longtext: 2WG0-pdb

content
HEADER    HYDROLASE                               15-APR-09   2WG0
TITLE     AGED CONJUGATE OF TORPEDO CALIFORNICA ACETYLCHOLINESTERASE
TITLE    2 WITH SOMAN (OBTAINED BY IN CRYSTALLO AGING)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYLCHOLINESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: RESIDUES 22-558;
COMPND   5 SYNONYM: ACHE;
COMPND   6 EC: 3.1.1.7
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: TORPEDO CALIFORNICA;
SOURCE   3 ORGANISM_COMMON: PACIFIC ELECTRIC RAY;
SOURCE   4 ORGANISM_TAXID: 7787;
SOURCE   5 VARIANT: G2 FORM;
SOURCE   6 ORGAN: ELECTRIC ORGAN;
SOURCE   7 TISSUE: ELECTROPLAQUE
KEYWDS    SOMAN, AGING, SYNAPSE, MEMBRANE, HYDROLASE, CELL MEMBRANE,
KEYWDS   2 DISULFIDE BOND, SERINE ESTERASE, NEUROTRANSMITTER
KEYWDS   3 CLEAVAGE, CONFORMATIONAL FLEXIBILITY, NEUROTRANSMITTER
KEYWDS   4 DEGRADATION, ALPHA/BETA HYDROLASE, ALTERNATIVE SPLICING,
KEYWDS   5 ORGANOPHOSPHATE, SYNAPSE MEMBRANE, ALZHEIMER DISEASE,
KEYWDS   6 GPI-ANCHOR, LIPOPROTEIN, GLYCOPROTEIN, CELL JUNCTION
EXPDTA    X-RAY DIFFRACTION
AUTHOR    B.SANSON,F.NACHON,J.P.COLLETIER,M.T.FROMENT,L.TOKER,
AUTHOR   2 H.M.GREENBLATT,J.L.SUSSMAN,Y.ASHANI,P.MASSON,I.SILMAN,
AUTHOR   3 M.WEIK
REVDAT   1   11-AUG-09 2WG0    0
JRNL        AUTH   B.SANSON,F.NACHON,J.P.COLLETIER,M.T.FROMENT,
JRNL        AUTH 2 L.TOKER,H.M.GREENBLATT,J.L.SUSSMAN,Y.ASHANI,
JRNL        AUTH 3 P.MASSON,I.SILMAN,M.WEIK
JRNL        TITL   CRYSTALLOGRAPHIC SNAPSHOTS OF NONAGED AND AGED
JRNL        TITL 2 CONJUGATES OF SOMAN WITH ACETYLCHOLINESTERASE, AND
JRNL        TITL 3 OF A TERNARY COMPLEX OF THE AGED CONJUGATE WITH
JRNL        TITL 4 PRALIDOXIME (DAGGER).
JRNL        REF    J.MED.CHEM.                                2009
JRNL        REFN                   ESSN 1520-4804
JRNL        PMID   19642642
JRNL        DOI    10.1021/JM900433T
REMARK   2
REMARK   2 RESOLUTION.    2.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.5.0072
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 48.34
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NONE
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.38
REMARK   3   NUMBER OF REFLECTIONS             : 47754
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.17964
REMARK   3   R VALUE            (WORKING SET) : 0.17718
REMARK   3   FREE R VALUE                     : 0.22892
REMARK   3   FREE R VALUE TEST SET SIZE   (%) :  4.9
REMARK   3   FREE R VALUE TEST SET COUNT      : 2459
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.200
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.257
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 3513
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.89
REMARK   3   BIN R VALUE           (WORKING SET) : 0.270
REMARK   3   BIN FREE R VALUE SET COUNT          : 169
REMARK   3   BIN FREE R VALUE                    : 0.327
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4297
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 196
REMARK   3   SOLVENT ATOMS            : 620
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 27.2
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 25.416
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 1.46
REMARK   3    B22 (A**2) : 1.46
REMARK   3    B33 (A**2) : -2.19
REMARK   3    B12 (A**2) : 0.73
REMARK   3    B13 (A**2) : 0.00
REMARK   3    B23 (A**2) : 0.00
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.172
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.168
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.118
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 4.787
REMARK   3
REMARK   3  CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.949
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.910
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4616 ; 0.014 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  6229 ; 1.506 ; 1.988
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   549 ; 6.287 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   215 ;34.240 ;24.047
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   732 ;14.745 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    25 ;13.880 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   647 ; 0.095 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  3488 ; 0.007 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2674 ; 0.719 ; 1.500
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  4331 ; 1.349 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1942 ; 2.110 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  1889 ; 3.390 ; 4.500
REMARK   3
REMARK   3  ANISOTROPIC THERMAL FACTOR RESTRAINTS.   COUNT   RMS    WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3   RIDING POSITIONS.
REMARK   4
REMARK   4 2WG0 COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 15-APR-09.
REMARK 100 THE PDBE ID CODE IS EBI-39475.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 22-FEB-07
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.8
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID14-2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.933
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 50212
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.20
REMARK 200  RESOLUTION RANGE LOW       (A) : 45.60
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.4
REMARK 200  DATA REDUNDANCY                : 5.6
REMARK 200  R MERGE                    (I) : 0.14
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.00
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.30
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.9
REMARK 200  R MERGE FOR SHELL          (I) : 0.49
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 3.25
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: OTHER
REMARK 200 SOFTWARE USED: REFMAC5
REMARK 200 STARTING MODEL: PDB ENTRY 1EA5
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 68
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.8
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: ACHE WAS CRYSTALLIZED IN 34%
REMARK 280  PEG200, 150 MM MES, PH 6, 4 DEG. C. THE CRYSTAL WAS SOAKED
REMARK 280  FOR 2 HOURS IN 1.3 MM SOMAN AT PH 6, 4 DEG C., THEN
REMARK 280  FLASH-COOLED IN LN2.
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 31 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+1/3
REMARK 290       3555   -X+Y,-X,Z+2/3
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z+2/3
REMARK 290       6555   -X,-X+Y,-Z+1/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       45.61333
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       91.22667
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       91.22667
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       45.61333
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY.  THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 GENERATING THE BIOMOLECULE
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PQS
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ASP A     1
REMARK 465     ASP A     2
REMARK 465     HIS A     3
REMARK 465     ALA A   536
REMARK 465     CYS A   537
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ARG A 388   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.4 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    LEU A  23       55.04     34.02
REMARK 500    SER A  24       -9.82     77.38
REMARK 500    SER A  25     -158.37   -112.42
REMARK 500    PHE A  45      -11.15     80.75
REMARK 500    CYS A  94       10.55   -142.95
REMARK 500    SER A 108       65.86   -152.62
REMARK 500    SER A 200     -125.23     67.09
REMARK 500    ILE A 296       93.66    -68.08
REMARK 500    GLU A 299      -72.16   -117.77
REMARK 500    ASP A 380       45.16   -158.28
REMARK 500    VAL A 400      -58.00   -128.35
REMARK 500    ASN A 457       43.64     72.48
REMARK 500    HIS A 486       81.56     50.63
REMARK 500    ASN A 506     -167.55   -165.68
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A2536
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A2557
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A2537
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A2538
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL A2539
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  GB A2540
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MES A2541
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PG4 A2542
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A2543
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A2544
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A2545
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A2546
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2547
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2548
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2549
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2550
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2551
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2552
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2553
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2554
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2555
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A2556
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 1ZGB   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF TORPEDO
REMARK 900  CALIFORNICAACETYLCHOLINESTERASE IN COMPLEX WITH
REMARK 900  AN (R)-TACRINE(10)-HUPYRIDONE INHIBITOR.
REMARK 900 RELATED ID: 1AMN   RELATED DB: PDB
REMARK 900  TRANSITION STATE ANALOG: ACETYLCHOLINESTERASE
REMARK 900  COMPLEXED WITH M-(N,N,N-TRIMETHYLAMMONIO)
REMARK 900  TRIFLUOROACETOPHENONE
REMARK 900 RELATED ID: 1QTI   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1E66   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE COMPLEXED WITH
REMARK 900   (-)-HUPRINE X AT 2.1A RESOLUTION
REMARK 900 RELATED ID: 2VQ6   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE COMPLEXED
REMARK 900   WITH 2-PAM
REMARK 900 RELATED ID: 2ACK   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH EDROPHONIUM,
REMARK 900  MONOCHROMATIC DATA
REMARK 900 RELATED ID: 2J3D   RELATED DB: PDB
REMARK 900  NATIVE MONOCLINIC FORM OF TORPEDO
REMARK 900  ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1QII   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT F) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 2CKM   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE COMPLEXED
REMARK 900   WITH ALKYLENE-LINKED BIS-TACRINE DIMER (7
REMARK 900   CARBON LINKER)
REMARK 900 RELATED ID: 1DX6   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE COMPLEXED WITH
REMARK 900   (-)-GALANTHAMINE AT 2.3A RESOLUTION
REMARK 900 RELATED ID: 1QIJ   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT G) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1QIE   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT B) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1ACL   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH DECAMETHONIUM
REMARK 900 RELATED ID: 1W4L   RELATED DB: PDB
REMARK 900  COMPLEX OF TCACHE WITH BIS-ACTING
REMARK 900  GALANTHAMINE DERIVATIVE
REMARK 900 RELATED ID: 1ODC   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE (E.C. 3.1
REMARK 900  .1.7) COMPLEXED WITH N-4'-QUINOLYL-N'-9
REMARK 900  "-(1",2",3",4"-TETRAHYDROACRIDINYL)-1,8-
REMARK 900  DIAMINOOCTANE AT 2.2A RESOLUTION
REMARK 900 RELATED ID: 2CMF   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE COMPLEXED
REMARK 900   WITH ALKYLENE-LINKED BIS-TACRINE DIMER (5
REMARK 900   CARBON LINKER)
REMARK 900 RELATED ID: 1GQS   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE (E.C. 3.1.1.7)
REMARK 900  COMPLEXED WITH NAP
REMARK 900 RELATED ID: 2J3Q   RELATED DB: PDB
REMARK 900  TORPEDO ACETYLCHOLINESTERASE COMPLEXED WITH
REMARK 900  FLUOROPHORE THIOFLAVIN T
REMARK 900 RELATED ID: 1E3Q   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE COMPLEXED
REMARK 900   WITH BW284C51
REMARK 900 RELATED ID: 2J4F   RELATED DB: PDB
REMARK 900  TORPEDO ACETYLCHOLINESTERASE - HG HEAVY-ATOM
REMARK 900  DERIVATIVE
REMARK 900 RELATED ID: 2DFP   RELATED DB: PDB
REMARK 900  X-RAY STRUCTURE OF AGED DI-ISOPROPYL-
REMARK 900  PHOSPHORO-FLUORIDATE (DFP) BOUND TO
REMARK 900  ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1QIK   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT H) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 2C5F   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH A NON HYDROLYSABLE SUBSTRATE
REMARK 900  ANALOGUE, 4-OXO-N,N,N-TRIMETHYLAMMONIUM
REMARK 900 RELATED ID: 1EA5   RELATED DB: PDB
REMARK 900  NATIVE ACETYLCHOLINESTERASE (E.C. 3.1.1.7
REMARK 900  ) FROM TORPEDO CALIFORNICA AT 1.8A
REMARK 900  RESOLUTION
REMARK 900 RELATED ID: 1EEA   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1QIF   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT C) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 2VJC   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH A NON HYDROLYSABLE SUBSTRATE
REMARK 900  ANALOGUE, 4-OXO-N,N,N-
REMARK 900  TRIMETHYLPENTANAMINIUM - ORTHORHOMBIC SPACE GROUP
REMARK 900   - DATASET A AT 150K
REMARK 900 RELATED ID: 2VJB   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH A NON HYDROLYSABLE SUBSTRATE
REMARK 900  ANALOGUE, 4-OXO-N,N,N-
REMARK 900  TRIMETHYLPENTANAMINIUM - ORTHORHOMBIC SPACE GROUP
REMARK 900   - DATASET D AT 100K
REMARK 900 RELATED ID: 1QIG   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT D) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1ZGC   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF TORPEDO
REMARK 900  CALIFORNICAACETYLCHOLINESTERASE IN COMPLEX WITH
REMARK 900  AN (RS)-TACRINE(10)-HUPYRIDONE INHIBITOR.
REMARK 900 RELATED ID: 1QID   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT A) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 2VJD   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH A NON HYDROLYSABLE SUBSTRATE
REMARK 900  ANALOGUE, 4-OXO-N,N,N-
REMARK 900  TRIMETHYLPENTANAMINIUM - ORTHORHOMBIC SPACE GROUP
REMARK 900   - DATASET C AT 150K
REMARK 900 RELATED ID: 1JJB   RELATED DB: PDB
REMARK 900  A NEUTRAL MOLECULE IN CATION-BINDING SITE:
REMARK 900  SPECIFIC BINDINGOF PEG-SH TO
REMARK 900  ACETYLCHOLINESTERASE FROM TORPEDO CALIFORNICA
REMARK 900 RELATED ID: 1UT6   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE (E.C. 3.1
REMARK 900  .1.7) COMPLEXED WITH N-9-(1',2',3',4
REMARK 900  '-TETRAHYDROACRIDINYL)-1,8- DIAMINOOCTANE AT
REMARK 900  2.4 ANGSTROMS RESOLUTION.
REMARK 900 RELATED ID: 2VT6   RELATED DB: PDB
REMARK 900  NATIVE TORPEDO CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900  COLLECTED WITH A CUMULATED DOSE OF 9400000
REMARK 900  GY
REMARK 900 RELATED ID: 2W9I   RELATED DB: PDB
REMARK 900  ACHE IN COMPLEX WITH METHYLENE BLUE
REMARK 900 RELATED ID: 2VT7   RELATED DB: PDB
REMARK 900  NATIVE TORPEDO CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900  COLLECTED WITH A CUMULATED DOSE OF 800000
REMARK 900  GY
REMARK 900 RELATED ID: 2CEK   RELATED DB: PDB
REMARK 900  CONFORMATIONAL FLEXIBILITY IN THE PERIPHERAL
REMARK 900  SITE OF TORPEDO CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900   REVEALED BY THE COMPLEX STRUCTURE WITH A
REMARK 900  BIFUNCTIONAL INHIBITOR
REMARK 900 RELATED ID: 1QIM   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT I) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1GPK   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE COMPLEXE WITH
REMARK 900   (+)-HUPERZINE A AT 2.1A RESOLUTION
REMARK 900 RELATED ID: 1JGA   RELATED DB: PDB
REMARK 900  THEORETICAL MODEL OF THE DIISOPROPYLPHOSPHORYL-
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH 1,7-
REMARK 900  HEPTYLENE-BIS-N,N'-SYN-2-PYRIDINIUMALDOXIME
REMARK 900 RELATED ID: 3ACE   RELATED DB: PDB
REMARK 900  THEORETICAL MODEL OF (R)-E2020 BOUND
REMARK 900  ACETYLCHOLINESTERASE COMPLEX, 3 STRUCTURES
REMARK 900 RELATED ID: 1OCE   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH MF268
REMARK 900 RELATED ID: 1W6R   RELATED DB: PDB
REMARK 900  COMPLEX OF TCACHE WITH GALANTHAMINE DERIVATIVE
REMARK 900 RELATED ID: 1SOM   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE INHIBITED
REMARK 900   BY NERVE AGENT GD (SOMAN).
REMARK 900 RELATED ID: 1VXO   RELATED DB: PDB
REMARK 900  METHYLPHOSPHONYLATED ACETYLCHOLINESTERASE (AGED)
REMARK 900  OBTAINED BY REACTION WITH O-ETHYL-S-[2-[
REMARK 900  BIS(1-METHYLETHYL) AMINO]ETHYL]
REMARK 900  METHYLPHOSPHONOTHIOATE (VX)
REMARK 900 RELATED ID: 2W6C   RELATED DB: PDB
REMARK 900  ACHE IN COMPLEX WITH A BIS-(-)-NOR-
REMARK 900  MEPTAZINOL DERIVATIVE
REMARK 900 RELATED ID: 2VJA   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH A NON HYDROLYSABLE SUBSTRATE
REMARK 900  ANALOGUE, 4-OXO-N,N,N-
REMARK 900  TRIMETHYLPENTANAMINIUM - ORTHORHOMBIC SPACE GROUP
REMARK 900   - DATASET A AT 100K
REMARK 900 RELATED ID: 1CFJ   RELATED DB: PDB
REMARK 900  METHYLPHOSPHONYLATED ACETYLCHOLINESTERASE (AGED)
REMARK 900  OBTAINED BY REACTION WITH O-
REMARK 900  ISOPROPYLMETHYLPHOSPHONOFLUORIDATE (GB, SARIN)
REMARK 900 RELATED ID: 2V96   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE UNPHOTOLYSED COMPLEX OF
REMARK 900  TCACHE WITH 1-(2-NITROPHENYL)-2,2,2-
REMARK 900  TRIFLUOROETHYL-ARSENOCHOLINE AT 100K
REMARK 900 RELATED ID: 1AX9   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH EDROPHONIUM,
REMARK 900  LAUE DATA
REMARK 900 RELATED ID: 1U65   RELATED DB: PDB
REMARK 900  ACHE W. CPT-11
REMARK 900 RELATED ID: 1W76   RELATED DB: PDB
REMARK 900  ORTHORHOMBIC FORM OF TORPEDO CALIFORNICA
REMARK 900  ACETYLCHOLINESTERASE (ACHE) COMPLEXED WITH BIS-
REMARK 900  ACTING GALANTHAMINE DERIVATIVE
REMARK 900 RELATED ID: 1H22   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE (E.C. 3.1
REMARK 900  .1.7) COMPLEXED WITH (S,S)-(-)-BIS(10)-
REMARK 900  HUPERZINE A-LIKE INHIBITOR AT 2.15A
REMARK 900  RESOLUTION
REMARK 900 RELATED ID: 1EVE   RELATED DB: PDB
REMARK 900  THREE DIMENSIONAL STRUCTURE OF THE ANTI-
REMARK 900  ALZHEIMER DRUG, E2020 (ARICEPT), COMPLEXED
REMARK 900  WITH ITS TARGET ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 2C4H   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH 500MM ACETYLTHIOCHOLINE
REMARK 900 RELATED ID: 2ACE   RELATED DB: PDB
REMARK 900  NATIVE ACETYLCHOLINESTERASE FROM TORPEDO
REMARK 900  CALIFORNICA
REMARK 900 RELATED ID: 2VA9   RELATED DB: PDB
REMARK 900  STRUCTURE OF NATIVE TCACHE AFTER A 9
REMARK 900  SECONDS ANNEALING TO ROOM TEMPERATURE DURING
REMARK 900  THE FIRST 5 SECONDS OF WHICH LASER
REMARK 900  IRRADIATION AT 266NM TOOK PLACE
REMARK 900 RELATED ID: 1GQR   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE (E.C. 3.1.1.7)
REMARK 900  COMPLEXED WITH RIVASTIGMINE
REMARK 900 RELATED ID: 1VXR   RELATED DB: PDB
REMARK 900  O-ETHYLMETHYLPHOSPHONYLATED ACETYLCHOLINESTERASE
REMARK 900  OBTAINED BY REACTION WITH O-ETHYL-S-[2-[
REMARK 900  BIS(1-METHYLETHYL) AMINO]ETHYL]
REMARK 900  METHYLPHOSPHONOTHIOATE (VX)
REMARK 900 RELATED ID: 4ACE   RELATED DB: PDB
REMARK 900  THEORETICAL MODEL OF (S)-E2020 BOUND
REMARK 900  ACETYLCHOLINESTERASE COMPLEX, 3 STRUCTURES
REMARK 900 RELATED ID: 2C58   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH 20MM ACETYLTHIOCHOLINE
REMARK 900 RELATED ID: 1HBJ   RELATED DB: PDB
REMARK 900  X-RAY CRYSTAL STRUCTURE OF COMPLEX BETWEEN
REMARK 900  TORPEDO CALIFORNICA ACHE AND A REVERSIBLE
REMARK 900  INHIBITOR, 4-AMINO-5-FLUORO-2-METHYL-3-(
REMARK 900  3-TRIFLUOROACETYLBENZYLTHIOMETHYL)QUINOLINE
REMARK 900 RELATED ID: 1W75   RELATED DB: PDB
REMARK 900  NATIVE ORTHORHOMBIC FORM OF TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE (ACHE)
REMARK 900 RELATED ID: 1VOT   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH HUPERZINE A
REMARK 900 RELATED ID: 2C5G   RELATED DB: PDB
REMARK 900  TORPEDO CALIFORNICA ACETYLCHOLINESTERASE IN
REMARK 900  COMPLEX WITH 20MM THIOCHOLINE
REMARK 900 RELATED ID: 1JGB   RELATED DB: PDB
REMARK 900  THEORETICAL MODEL OF THE DIISOPROPYLPHOSPHORYL-
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH 1,3-
REMARK 900  PROPYLENE-BIS-N,N'-SYN-4-PYRIDINIUMALDOXIME
REMARK 900 RELATED ID: 2V98   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE COMPLEX OF TCACHE WITH 1
REMARK 900  -(2-NITROPHENYL)-2,2,2-TRIFLUOROETHYL-
REMARK 900  ARSENOCHOLINE AFTER A 9 SECONDS ANNEALING TO
REMARK 900   ROOM TEMPERATURE, DURING HTE FIRST 5
REMARK 900  SECONDS OF WHICH LASER IRRADIATION AT 266NM
REMARK 900   TOOK PLACE
REMARK 900 RELATED ID: 1GPN   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE COMPLEXED WITH
REMARK 900   HUPERZINE B AT 2.35A RESOLUTION
REMARK 900 RELATED ID: 1QIH   RELATED DB: PDB
REMARK 900  SPECIFIC CHEMICAL AND STRUCTURAL DAMAGE AT
REMARK 900  NINE TIME POINTS (POINT E) CAUSED BY
REMARK 900  INTENSE SYNCHROTRON RADIATION TO TORPEDO
REMARK 900  CALIFORNICA ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1H23   RELATED DB: PDB
REMARK 900  STRUCTURE OF ACETYLCHOLINESTERASE (E.C. 3.1
REMARK 900  .1.7) COMPLEXED WITH (S,S)-(-)-BIS(12)-
REMARK 900  HUPERZINE A-LIKE INHIBITOR AT 2.15A
REMARK 900  RESOLUTION
REMARK 900 RELATED ID: 1ACJ   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH TACRINE
REMARK 900 RELATED ID: 1FSS   RELATED DB: PDB
REMARK 900  ACETYLCHOLINESTERASE COMPLEXED WITH FASCICULIN-
REMARK 900  II
REMARK 900 RELATED ID: 2V97   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE UNPHOTOLYSED COMPLEX OF
REMARK 900  TCACHE WITH 1-(2-NITROPHENYL)-2,2,2-
REMARK 900  TRIFLUOROETHYL-ARSENOCHOLINE AFTER A 9 SECONDS
REMARK 900   ANNEALING TO ROOM TEMPERATURE
REMARK 900 RELATED ID: 2WFZ   RELATED DB: PDB
REMARK 900  NON-AGED CONJUGATE OF TORPEDO CALIFORNICA
REMARK 900  ACETYLCHOLINESTERASE WITH SOMAN
REMARK 900 RELATED ID: 2WG1   RELATED DB: PDB
REMARK 900  TERNARY COMPLEX OF THE AGED CONJUGATE OF
REMARK 900  TORPEDO CALIFORNICA ACEYLCHOLINESTERASE WITH
REMARK 900  SOMAN AND 2-PAM
REMARK 900 RELATED ID: 2WG2   RELATED DB: PDB
REMARK 900  NON-AGED CONJUGATE OF TORPEDO CALIFORNICA
REMARK 900  ACETYLCHOLINESTERASE WITH SOMAN (ALTERNATIVE
REMARK 900  REFINEMENT)
DBREF  2WG0 A    1   537  UNP    P04058   ACES_TORCA      22    558
SEQRES   1 A  537  ASP ASP HIS SER GLU LEU LEU VAL ASN THR LYS SER GLY
SEQRES   2 A  537  LYS VAL MET GLY THR ARG VAL PRO VAL LEU SER SER HIS
SEQRES   3 A  537  ILE SER ALA PHE LEU GLY ILE PRO PHE ALA GLU PRO PRO
SEQRES   4 A  537  VAL GLY ASN MET ARG PHE ARG ARG PRO GLU PRO LYS LYS
SEQRES   5 A  537  PRO TRP SER GLY VAL TRP ASN ALA SER THR TYR PRO ASN
SEQRES   6 A  537  ASN CYS GLN GLN TYR VAL ASP GLU GLN PHE PRO GLY PHE
SEQRES   7 A  537  SER GLY SER GLU MET TRP ASN PRO ASN ARG GLU MET SER
SEQRES   8 A  537  GLU ASP CYS LEU TYR LEU ASN ILE TRP VAL PRO SER PRO
SEQRES   9 A  537  ARG PRO LYS SER THR THR VAL MET VAL TRP ILE TYR GLY
SEQRES  10 A  537  GLY GLY PHE TYR SER GLY SER SER THR LEU ASP VAL TYR
SEQRES  11 A  537  ASN GLY LYS TYR LEU ALA TYR THR GLU GLU VAL VAL LEU
SEQRES  12 A  537  VAL SER LEU SER TYR ARG VAL GLY ALA PHE GLY PHE LEU
SEQRES  13 A  537  ALA LEU HIS GLY SER GLN GLU ALA PRO GLY ASN VAL GLY
SEQRES  14 A  537  LEU LEU ASP GLN ARG MET ALA LEU GLN TRP VAL HIS ASP
SEQRES  15 A  537  ASN ILE GLN PHE PHE GLY GLY ASP PRO LYS THR VAL THR
SEQRES  16 A  537  ILE PHE GLY GLU SER ALA GLY GLY ALA SER VAL GLY MET
SEQRES  17 A  537  HIS ILE LEU SER PRO GLY SER ARG ASP LEU PHE ARG ARG
SEQRES  18 A  537  ALA ILE LEU GLN SER GLY SER PRO ASN CYS PRO TRP ALA
SEQRES  19 A  537  SER VAL SER VAL ALA GLU GLY ARG ARG ARG ALA VAL GLU
SEQRES  20 A  537  LEU GLY ARG ASN LEU ASN CYS ASN LEU ASN SER ASP GLU
SEQRES  21 A  537  GLU LEU ILE HIS CYS LEU ARG GLU LYS LYS PRO GLN GLU
SEQRES  22 A  537  LEU ILE ASP VAL GLU TRP ASN VAL LEU PRO PHE ASP SER
SEQRES  23 A  537  ILE PHE ARG PHE SER PHE VAL PRO VAL ILE ASP GLY GLU
SEQRES  24 A  537  PHE PHE PRO THR SER LEU GLU SER MET LEU ASN SER GLY
SEQRES  25 A  537  ASN PHE LYS LYS THR GLN ILE LEU LEU GLY VAL ASN LYS
SEQRES  26 A  537  ASP GLU GLY SER PHE PHE LEU LEU TYR GLY ALA PRO GLY
SEQRES  27 A  537  PHE SER LYS ASP SER GLU SER LYS ILE SER ARG GLU ASP
SEQRES  28 A  537  PHE MET SER GLY VAL LYS LEU SER VAL PRO HIS ALA ASN
SEQRES  29 A  537  ASP LEU GLY LEU ASP ALA VAL THR LEU GLN TYR THR ASP
SEQRES  30 A  537  TRP MET ASP ASP ASN ASN GLY ILE LYS ASN ARG ASP GLY
SEQRES  31 A  537  LEU ASP ASP ILE VAL GLY ASP HIS ASN VAL ILE CYS PRO
SEQRES  32 A  537  LEU MET HIS PHE VAL ASN LYS TYR THR LYS PHE GLY ASN
SEQRES  33 A  537  GLY THR TYR LEU TYR PHE PHE ASN HIS ARG ALA SER ASN
SEQRES  34 A  537  LEU VAL TRP PRO GLU TRP MET GLY VAL ILE HIS GLY TYR
SEQRES  35 A  537  GLU ILE GLU PHE VAL PHE GLY LEU PRO LEU VAL LYS GLU
SEQRES  36 A  537  LEU ASN TYR THR ALA GLU GLU GLU ALA LEU SER ARG ARG
SEQRES  37 A  537  ILE MET HIS TYR TRP ALA THR PHE ALA LYS THR GLY ASN
SEQRES  38 A  537  PRO ASN GLU PRO HIS SER GLN GLU SER LYS TRP PRO LEU
SEQRES  39 A  537  PHE THR THR LYS GLU GLN LYS PHE ILE ASP LEU ASN THR
SEQRES  40 A  537  GLU PRO MET LYS VAL HIS GLN ARG LEU ARG VAL GLN MET
SEQRES  41 A  537  CYS VAL PHE TRP ASN GLN PHE LEU PRO LYS LEU LEU ASN
SEQRES  42 A  537  ALA THR ALA CYS
HET    NAG  A2536      14
HET    NAG  A2557      14
HET    NAG  A2537      14
HET    NAG  A2538      14
HET     CL  A2539       1
HET     GB  A2540       4
HET    MES  A2541      12
HET    PG4  A2542      13
HET    PGE  A2543      10
HET    PGE  A2544      10
HET    PGE  A2545      10
HET    PGE  A2546      10
HET    PEG  A2547       7
HET    PEG  A2548       7
HET    PEG  A2549       7
HET    PEG  A2550       7
HET    PEG  A2551       7
HET    PEG  A2552       7
HET    PEG  A2553       7
HET    PEG  A2554       7
HET    PEG  A2555       7
HET    PEG  A2556       7
HETNAM     PG4 TETRAETHYLENE GLYCOL
HETNAM     PGE TRIETHYLENE GLYCOL
HETNAM      CL CHLORIDE ION
HETNAM     PEG DI(HYDROXYETHYL)ETHER
HETNAM      GB METHYLPHOSPHONIC ACID ESTER GROUP
HETNAM     MES 2-(N-MORPHOLINO)-ETHANESULFONIC ACID
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   2  PG4    C8 H18 O5
FORMUL   3  PGE    4(C6 H14 O4)
FORMUL   4   CL    CL 1-
FORMUL   5  PEG    10(C4 H10 O3)
FORMUL   6   GB    C H5 O3 P
FORMUL   7  MES    C6 H13 N O4 S
FORMUL   8  NAG    4(C8 H15 N O6)
FORMUL   9  HOH   *620(H2 O1)
HELIX    1   1 VAL A   40  ARG A   44  5                                   5
HELIX    2   2 PHE A   78  MET A   83  1                                   6
HELIX    3   3 LEU A  127  ASN A  131  5                                   5
HELIX    4   4 GLY A  132  GLU A  140  1                                   9
HELIX    5   5 VAL A  150  LEU A  156  1                                   7
HELIX    6   6 ASN A  167  ILE A  184  1                                  18
HELIX    7   7 GLN A  185  PHE A  187  5                                   3
HELIX    8   8 SER A  200  SER A  212  1                                  13
HELIX    9   9 SER A  215  PHE A  219  5                                   5
HELIX   10  10 SER A  237  LEU A  252  1                                  16
HELIX   11  11 SER A  258  LYS A  269  1                                  12
HELIX   12  12 LYS A  270  GLU A  278  1                                   9
HELIX   13  13 TRP A  279  LEU A  282  5                                   4
HELIX   14  14 SER A  304  GLY A  312  1                                   9
HELIX   15  15 GLY A  328  ALA A  336  1                                   9
HELIX   16  16 SER A  348  VAL A  360  1                                  13
HELIX   17  17 ASN A  364  TYR A  375  1                                  12
HELIX   18  18 ASN A  383  VAL A  400  1                                  18
HELIX   19  19 VAL A  400  LYS A  413  1                                  14
HELIX   20  20 PRO A  433  GLY A  437  5                                   5
HELIX   21  21 GLU A  443  PHE A  448  1                                   6
HELIX   22  22 GLY A  449  VAL A  453  5                                   5
HELIX   23  23 VAL A  453  ASN A  457  5                                   5
HELIX   24  24 THR A  459  GLY A  480  1                                  22
HELIX   25  25 ARG A  517  GLN A  526  1                                  10
HELIX   26  26 GLN A  526  THR A  535  1                                  10
SHEET    1  AA 3 LEU A   7  THR A  10  0
SHEET    2  AA 3 GLY A  13  MET A  16 -1  O  GLY A  13   N  THR A  10
SHEET    3  AA 3 VAL A  57  ASN A  59  1  O  TRP A  58   N  MET A  16
SHEET    1  AB11 THR A  18  VAL A  22  0
SHEET    2  AB11 SER A  25  PRO A  34 -1  O  SER A  25   N  VAL A  22
SHEET    3  AB11 TYR A  96  VAL A 101 -1  O  LEU A  97   N  ILE A  33
SHEET    4  AB11 VAL A 142  SER A 145 -1  O  LEU A 143   N  TRP A 100
SHEET    5  AB11 THR A 109  ILE A 115  1  O  THR A 110   N  VAL A 142
SHEET    6  AB11 GLY A 189  GLU A 199  1  N  ASP A 190   O  THR A 109
SHEET    7  AB11 ARG A 221  GLN A 225  1  O  ARG A 221   N  ILE A 196
SHEET    8  AB11 GLN A 318  ASN A 324  1  O  GLN A 318   N  ALA A 222
SHEET    9  AB11 GLY A 417  PHE A 423  1  O  GLY A 417   N  ILE A 319
SHEET   10  AB11 LYS A 501  LEU A 505  1  O  ILE A 503   N  PHE A 422
SHEET   11  AB11 VAL A 512  GLN A 514 -1  O  HIS A 513   N  PHE A 502
SSBOND   1 CYS A   67    CYS A   94                          1555   1555  2.05
SSBOND   2 CYS A  254    CYS A  265                          1555   1555  2.05
SSBOND   3 CYS A  402    CYS A  521                          1555   1555  2.02
LINK         ND2 ASN A  59                 C1  NAG A2538     1555   1555  1.45
LINK         OG  SER A 200                 P1   GB A2540     1555   1555  1.62
LINK         ND2 ASN A 416                 C1  NAG A2537     1555   1555  1.45
LINK         ND2 ASN A 457                 C1  NAG A2536     1555   1555  1.45
LINK         O4  NAG A2537                 C1  NAG A2557     1555   1555  1.44
CISPEP   1 SER A  103    PRO A  104          0         2.17
SITE     1 AC1  3 ASN A 457  HOH A1586  HOH A1587
SITE     1 AC2  2 HOH A1620  NAG A2537
SITE     1 AC3  7 ASN A 416  HOH A1588  HOH A1590  HOH A1591
SITE     2 AC3  7 HOH A1592  HOH A1593  NAG A2557
SITE     1 AC4  6 ASN A  59  SER A  61  HOH A1594  HOH A1595
SITE     2 AC4  6 HOH A1596  HOH A1597
SITE     1 AC5  3 LYS A 133  TYR A 134  HOH A1494
SITE     1 AC6  8 GLY A 118  GLY A 119  SER A 200  ALA A 201
SITE     2 AC6  8 PHE A 288  PHE A 290  HIS A 440  PEG A2549
SITE     1 AC7 10 ASN A   9  THR A  10  LYS A  11  ARG A 243
SITE     2 AC7 10 GLU A 247  ASN A 280  VAL A 281  LEU A 282
SITE     3 AC7 10 PRO A 283  HOH A1600
SITE     1 AC8  5 TYR A 121  PHE A 330  TYR A 334  HOH A1602
SITE     2 AC8  5 PEG A2549
SITE     1 AC9  5 ILE A 296  ASP A 297  HOH A1114  HOH A1603
SITE     2 AC9  5 HOH A1605
SITE     1 BC1  8 SER A   4  GLU A   5  ARG A 105  ARG A 250
SITE     2 BC1  8 ASN A 251  ASN A 253  HOH A1149  PEG A2550
SITE     1 BC2  4 GLU A 499  LYS A 501  VAL A 512  GLN A 514
SITE     1 BC3  4 LYS A  11  TRP A  54  PHE A 284  HOH A1610
SITE     1 BC4  4 THR A 138  ARG A 467  ARG A 468  HOH A1189
SITE     1 BC5  7 ARG A 242  VAL A 246  LEU A 256  ASN A 257
SITE     2 BC5  7 SER A 258  ASP A 259  HOH A1611
SITE     1 BC6  9 TRP A  84  GLY A 117  GLY A 118  GLU A 199
SITE     2 BC6  9 PHE A 330  HOH A1612  HOH A1613   GB A2540
SITE     3 BC6  9 PG4 A2542
SITE     1 BC7 10 VAL A   8  ASN A   9  ARG A 105  PHE A 186
SITE     2 BC7 10 ASN A 251  ASN A 280  HOH A1149  HOH A1150
SITE     3 BC7 10 HOH A1151  PGE A2544
SITE     1 BC8 10 MET A 405  HIS A 406  ASN A 409  ARG A 515
SITE     2 BC8 10 LEU A 516  VAL A 518  ASN A 525  GLN A 526
SITE     3 BC8 10 HOH A1546  HOH A1575
SITE     1 BC9  8 VAL A  20  TYR A  63  ARG A  88  TRP A 100
SITE     2 BC9  8 ASN A 131  LYS A 133  HOH A1074  HOH A1614
SITE     1 CC1  1 THR A 376
SITE     1 CC2  3 SER A  55  VAL A  57  TRP A  58
SITE     1 CC3  9 SER A  79  GLY A  80  MET A  83  ASN A 429
SITE     2 CC3  9 LEU A 430  VAL A 431  TYR A 442  HOH A1489
SITE     3 CC3  9 HOH A1490
SITE     1 CC4  7 ASP A 326  ASP A 389  GLU A 434  TRP A 435
SITE     2 CC4  7 HOH A1617  HOH A1618  HOH A1619
CRYST1  111.610  111.610  136.840  90.00  90.00 120.00 P 31 2 1      6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008960  0.005173  0.000000        0.00000
SCALE2      0.000000  0.010346  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007308        0.00000
TER    4298      THR A 535
MASTER      698    0   22   26   14    0   41    6 5113    1  205   42
END