longtext: 2QZP-pdb

content
HEADER    HYDROLASE                               17-AUG-07   2QZP
TITLE     CRYSTAL STRUCTURE OF MUTATION OF AN ACYLPTIDE
TITLE    2 HYDROLASE/ESTERASE FROM AEROPYRUM PERNIX K1
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACYLAMINO-ACID-RELEASING ENZYME;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: RESIDUES 21-582;
COMPND   5 SYNONYM: AARE, ACYL-PEPTIDE HYDROLASE, APH, ACYLAMINOACYL-
COMPND   6 PEPTIDASE;
COMPND   7 EC: 3.4.19.1;
COMPND   8 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: AEROPYRUM PERNIX;
SOURCE   3 ORGANISM_COMMON: ARCHAEA;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   5 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   6 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   7 EXPRESSION_SYSTEM_PLASMID: PET11A
KEYWDS    CRYSTAL STRUCTURE, TRUNCATED, ACYLPEPTIDE HYDROLASE,
KEYWDS   2 AEROPYRUM PERNIX K1, CYTOPLASM
EXPDTA    X-RAY DIFFRACTION
AUTHOR    H.F.ZHANG,B.S.ZHENG,Z.RAO
REVDAT   1   02-OCT-07 2QZP    0
JRNL        AUTH   H.F.ZHANG,B.S.ZHENG,Z.RAO
JRNL        TITL   EXPRESSION, PURIFICATION AND CRYSTAL STRUCTURE OF
JRNL        TITL 2 A TRUNCATED ACYLPEPTIDE HYDROLASE FROM AEROPYRUM
JRNL        TITL 3 PERNIX K1
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   1
REMARK   2
REMARK   2 RESOLUTION. 2.70 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.0
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.70
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 48.11
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 92.0
REMARK   3   NUMBER OF REFLECTIONS             : 27467
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.226
REMARK   3   FREE R VALUE                     : 0.277
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 1393
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.007
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.70
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.82
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 89.60
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 3106
REMARK   3   BIN R VALUE           (WORKING SET) : 0.3160
REMARK   3   BIN FREE R VALUE                    : 0.4030
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 5.40
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 176
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.030
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 8515
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 0
REMARK   3   SOLVENT ATOMS            : 366
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 45.00
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 30.14
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -1.44000
REMARK   3    B22 (A**2) : -6.83000
REMARK   3    B33 (A**2) : 8.27000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.33
REMARK   3   ESD FROM SIGMAA              (A) : 0.41
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.40
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.52
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.009
REMARK   3   BOND ANGLES            (DEGREES) : 1.60
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : CNS BULK SOLVENT MODEL USED
REMARK   3   KSOL        : 0.34
REMARK   3   BSOL        : 67.97
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM
REMARK   3  PARAMETER FILE  2  : WATER_REP.PARAM
REMARK   3  PARAMETER FILE  3  : NULL
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP
REMARK   3  TOPOLOGY FILE  2   : WATER_REP.TOP
REMARK   3  TOPOLOGY FILE  3   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 2QZP COMPLIES WITH FORMAT V. 3.1, 1-AUG-2007
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ.
REMARK 100 THE RCSB ID CODE IS RCSB044247.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 05-NOV-2004
REMARK 200  TEMPERATURE           (KELVIN) : 100.0
REMARK 200  PH                             : 4.60
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : ROTATING ANODE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU MICROMAX-007
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418
REMARK 200  MONOCHROMATOR                  : OSMIC MIRROR
REMARK 200  OPTICS                         : OSMIC MIRROR
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : MAR SCANNER 345 MM PLATE
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 35191
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.700
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.0
REMARK 200  DATA REDUNDANCY                : 5.800
REMARK 200  R MERGE                    (I) : 0.17700
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.70
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.80
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.80
REMARK 200  R MERGE FOR SHELL          (I) : 0.77700
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: CNS
REMARK 200 STARTING MODEL: 1VE6
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 47.13
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.33
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NAAC, PEG3000, PH 4.6, VAPOR
REMARK 280  DIFFUSION, HANGING DROP, TEMPERATURE 291K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   1/2-X,-Y,1/2+Z
REMARK 290       3555   -X,1/2+Y,1/2-Z
REMARK 290       4555   1/2+X,1/2-Y,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       31.55850
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       81.79700
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       51.09150
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       81.79700
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       31.55850
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       51.09150
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMER
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMER
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C  SSEQI
REMARK 465     ALA A    21
REMARK 465     ARG A   582
REMARK 465     ARG B   582
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI
REMARK 500   OG1  THR A   130     OD1  ASP A   132              2.10
REMARK 500   O    ARG A   511     O    HOH A   630              2.10
REMARK 500   O    ARG B   327     O    HOH B   720              2.11
REMARK 500   N    SER A   116     O    HOH A   677              2.13
REMARK 500   O    GLU A   562     N    ALA A   564              2.17
REMARK 500   OD2  ASP A   563     O    HOH A   585              2.18
REMARK 500   N    GLY A   417     OE2  GLU A   419              2.18
REMARK 500   O    THR B   451     O    HOH B   615              2.18
REMARK 500   O    LYS B   463     O    HOH B   689              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP A  32       18.14     45.66
REMARK 500    SER A  42      171.81    -48.65
REMARK 500    GLU A  43       79.34      4.09
REMARK 500    ARG A  61      -72.77    -90.75
REMARK 500    ILE A  64     -157.55    -87.52
REMARK 500    ASN A  65      -39.35   -132.19
REMARK 500    TYR A  72      131.60    -13.45
REMARK 500    ARG A  76      155.53    176.66
REMARK 500    LYS A  85       79.23     66.24
REMARK 500    ALA A  87       -9.09    -56.09
REMARK 500    GLU A  88        6.77     81.35
REMARK 500    ARG A  99       73.74   -112.01
REMARK 500    LEU A 106       82.62    -57.19
REMARK 500    ALA A 108       37.27    -82.15
REMARK 500    MET A 112     -158.34   -157.05
REMARK 500    SER A 116      158.36    166.25
REMARK 500    PHE A 126      161.27    173.83
REMARK 500    ARG A 133      162.70    177.49
REMARK 500    PHE A 153      119.81    -33.69
REMARK 500    LEU A 187      138.63   -177.23
REMARK 500    SER A 199      155.11    171.46
REMARK 500    SER A 201      131.78   -171.39
REMARK 500    ARG A 216     -102.17    -76.90
REMARK 500    GLU A 217     -130.40   -121.60
REMARK 500    THR A 222       72.68   -118.28
REMARK 500    ALA A 247      114.10   -161.71
REMARK 500    ARG A 264      157.59    179.45
REMARK 500    GLU A 276       99.56    -56.94
REMARK 500    ARG A 287      126.40    -31.63
REMARK 500    THR A 297     -169.63   -127.99
REMARK 500    THR A 300      135.27   -173.98
REMARK 500    PRO A 306      132.74    -38.43
REMARK 500    PRO A 323      171.02    -58.37
REMARK 500    ASP A 342       20.92    -74.16
REMARK 500    PRO A 347      101.17    -58.09
REMARK 500    SER A 353       76.23   -103.11
REMARK 500    ARG A 355       22.47    -75.13
REMARK 500    PHE A 371       52.46    -62.31
REMARK 500    ALA A 372     -154.12   -128.05
REMARK 500    GLU A 373      161.66    176.86
REMARK 500    ASN A 396       70.88   -104.54
REMARK 500    TRP A 429      -70.86    -33.84
REMARK 500    ALA A 430      -68.43    -26.60
REMARK 500    SER A 445     -121.25     67.27
REMARK 500    PRO A 459       63.23    -61.55
REMARK 500    GLU A 479        5.83    -69.23
REMARK 500    LEU A 492       -3.08    -58.56
REMARK 500    GLU A 498       -5.99    -44.30
REMARK 500    ARG A 501      -61.84    -97.45
REMARK 500    PRO A 515      179.34    -55.41
REMARK 500    LEU A 516      126.41   -172.18
REMARK 500    ARG A 526      -61.67    -98.89
REMARK 500    ARG A 542       28.13    -79.55
REMARK 500    ASP A 553       40.78     37.71
REMARK 500    GLU A 562      -73.81    -56.93
REMARK 500    ASP A 563        8.49    -44.12
REMARK 500    ILE A 567       62.66   -104.79
REMARK 500    LEU A 568        6.09   -179.40
REMARK 500    ARG A 579      -61.16    -23.47
REMARK 500    GLU A 580       41.92    -74.62
REMARK 500    ASP B  34       82.06   -150.23
REMARK 500    LYS B  35      109.35   -160.38
REMARK 500    PHE B  41       79.95   -107.13
REMARK 500    SER B  42     -154.72   -167.67
REMARK 500    SER B  45     -140.77   -177.41
REMARK 500    ASN B  60     -144.13    -69.47
REMARK 500    ARG B  61      -87.76   -159.24
REMARK 500    LEU B  68      176.02    -57.51
REMARK 500    TYR B  72      122.07    -25.47
REMARK 500    VAL B  74      145.47    -33.67
REMARK 500    GLU B  88       11.95     83.78
REMARK 500    THR B  97      -37.94    -37.06
REMARK 500    ARG B  99       83.03   -150.21
REMARK 500    GLU B 107      -22.96    -35.06
REMARK 500    MET B 112     -154.10   -136.12
REMARK 500    SER B 116     -161.89   -168.70
REMARK 500    THR B 120      -83.45    -83.69
REMARK 500    GLU B 122      -17.59   -165.66
REMARK 500    VAL B 124       79.50   -111.43
REMARK 500    ARG B 133      145.53   -173.89
REMARK 500    ASP B 140     -168.96   -163.01
REMARK 500    ARG B 160      103.43   -165.05
REMARK 500    SER B 180     -177.92    179.88
REMARK 500    LEU B 187      119.30    169.35
REMARK 500    SER B 201      121.36   -172.08
REMARK 500    MET B 206       47.66    101.61
REMARK 500    ALA B 210       94.22    174.72
REMARK 500    ASP B 232      169.91    -40.86
REMARK 500    LYS B 238       64.61   -109.67
REMARK 500    ARG B 244       60.73     25.58
REMARK 500    THR B 249      -72.46    -68.02
REMARK 500    ASP B 256       -2.52    -46.12
REMARK 500    ARG B 292       49.54     77.67
REMARK 500    SER B 301      159.32    168.82
REMARK 500    LEU B 302      -35.73    -39.85
REMARK 500    THR B 304      120.26    179.68
REMARK 500    PRO B 306      104.91    -49.76
REMARK 500    PRO B 323      163.93    -39.63
REMARK 500    PRO B 361      108.42    -52.66
REMARK 500    HIS B 367     -164.59    -69.39
REMARK 500    PHE B 371       72.79    -47.07
REMARK 500    SER B 375     -151.00   -131.28
REMARK 500    ASP B 376       48.23    -97.61
REMARK 500    THR B 380      -75.00    -39.10
REMARK 500    PHE B 381      -38.09    -37.95
REMARK 500    ASN B 396       58.24   -108.63
REMARK 500    CYS B 416       68.82     69.97
REMARK 500    ALA B 427      -70.94    -53.88
REMARK 500    TRP B 429      -72.35    -35.40
REMARK 500    ALA B 430      -27.10    -39.41
REMARK 500    SER B 445     -124.47     53.33
REMARK 500    VAL B 472      -73.86    -67.51
REMARK 500    LEU B 480       41.94   -106.63
REMARK 500    SER B 496      125.77     -5.72
REMARK 500    GLU B 498      -44.18    -27.79
REMARK 500    PRO B 505      -10.30    -40.21
REMARK 500    ASP B 510        1.85    -64.96
REMARK 500    PRO B 521      106.93    -57.58
REMARK 500    LYS B 530      -72.78    -58.31
REMARK 500    PRO B 531      -37.35    -39.94
REMARK 500    LYS B 544      110.90    -16.54
REMARK 500    PHE B 546      131.89   -172.87
REMARK 500    THR B 560      160.46    167.03
REMARK 500    GLU B 562       33.79    -68.71
REMARK 500    ASP B 563       15.51   -154.54
REMARK 500    LYS B 566        0.99    -58.55
REMARK 500    GLU B 580       14.46    -59.77
REMARK 500
REMARK 500 REMARK: NULL
DBREF  2QZP A   21   582  UNP    Q9YBQ2   APEH_AERPE      21    582
DBREF  2QZP B   21   582  UNP    Q9YBQ2   APEH_AERPE      21    582
SEQRES   1 A  562  ALA VAL GLU LYS TYR SER LEU GLN GLY VAL VAL ASP GLY
SEQRES   2 A  562  ASP LYS LEU LEU VAL VAL GLY PHE SER GLU GLY SER VAL
SEQRES   3 A  562  ASN ALA TYR LEU TYR ASP GLY GLY GLU THR VAL LYS LEU
SEQRES   4 A  562  ASN ARG GLU PRO ILE ASN SER VAL LEU ASP PRO HIS TYR
SEQRES   5 A  562  GLY VAL GLY ARG VAL ILE LEU VAL ARG ASP VAL SER LYS
SEQRES   6 A  562  GLY ALA GLU GLN HIS ALA LEU PHE LYS VAL ASN THR SER
SEQRES   7 A  562  ARG PRO GLY GLU GLU GLN ARG LEU GLU ALA VAL LYS PRO
SEQRES   8 A  562  MET ARG ILE LEU SER GLY VAL ASP THR GLY GLU ALA VAL
SEQRES   9 A  562  VAL PHE THR GLY ALA THR GLU ASP ARG VAL ALA LEU TYR
SEQRES  10 A  562  ALA LEU ASP GLY GLY GLY LEU ARG GLU LEU ALA ARG LEU
SEQRES  11 A  562  PRO GLY PHE GLY PHE VAL SER ASP ILE ARG GLY ASP LEU
SEQRES  12 A  562  ILE ALA GLY LEU GLY PHE PHE GLY GLY GLY ARG VAL SER
SEQRES  13 A  562  LEU PHE THR SER ASN LEU SER SER GLY GLY LEU ARG VAL
SEQRES  14 A  562  PHE ASP SER GLY GLU GLY SER PHE SER SER ALA SER ILE
SEQRES  15 A  562  SER PRO GLY MET LYS VAL THR ALA GLY LEU GLU THR ALA
SEQRES  16 A  562  ARG GLU ALA ARG LEU VAL THR VAL ASP PRO ARG ASP GLY
SEQRES  17 A  562  SER VAL GLU ASP LEU GLU LEU PRO SER LYS ASP PHE SER
SEQRES  18 A  562  SER TYR ARG PRO THR ALA ILE THR TRP LEU GLY TYR LEU
SEQRES  19 A  562  PRO ASP GLY ARG LEU ALA VAL VAL ALA ARG ARG GLU GLY
SEQRES  20 A  562  ARG SER ALA VAL PHE ILE ASP GLY GLU ARG VAL GLU ALA
SEQRES  21 A  562  PRO GLN GLY ASN HIS GLY ARG VAL VAL LEU TRP ARG GLY
SEQRES  22 A  562  LYS LEU VAL THR SER HIS THR SER LEU SER THR PRO PRO
SEQRES  23 A  562  ARG ILE VAL SER LEU PRO SER GLY GLU PRO LEU LEU GLU
SEQRES  24 A  562  GLY GLY LEU PRO GLU ASP LEU ARG ARG SER ILE ALA GLY
SEQRES  25 A  562  SER ARG LEU VAL TRP VAL GLU SER PHE ASP GLY SER ARG
SEQRES  26 A  562  VAL PRO THR TYR VAL LEU GLU SER GLY ARG ALA PRO THR
SEQRES  27 A  562  PRO GLY PRO THR VAL VAL LEU VAL HIS GLY GLY PRO PHE
SEQRES  28 A  562  ALA GLU ASP SER ASP SER TRP ASP THR PHE ALA ALA SER
SEQRES  29 A  562  LEU ALA ALA ALA GLY PHE HIS VAL VAL MET PRO ASN TYR
SEQRES  30 A  562  ARG GLY SER THR GLY TYR GLY GLU GLU TRP ARG LEU LYS
SEQRES  31 A  562  ILE ILE GLY ASP PRO CYS GLY GLY GLU LEU GLU ASP VAL
SEQRES  32 A  562  SER ALA ALA ALA ARG TRP ALA ARG GLU SER GLY LEU ALA
SEQRES  33 A  562  SER GLU LEU TYR ILE MET GLY TYR SER TYR GLY GLY TYR
SEQRES  34 A  562  MET THR LEU CYS ALA LEU THR MET LYS PRO GLY LEU PHE
SEQRES  35 A  562  LYS ALA GLY VAL ALA GLY ALA SER VAL VAL ASP TRP GLU
SEQRES  36 A  562  GLU MET TYR GLU LEU SER ASP ALA ALA PHE ARG ASN PHE
SEQRES  37 A  562  ILE GLU GLN LEU THR GLY GLY SER ARG GLU ILE MET ARG
SEQRES  38 A  562  SER ARG SER PRO ILE ASN HIS VAL ASP ARG ILE LYS GLU
SEQRES  39 A  562  PRO LEU ALA LEU ILE HIS PRO GLN ASN ASP SER ARG THR
SEQRES  40 A  562  PRO LEU LYS PRO LEU LEU ARG LEU MET GLY GLU LEU LEU
SEQRES  41 A  562  ALA ARG GLY LYS THR PHE GLU ALA HIS ILE ILE PRO ASP
SEQRES  42 A  562  ALA GLY HIS ALA ILE ASN THR MET GLU ASP ALA VAL LYS
SEQRES  43 A  562  ILE LEU LEU PRO ALA VAL PHE PHE LEU ALA THR GLN ARG
SEQRES  44 A  562  GLU ARG ARG
SEQRES   1 B  562  ALA VAL GLU LYS TYR SER LEU GLN GLY VAL VAL ASP GLY
SEQRES   2 B  562  ASP LYS LEU LEU VAL VAL GLY PHE SER GLU GLY SER VAL
SEQRES   3 B  562  ASN ALA TYR LEU TYR ASP GLY GLY GLU THR VAL LYS LEU
SEQRES   4 B  562  ASN ARG GLU PRO ILE ASN SER VAL LEU ASP PRO HIS TYR
SEQRES   5 B  562  GLY VAL GLY ARG VAL ILE LEU VAL ARG ASP VAL SER LYS
SEQRES   6 B  562  GLY ALA GLU GLN HIS ALA LEU PHE LYS VAL ASN THR SER
SEQRES   7 B  562  ARG PRO GLY GLU GLU GLN ARG LEU GLU ALA VAL LYS PRO
SEQRES   8 B  562  MET ARG ILE LEU SER GLY VAL ASP THR GLY GLU ALA VAL
SEQRES   9 B  562  VAL PHE THR GLY ALA THR GLU ASP ARG VAL ALA LEU TYR
SEQRES  10 B  562  ALA LEU ASP GLY GLY GLY LEU ARG GLU LEU ALA ARG LEU
SEQRES  11 B  562  PRO GLY PHE GLY PHE VAL SER ASP ILE ARG GLY ASP LEU
SEQRES  12 B  562  ILE ALA GLY LEU GLY PHE PHE GLY GLY GLY ARG VAL SER
SEQRES  13 B  562  LEU PHE THR SER ASN LEU SER SER GLY GLY LEU ARG VAL
SEQRES  14 B  562  PHE ASP SER GLY GLU GLY SER PHE SER SER ALA SER ILE
SEQRES  15 B  562  SER PRO GLY MET LYS VAL THR ALA GLY LEU GLU THR ALA
SEQRES  16 B  562  ARG GLU ALA ARG LEU VAL THR VAL ASP PRO ARG ASP GLY
SEQRES  17 B  562  SER VAL GLU ASP LEU GLU LEU PRO SER LYS ASP PHE SER
SEQRES  18 B  562  SER TYR ARG PRO THR ALA ILE THR TRP LEU GLY TYR LEU
SEQRES  19 B  562  PRO ASP GLY ARG LEU ALA VAL VAL ALA ARG ARG GLU GLY
SEQRES  20 B  562  ARG SER ALA VAL PHE ILE ASP GLY GLU ARG VAL GLU ALA
SEQRES  21 B  562  PRO GLN GLY ASN HIS GLY ARG VAL VAL LEU TRP ARG GLY
SEQRES  22 B  562  LYS LEU VAL THR SER HIS THR SER LEU SER THR PRO PRO
SEQRES  23 B  562  ARG ILE VAL SER LEU PRO SER GLY GLU PRO LEU LEU GLU
SEQRES  24 B  562  GLY GLY LEU PRO GLU ASP LEU ARG ARG SER ILE ALA GLY
SEQRES  25 B  562  SER ARG LEU VAL TRP VAL GLU SER PHE ASP GLY SER ARG
SEQRES  26 B  562  VAL PRO THR TYR VAL LEU GLU SER GLY ARG ALA PRO THR
SEQRES  27 B  562  PRO GLY PRO THR VAL VAL LEU VAL HIS GLY GLY PRO PHE
SEQRES  28 B  562  ALA GLU ASP SER ASP SER TRP ASP THR PHE ALA ALA SER
SEQRES  29 B  562  LEU ALA ALA ALA GLY PHE HIS VAL VAL MET PRO ASN TYR
SEQRES  30 B  562  ARG GLY SER THR GLY TYR GLY GLU GLU TRP ARG LEU LYS
SEQRES  31 B  562  ILE ILE GLY ASP PRO CYS GLY GLY GLU LEU GLU ASP VAL
SEQRES  32 B  562  SER ALA ALA ALA ARG TRP ALA ARG GLU SER GLY LEU ALA
SEQRES  33 B  562  SER GLU LEU TYR ILE MET GLY TYR SER TYR GLY GLY TYR
SEQRES  34 B  562  MET THR LEU CYS ALA LEU THR MET LYS PRO GLY LEU PHE
SEQRES  35 B  562  LYS ALA GLY VAL ALA GLY ALA SER VAL VAL ASP TRP GLU
SEQRES  36 B  562  GLU MET TYR GLU LEU SER ASP ALA ALA PHE ARG ASN PHE
SEQRES  37 B  562  ILE GLU GLN LEU THR GLY GLY SER ARG GLU ILE MET ARG
SEQRES  38 B  562  SER ARG SER PRO ILE ASN HIS VAL ASP ARG ILE LYS GLU
SEQRES  39 B  562  PRO LEU ALA LEU ILE HIS PRO GLN ASN ASP SER ARG THR
SEQRES  40 B  562  PRO LEU LYS PRO LEU LEU ARG LEU MET GLY GLU LEU LEU
SEQRES  41 B  562  ALA ARG GLY LYS THR PHE GLU ALA HIS ILE ILE PRO ASP
SEQRES  42 B  562  ALA GLY HIS ALA ILE ASN THR MET GLU ASP ALA VAL LYS
SEQRES  43 B  562  ILE LEU LEU PRO ALA VAL PHE PHE LEU ALA THR GLN ARG
SEQRES  44 B  562  GLU ARG ARG
FORMUL   3  HOH   *366(H2 O)
HELIX    1   1 ASP A  239  ARG A  244  1                                   6
HELIX    2   2 ASP A  379  ALA A  388  1                                  10
HELIX    3   3 GLY A  404  LEU A  409  1                                   6
HELIX    4   4 GLY A  417  SER A  433  1                                  17
HELIX    5   5 SER A  445  MET A  457  1                                  13
HELIX    6   6 ASP A  473  LEU A  480  1                                   8
HELIX    7   7 ASP A  482  THR A  493  1                                  12
HELIX    8   8 SER A  496  ARG A  503  1                                   8
HELIX    9   9 SER A  504  VAL A  509  5                                   6
HELIX   10  10 LEU A  529  ARG A  542  1                                  14
HELIX   11  11 MET A  561  LEU A  568  5                                   8
HELIX   12  12 LEU A  569  GLU A  580  1                                  12
HELIX   13  13 GLU B  324  SER B  329  1                                   6
HELIX   14  14 ASP B  379  ALA B  388  1                                  10
HELIX   15  15 GLY B  404  LEU B  409  1                                   6
HELIX   16  16 GLY B  417  SER B  433  1                                  17
HELIX   17  17 SER B  445  LYS B  458  1                                  14
HELIX   18  18 TRP B  474  LEU B  480  1                                   7
HELIX   19  19 ASP B  482  THR B  493  1                                  12
HELIX   20  20 SER B  496  ARG B  503  1                                   8
HELIX   21  21 SER B  504  ILE B  512  5                                   9
HELIX   22  22 PRO B  528  ARG B  542  1                                  15
HELIX   23  23 LEU B  568  GLU B  580  1                                  13
SHEET    1   A 4 TYR A  25  VAL A  31  0
SHEET    2   A 4 LYS A  35  GLY A  40 -1  O  LEU A  37   N  GLY A  29
SHEET    3   A 4 ASN A  47  TYR A  51 -1  O  TYR A  49   N  VAL A  38
SHEET    4   A 4 VAL A  57  LYS A  58 -1  O  VAL A  57   N  LEU A  50
SHEET    1   B 3 SER A  66  VAL A  67  0
SHEET    2   B 3 VAL A  77  ASP A  82 -1  O  VAL A  80   N  SER A  66
SHEET    3   B 3 HIS A  90  VAL A  95 -1  O  VAL A  95   N  VAL A  77
SHEET    1   C 5 ASP A  69  PRO A  70  0
SHEET    2   C 5 ARG A 113  ASP A 119  1  O  ASP A 119   N  ASP A  69
SHEET    3   C 5 VAL A 124  ALA A 129 -1  O  THR A 127   N  SER A 116
SHEET    4   C 5 VAL A 134  ASP A 140 -1  O  LEU A 139   N  VAL A 124
SHEET    5   C 5 GLY A 143  LEU A 150 -1  O  LEU A 147   N  LEU A 136
SHEET    1   D 4 GLY A 154  ARG A 160  0
SHEET    2   D 4 LEU A 163  GLY A 171 -1  O  ALA A 165   N  ASP A 158
SHEET    3   D 4 ARG A 174  ASN A 181 -1  O  SER A 176   N  GLY A 168
SHEET    4   D 4 ARG A 188  PHE A 190 -1  O  PHE A 190   N  LEU A 177
SHEET    1   E 4 GLY A 195  SER A 201  0
SHEET    2   E 4 VAL A 208  THR A 214 -1  O  GLU A 213   N  SER A 196
SHEET    3   E 4 ALA A 218  VAL A 223 -1  O  ARG A 219   N  LEU A 212
SHEET    4   E 4 VAL A 230  ASP A 232 -1  O  GLU A 231   N  THR A 222
SHEET    1   F 4 TRP A 250  TYR A 253  0
SHEET    2   F 4 LEU A 259  ARG A 265 -1  O  VAL A 262   N  TRP A 250
SHEET    3   F 4 ARG A 268  ILE A 273 -1  O  ARG A 268   N  ARG A 265
SHEET    4   F 4 GLU A 276  VAL A 278 -1  O  VAL A 278   N  VAL A 271
SHEET    1   G 4 VAL A 289  TRP A 291  0
SHEET    2   G 4 LYS A 294  THR A 297 -1  O  LYS A 294   N  TRP A 291
SHEET    3   G 4 ILE A 308  SER A 310 -1  O  VAL A 309   N  THR A 297
SHEET    4   G 4 PRO A 316  LEU A 318 -1  O  LEU A 318   N  ILE A 308
SHEET    1   H12 ARG A 334  GLU A 339  0
SHEET    2   H12 ARG A 345  LEU A 351 -1  O  VAL A 350   N  ARG A 334
SHEET    3   H12 HIS A 391  PRO A 395 -1  O  MET A 394   N  TYR A 349
SHEET    4   H12 GLY A 360  VAL A 366  1  N  VAL A 363   O  HIS A 391
SHEET    5   H12 ALA A 436  TYR A 444  1  O  TYR A 440   N  VAL A 364
SHEET    6   H12 ALA A 464  GLY A 468  1  O  ALA A 464   N  ILE A 441
SHEET    7   H12 LEU A 516  PRO A 521  1  O  ALA A 517   N  ALA A 467
SHEET    8   H12 GLU A 547  ILE A 551  1  O  HIS A 549   N  HIS A 520
SHEET    9   H12 PHE B 546  ILE B 551 -1  O  ALA B 548   N  ILE A 550
SHEET   10   H12 LEU B 516  PRO B 521  1  N  HIS B 520   O  HIS B 549
SHEET   11   H12 ALA B 464  GLY B 468  1  N  ALA B 467   O  ALA B 517
SHEET   12   H12 TYR B 440  TYR B 444  1  N  ILE B 441   O  VAL B 466
SHEET    1   I 3 LYS B  24  TYR B  25  0
SHEET    2   I 3 LEU B  36  PHE B  41 -1  O  PHE B  41   N  LYS B  24
SHEET    3   I 3 ASN B  47  TYR B  51 -1  O  ASN B  47   N  GLY B  40
SHEET    1   J 4 SER B  66  VAL B  67  0
SHEET    2   J 4 ARG B  76  ASP B  82 -1  O  VAL B  80   N  SER B  66
SHEET    3   J 4 HIS B  90  ASN B  96 -1  O  PHE B  93   N  LEU B  79
SHEET    4   J 4 GLN B 104  ARG B 105 -1  O  GLN B 104   N  LYS B  94
SHEET    1   K 4 ASP B  69  PRO B  70  0
SHEET    2   K 4 VAL B 118  ASP B 119  1  O  ASP B 119   N  ASP B  69
SHEET    3   K 4 VAL B 124  ALA B 129 -1  O  VAL B 125   N  VAL B 118
SHEET    4   K 4 ARG B 113  ILE B 114 -1  N  ARG B 113   O  ALA B 129
SHEET    1   L 5 ASP B  69  PRO B  70  0
SHEET    2   L 5 VAL B 118  ASP B 119  1  O  ASP B 119   N  ASP B  69
SHEET    3   L 5 VAL B 124  ALA B 129 -1  O  VAL B 125   N  VAL B 118
SHEET    4   L 5 VAL B 134  ASP B 140 -1  O  TYR B 137   N  PHE B 126
SHEET    5   L 5 GLY B 143  LEU B 150 -1  O  GLY B 143   N  ASP B 140
SHEET    1   M 4 PHE B 153  ARG B 160  0
SHEET    2   M 4 LEU B 163  GLY B 171 -1  O  ALA B 165   N  ASP B 158
SHEET    3   M 4 ARG B 174  PHE B 178 -1  O  SER B 176   N  GLY B 168
SHEET    4   M 4 VAL B 189  PHE B 190 -1  O  PHE B 190   N  LEU B 177
SHEET    1   N 3 GLY B 195  PHE B 197  0
SHEET    2   N 3 LEU B 212  THR B 214 -1  O  GLU B 213   N  SER B 196
SHEET    3   N 3 ALA B 218  ARG B 219 -1  O  ARG B 219   N  LEU B 212
SHEET    1   O 2 SER B 201  ILE B 202  0
SHEET    2   O 2 VAL B 208  THR B 209 -1  O  THR B 209   N  SER B 201
SHEET    1   P 4 ILE B 248  TYR B 253  0
SHEET    2   P 4 LEU B 259  ARG B 265 -1  O  VAL B 262   N  TRP B 250
SHEET    3   P 4 ARG B 268  ILE B 273 -1  O  PHE B 272   N  VAL B 261
SHEET    4   P 4 GLU B 276  ARG B 277 -1  O  GLU B 276   N  ILE B 273
SHEET    1   Q 4 VAL B 289  LEU B 290  0
SHEET    2   Q 4 LEU B 295  SER B 301 -1  O  VAL B 296   N  VAL B 289
SHEET    3   Q 4 THR B 304  SER B 310 -1  O  THR B 304   N  SER B 301
SHEET    4   Q 4 GLU B 315  LEU B 318 -1  O  GLU B 315   N  SER B 310
SHEET    1   R 5 ARG B 334  VAL B 338  0
SHEET    2   R 5 VAL B 346  LEU B 351 -1  O  VAL B 346   N  VAL B 338
SHEET    3   R 5 HIS B 391  PRO B 395 -1  O  VAL B 392   N  LEU B 351
SHEET    4   R 5 GLY B 360  VAL B 364  1  N  VAL B 363   O  VAL B 393
SHEET    5   R 5 ALA B 436  GLU B 438  1  O  GLU B 438   N  THR B 362
SSBOND   1 CYS A  416    CYS A  453                        1555   1555    2.03
SSBOND   2 CYS B  416    CYS B  453                        1555   1555    2.04
CISPEP   1 LEU A  311    PRO A  312          0        -0.50
CISPEP   2 THR A  358    PRO A  359          0        -0.05
CISPEP   3 GLY A  369    PRO A  370          0         0.42
CISPEP   4 LEU B  311    PRO B  312          0        -0.18
CISPEP   5 THR B  358    PRO B  359          0         0.36
CISPEP   6 GLY B  369    PRO B  370          0        -0.23
CRYST1   63.117  102.183  163.594  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.015844  0.000000  0.000000        0.00000
SCALE2      0.000000  0.009786  0.000000        0.00000
SCALE3      0.000000  0.000000  0.006113        0.00000
TER    4256      ARG A 581
TER    8517      ARG B 581
MASTER      392    0    0   23   78    0    0    6 8881    2    4   88
END