116.805   87.528  90.00  90.00  90.00 I 41 2 2     16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008561  0.000000  0.000000        0.00000
SCALE2      0.000000  0.008561  0.000000        0.00000
SCALE3      0.000000  0.000000  0.011425        0.00000
TER    2260      ALA A 286
MASTER      357    0    3   15    8    0    8    6 2436    1   26   22
END