longtext: 3HI4-pdb

content
HEADER    HYDROLASE                               18-MAY-09   3HI4
TITLE     SWITCHING CATALYSIS FROM HYDROLYSIS TO PERHYDROLYSIS IN P.
TITLE    2 FLUORESCENS ESTERASE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ARYLESTERASE;
COMPND   3 CHAIN: A, B, C, D, E, F;
COMPND   4 SYNONYM: ARYL-ESTER HYDROLASE, PFE, PUTATIVE
COMPND   5 BROMOPEROXIDASE;
COMPND   6 EC: 3.1.1.2, 1.-.-.-;
COMPND   7 ENGINEERED: YES;
COMPND   8 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOMONAS FLUORESCENS;
SOURCE   3 ORGANISM_TAXID: 294;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   6 EXPRESSION_SYSTEM_STRAIN: DH5ALPHA;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   8 EXPRESSION_SYSTEM_PLASMID: PJOE2792
KEYWDS    ESTERASE, HYDROLYSIS, PERHYDROLYSIS, HYDROLASE, PEROXIDASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    V.M.PURPERO
REVDAT   1   23-FEB-10 3HI4    0
JRNL        AUTH   D.L.YIN,P.BERNHARDT,K.L.MORLEY,Y.JIANG,
JRNL        AUTH 2 J.D.CHEESEMAN,V.PURPERO,J.D.SCHRAG,R.J.KAZLAUSKAS
JRNL        TITL   SWITCHING CATALYSIS FROM HYDROLYSIS TO
JRNL        TITL 2 PERHYDROLYSIS IN PSEUDOMONAS FLUORESCENS ESTERASE.
JRNL        REF    BIOCHEMISTRY                               2010
JRNL        REFN                   ISSN 0006-2960
JRNL        PMID   20112920
JRNL        DOI    10.1021/BI9021268
REMARK   2
REMARK   2 RESOLUTION.    2.25 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 6.1.1
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.25
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 39.97
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.6
REMARK   3   NUMBER OF REFLECTIONS             : 146673
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.166
REMARK   3   R VALUE            (WORKING SET) : 0.165
REMARK   3   FREE R VALUE                     : 0.202
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 7330
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 12802
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 185
REMARK   3   SOLVENT ATOMS            : 1036
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 37.65
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 29.39
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.00000
REMARK   3    B22 (A**2) : 0.00000
REMARK   3    B33 (A**2) : 0.00000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.155
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.147
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.097
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 3.939
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   DISTANCE RESTRAINTS.                    RMS    SIGMA
REMARK   3    BOND LENGTH                     (A) : NULL  ; NULL
REMARK   3    ANGLE DISTANCE                  (A) : NULL  ; NULL
REMARK   3    INTRAPLANAR 1-4 DISTANCE        (A) : NULL  ; NULL
REMARK   3    H-BOND OR METAL COORDINATION    (A) : NULL  ; NULL
REMARK   3
REMARK   3   PLANE RESTRAINT                  (A) : NULL  ; NULL
REMARK   3   CHIRAL-CENTER RESTRAINT       (A**3) : NULL  ; NULL
REMARK   3
REMARK   3   NON-BONDED CONTACT RESTRAINTS.
REMARK   3    SINGLE TORSION                  (A) : NULL  ; NULL
REMARK   3    MULTIPLE TORSION                (A) : NULL  ; NULL
REMARK   3    H-BOND (X...Y)                  (A) : NULL  ; NULL
REMARK   3    H-BOND (X-H...Y)                (A) : NULL  ; NULL
REMARK   3
REMARK   3   CONFORMATIONAL TORSION ANGLE RESTRAINTS.
REMARK   3    SPECIFIED                 (DEGREES) : NULL  ; NULL
REMARK   3    PLANAR                    (DEGREES) : NULL  ; NULL
REMARK   3    STAGGERED                 (DEGREES) : NULL  ; NULL
REMARK   3    TRANSVERSE                (DEGREES) : NULL  ; NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3  RIDING POSITIONS
REMARK   4
REMARK   4 3HI4 COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 28-MAY-09.
REMARK 100 THE RCSB ID CODE IS RCSB053161.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 08-DEC-08
REMARK 200  TEMPERATURE           (KELVIN) : 120
REMARK 200  PH                             : 7.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : ROTATING ANODE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU MICROMAX-007 HF
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.54
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : AREA DETECTOR
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU RAXIS IV++
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 146673
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.250
REMARK 200  RESOLUTION RANGE LOW       (A) : 39.970
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.6
REMARK 200  DATA REDUNDANCY                : 2.700
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : 0.07900
REMARK 200   FOR THE DATA SET  : 12.3000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.25
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.29
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.6
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.50
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : 0.35600
REMARK 200   FOR SHELL         : 2.150
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: 1VA4
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 72.15
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 4.42
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 1.75M AMMONIUM SULFATE 1% PEG 400,
REMARK 280  0.1M HEPES PH 7.0, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE
REMARK 280  291K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 32
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+2/3
REMARK 290       3555   -X+Y,-X,Z+1/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       86.66000
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       43.33000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TRIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 8560 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 27800 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -38.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: TRIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 8990 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 27440 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -31.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D, E, F
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OD2  ASP C   138     O    HOH C   380              2.03
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    SER A  44   CB    SER A  44   OG     -0.081
REMARK 500    SER C  44   CB    SER C  44   OG     -0.093
REMARK 500    SER D  44   CB    SER D  44   OG     -0.115
REMARK 500    GLU D  87   CD    GLU D  87   OE1     0.069
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ASP A  15   CB  -  CG  -  OD1 ANGL. DEV. =   5.8 DEGREES
REMARK 500    ASP A  15   CB  -  CG  -  OD2 ANGL. DEV. =  -6.5 DEGREES
REMARK 500    ARG A  59   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.2 DEGREES
REMARK 500    ASP A  74   CB  -  CG  -  OD1 ANGL. DEV. =   5.7 DEGREES
REMARK 500    LYS A 189   CD  -  CE  -  NZ  ANGL. DEV. = -14.5 DEGREES
REMARK 500    ASP B 138   CB  -  CG  -  OD1 ANGL. DEV. =  -5.7 DEGREES
REMARK 500    ASP B 220   CB  -  CG  -  OD1 ANGL. DEV. =   5.7 DEGREES
REMARK 500    ASP B 220   CB  -  CG  -  OD2 ANGL. DEV. =  -7.0 DEGREES
REMARK 500    ARG C 102   NE  -  CZ  -  NH2 ANGL. DEV. =   3.1 DEGREES
REMARK 500    ARG D  59   NE  -  CZ  -  NH1 ANGL. DEV. =   3.0 DEGREES
REMARK 500    ARG E 151   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.1 DEGREES
REMARK 500    ASP E 202   CB  -  CG  -  OD1 ANGL. DEV. =   5.8 DEGREES
REMARK 500    ARG F 111   NE  -  CZ  -  NH1 ANGL. DEV. =  -3.7 DEGREES
REMARK 500    ARG F 111   NE  -  CZ  -  NH2 ANGL. DEV. =   4.9 DEGREES
REMARK 500    ASP F 211   CB  -  CG  -  OD1 ANGL. DEV. =   6.1 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PRO A  29       65.37   -105.41
REMARK 500    LEU A  30     -147.19   -102.69
REMARK 500    ASP A  31     -168.88   -161.94
REMARK 500    GLN A  62       76.09   -118.57
REMARK 500    SER A  94     -117.20     57.20
REMARK 500    PHE A 125      -63.69   -101.11
REMARK 500    ASP A 150       87.11   -158.72
REMARK 500    THR A 230      -91.47   -127.39
REMARK 500    PRO B  29       72.83   -102.03
REMARK 500    LEU B  30     -148.06   -109.27
REMARK 500    GLN B  62       68.82   -116.67
REMARK 500    SER B  94     -126.00     63.32
REMARK 500    THR B 230      -92.80   -126.88
REMARK 500    PRO C  29       65.23   -110.77
REMARK 500    LEU C  30     -145.40    -98.92
REMARK 500    GLN C  62       68.80   -117.90
REMARK 500    SER C  94     -122.24     55.35
REMARK 500    TYR C 131       57.56   -146.28
REMARK 500    ASP C 150       89.35   -153.69
REMARK 500    THR C 230      -91.45   -129.70
REMARK 500    ASP C 248        2.52     56.09
REMARK 500    PRO D  29       65.45   -107.09
REMARK 500    LEU D  30     -146.38   -100.50
REMARK 500    ASP D  31     -168.91   -161.21
REMARK 500    SER D  94     -122.50     63.72
REMARK 500    PHE D 125      -63.64   -121.42
REMARK 500    TYR D 131       59.85   -140.09
REMARK 500    ASP D 150       84.67   -150.73
REMARK 500    THR D 230      -86.53   -125.49
REMARK 500    ASP D 248        8.63     58.61
REMARK 500    PRO E  29       64.62   -108.00
REMARK 500    LEU E  30     -147.19   -103.06
REMARK 500    GLN E  62       77.84   -119.98
REMARK 500    SER E  94     -125.20     56.67
REMARK 500    PHE E 125      -61.86   -102.17
REMARK 500    TYR E 131       54.15   -142.28
REMARK 500    ASP E 150       89.01   -153.47
REMARK 500    THR E 230      -89.39   -129.63
REMARK 500    ASP E 248        9.18     57.03
REMARK 500    PRO F  29       64.39   -106.06
REMARK 500    LEU F  30     -147.24   -101.91
REMARK 500    GLN F  62       71.69   -112.29
REMARK 500    SER F  94     -126.98     65.39
REMARK 500    TYR F 131       56.32   -153.08
REMARK 500    ASP F 150       88.03   -160.98
REMARK 500    THR F 230      -93.38   -129.97
REMARK 500    ASP F 248       -0.70     59.50
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    VAL A 255        20.5      L          L   OUTSIDE RANGE
REMARK 500    PHE B 198        24.0      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH D1000        DISTANCE =  5.07 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT A 272
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A 273
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 274
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 275
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 276
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT B 272
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 B 273
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL B 274
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL B 275
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL B 276
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT C 272
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 C 273
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT C 274
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL C 275
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL C 276
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL C 277
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL C 278
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT D 272
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 D 273
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL D 274
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL D 275
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL D 276
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT E 272
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 E 273
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT E 274
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL E 275
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL E 276
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL E 277
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT F 272
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT F 273
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE ACT F 274
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL F 275
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL F 276
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL F 277
REMARK 800
REMARK 800 SITE_IDENTIFIER: DC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL F 278
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 1VA4   RELATED DB: PDB
REMARK 900 PFE WITH MUTATION AT L29P
DBREF  3HI4 A    1   271  UNP    P22862   ESTE_PSEFL       2    272
DBREF  3HI4 B    1   271  UNP    P22862   ESTE_PSEFL       2    272
DBREF  3HI4 C    1   271  UNP    P22862   ESTE_PSEFL       2    272
DBREF  3HI4 D    1   271  UNP    P22862   ESTE_PSEFL       2    272
DBREF  3HI4 E    1   271  UNP    P22862   ESTE_PSEFL       2    272
DBREF  3HI4 F    1   271  UNP    P22862   ESTE_PSEFL       2    272
SEQADV 3HI4 PRO A   29  UNP  P22862    LEU    30 ENGINEERED
SEQADV 3HI4 PRO B   29  UNP  P22862    LEU    30 ENGINEERED
SEQADV 3HI4 PRO C   29  UNP  P22862    LEU    30 ENGINEERED
SEQADV 3HI4 PRO D   29  UNP  P22862    LEU    30 ENGINEERED
SEQADV 3HI4 PRO E   29  UNP  P22862    LEU    30 ENGINEERED
SEQADV 3HI4 PRO F   29  UNP  P22862    LEU    30 ENGINEERED
SEQRES   1 A  271  SER THR PHE VAL ALA LYS ASP GLY THR GLN ILE TYR PHE
SEQRES   2 A  271  LYS ASP TRP GLY SER GLY LYS PRO VAL LEU PHE SER HIS
SEQRES   3 A  271  GLY TRP PRO LEU ASP ALA ASP MET TRP GLU TYR GLN MET
SEQRES   4 A  271  GLU TYR LEU SER SER ARG GLY TYR ARG THR ILE ALA PHE
SEQRES   5 A  271  ASP ARG ARG GLY PHE GLY ARG SER ASP GLN PRO TRP THR
SEQRES   6 A  271  GLY ASN ASP TYR ASP THR PHE ALA ASP ASP ILE ALA GLN
SEQRES   7 A  271  LEU ILE GLU HIS LEU ASP LEU LYS GLU VAL THR LEU VAL
SEQRES   8 A  271  GLY PHE SER MET GLY GLY GLY ASP VAL ALA ARG TYR ILE
SEQRES   9 A  271  ALA ARG HIS GLY SER ALA ARG VAL ALA GLY LEU VAL LEU
SEQRES  10 A  271  LEU GLY ALA VAL THR PRO LEU PHE GLY GLN LYS PRO ASP
SEQRES  11 A  271  TYR PRO GLN GLY VAL PRO LEU ASP VAL PHE ALA ARG PHE
SEQRES  12 A  271  LYS THR GLU LEU LEU LYS ASP ARG ALA GLN PHE ILE SER
SEQRES  13 A  271  ASP PHE ASN ALA PRO PHE TYR GLY ILE ASN LYS GLY GLN
SEQRES  14 A  271  VAL VAL SER GLN GLY VAL GLN THR GLN THR LEU GLN ILE
SEQRES  15 A  271  ALA LEU LEU ALA SER LEU LYS ALA THR VAL ASP CYS VAL
SEQRES  16 A  271  THR ALA PHE ALA GLU THR ASP PHE ARG PRO ASP MET ALA
SEQRES  17 A  271  LYS ILE ASP VAL PRO THR LEU VAL ILE HIS GLY ASP GLY
SEQRES  18 A  271  ASP GLN ILE VAL PRO PHE GLU THR THR GLY LYS VAL ALA
SEQRES  19 A  271  ALA GLU LEU ILE LYS GLY ALA GLU LEU LYS VAL TYR LYS
SEQRES  20 A  271  ASP ALA PRO HIS GLY PHE ALA VAL THR HIS ALA GLN GLN
SEQRES  21 A  271  LEU ASN GLU ASP LEU LEU ALA PHE LEU LYS ARG
SEQRES   1 B  271  SER THR PHE VAL ALA LYS ASP GLY THR GLN ILE TYR PHE
SEQRES   2 B  271  LYS ASP TRP GLY SER GLY LYS PRO VAL LEU PHE SER HIS
SEQRES   3 B  271  GLY TRP PRO LEU ASP ALA ASP MET TRP GLU TYR GLN MET
SEQRES   4 B  271  GLU TYR LEU SER SER ARG GLY TYR ARG THR ILE ALA PHE
SEQRES   5 B  271  ASP ARG ARG GLY PHE GLY ARG SER ASP GLN PRO TRP THR
SEQRES   6 B  271  GLY ASN ASP TYR ASP THR PHE ALA ASP ASP ILE ALA GLN
SEQRES   7 B  271  LEU ILE GLU HIS LEU ASP LEU LYS GLU VAL THR LEU VAL
SEQRES   8 B  271  GLY PHE SER MET GLY GLY GLY ASP VAL ALA ARG TYR ILE
SEQRES   9 B  271  ALA ARG HIS GLY SER ALA ARG VAL ALA GLY LEU VAL LEU
SEQRES  10 B  271  LEU GLY ALA VAL THR PRO LEU PHE GLY GLN LYS PRO ASP
SEQRES  11 B  271  TYR PRO GLN GLY VAL PRO LEU ASP VAL PHE ALA ARG PHE
SEQRES  12 B  271  LYS THR GLU LEU LEU LYS ASP ARG ALA GLN PHE ILE SER
SEQRES  13 B  271  ASP PHE ASN ALA PRO PHE TYR GLY ILE ASN LYS GLY GLN
SEQRES  14 B  271  VAL VAL SER GLN GLY VAL GLN THR GLN THR LEU GLN ILE
SEQRES  15 B  271  ALA LEU LEU ALA SER LEU LYS ALA THR VAL ASP CYS VAL
SEQRES  16 B  271  THR ALA PHE ALA GLU THR ASP PHE ARG PRO ASP MET ALA
SEQRES  17 B  271  LYS ILE ASP VAL PRO THR LEU VAL ILE HIS GLY ASP GLY
SEQRES  18 B  271  ASP GLN ILE VAL PRO PHE GLU THR THR GLY LYS VAL ALA
SEQRES  19 B  271  ALA GLU LEU ILE LYS GLY ALA GLU LEU LYS VAL TYR LYS
SEQRES  20 B  271  ASP ALA PRO HIS GLY PHE ALA VAL THR HIS ALA GLN GLN
SEQRES  21 B  271  LEU ASN GLU ASP LEU LEU ALA PHE LEU LYS ARG
SEQRES   1 C  271  SER THR PHE VAL ALA LYS ASP GLY THR GLN ILE TYR PHE
SEQRES   2 C  271  LYS ASP TRP GLY SER GLY LYS PRO VAL LEU PHE SER HIS
SEQRES   3 C  271  GLY TRP PRO LEU ASP ALA ASP MET TRP GLU TYR GLN MET
SEQRES   4 C  271  GLU TYR LEU SER SER ARG GLY TYR ARG THR ILE ALA PHE
SEQRES   5 C  271  ASP ARG ARG GLY PHE GLY ARG SER ASP GLN PRO TRP THR
SEQRES   6 C  271  GLY ASN ASP TYR ASP THR PHE ALA ASP ASP ILE ALA GLN
SEQRES   7 C  271  LEU ILE GLU HIS LEU ASP LEU LYS GLU VAL THR LEU VAL
SEQRES   8 C  271  GLY PHE SER MET GLY GLY GLY ASP VAL ALA ARG TYR ILE
SEQRES   9 C  271  ALA ARG HIS GLY SER ALA ARG VAL ALA GLY LEU VAL LEU
SEQRES  10 C  271  LEU GLY ALA VAL THR PRO LEU PHE GLY GLN LYS PRO ASP
SEQRES  11 C  271  TYR PRO GLN GLY VAL PRO LEU ASP VAL PHE ALA ARG PHE
SEQRES  12 C  271  LYS THR GLU LEU LEU LYS ASP ARG ALA GLN PHE ILE SER
SEQRES  13 C  271  ASP PHE ASN ALA PRO PHE TYR GLY ILE ASN LYS GLY GLN
SEQRES  14 C  271  VAL VAL SER GLN GLY VAL GLN THR GLN THR LEU GLN ILE
SEQRES  15 C  271  ALA LEU LEU ALA SER LEU LYS ALA THR VAL ASP CYS VAL
SEQRES  16 C  271  THR ALA PHE ALA GLU THR ASP PHE ARG PRO ASP MET ALA
SEQRES  17 C  271  LYS ILE ASP VAL PRO THR LEU VAL ILE HIS GLY ASP GLY
SEQRES  18 C  271  ASP GLN ILE VAL PRO PHE GLU THR THR GLY LYS VAL ALA
SEQRES  19 C  271  ALA GLU LEU ILE LYS GLY ALA GLU LEU LYS VAL TYR LYS
SEQRES  20 C  271  ASP ALA PRO HIS GLY PHE ALA VAL THR HIS ALA GLN GLN
SEQRES  21 C  271  LEU ASN GLU ASP LEU LEU ALA PHE LEU LYS ARG
SEQRES   1 D  271  SER THR PHE VAL ALA LYS ASP GLY THR GLN ILE TYR PHE
SEQRES   2 D  271  LYS ASP TRP GLY SER GLY LYS PRO VAL LEU PHE SER HIS
SEQRES   3 D  271  GLY TRP PRO LEU ASP ALA ASP MET TRP GLU TYR GLN MET
SEQRES   4 D  271  GLU TYR LEU SER SER ARG GLY TYR ARG THR ILE ALA PHE
SEQRES   5 D  271  ASP ARG ARG GLY PHE GLY ARG SER ASP GLN PRO TRP THR
SEQRES   6 D  271  GLY ASN ASP TYR ASP THR PHE ALA ASP ASP ILE ALA GLN
SEQRES   7 D  271  LEU ILE GLU HIS LEU ASP LEU LYS GLU VAL THR LEU VAL
SEQRES   8 D  271  GLY PHE SER MET GLY GLY GLY ASP VAL ALA ARG TYR ILE
SEQRES   9 D  271  ALA ARG HIS GLY SER ALA ARG VAL ALA GLY LEU VAL LEU
SEQRES  10 D  271  LEU GLY ALA VAL THR PRO LEU PHE GLY GLN LYS PRO ASP
SEQRES  11 D  271  TYR PRO GLN GLY VAL PRO LEU ASP VAL PHE ALA ARG PHE
SEQRES  12 D  271  LYS THR GLU LEU LEU LYS ASP ARG ALA GLN PHE ILE SER
SEQRES  13 D  271  ASP PHE ASN ALA PRO PHE TYR GLY ILE ASN LYS GLY GLN
SEQRES  14 D  271  VAL VAL SER GLN GLY VAL GLN THR GLN THR LEU GLN ILE
SEQRES  15 D  271  ALA LEU LEU ALA SER LEU LYS ALA THR VAL ASP CYS VAL
SEQRES  16 D  271  THR ALA PHE ALA GLU THR ASP PHE ARG PRO ASP MET ALA
SEQRES  17 D  271  LYS ILE ASP VAL PRO THR LEU VAL ILE HIS GLY ASP GLY
SEQRES  18 D  271  ASP GLN ILE VAL PRO PHE GLU THR THR GLY LYS VAL ALA
SEQRES  19 D  271  ALA GLU LEU ILE LYS GLY ALA GLU LEU LYS VAL TYR LYS
SEQRES  20 D  271  ASP ALA PRO HIS GLY PHE ALA VAL THR HIS ALA GLN GLN
SEQRES  21 D  271  LEU ASN GLU ASP LEU LEU ALA PHE LEU LYS ARG
SEQRES   1 E  271  SER THR PHE VAL ALA LYS ASP GLY THR GLN ILE TYR PHE
SEQRES   2 E  271  LYS ASP TRP GLY SER GLY LYS PRO VAL LEU PHE SER HIS
SEQRES   3 E  271  GLY TRP PRO LEU ASP ALA ASP MET TRP GLU TYR GLN MET
SEQRES   4 E  271  GLU TYR LEU SER SER ARG GLY TYR ARG THR ILE ALA PHE
SEQRES   5 E  271  ASP ARG ARG GLY PHE GLY ARG SER ASP GLN PRO TRP THR
SEQRES   6 E  271  GLY ASN ASP TYR ASP THR PHE ALA ASP ASP ILE ALA GLN
SEQRES   7 E  271  LEU ILE GLU HIS LEU ASP LEU LYS GLU VAL THR LEU VAL
SEQRES   8 E  271  GLY PHE SER MET GLY GLY GLY ASP VAL ALA ARG TYR ILE
SEQRES   9 E  271  ALA ARG HIS GLY SER ALA ARG VAL ALA GLY LEU VAL LEU
SEQRES  10 E  271  LEU GLY ALA VAL THR PRO LEU PHE GLY GLN LYS PRO ASP
SEQRES  11 E  271  TYR PRO GLN GLY VAL PRO LEU ASP VAL PHE ALA ARG PHE
SEQRES  12 E  271  LYS THR GLU LEU LEU LYS ASP ARG ALA GLN PHE ILE SER
SEQRES  13 E  271  ASP PHE ASN ALA PRO PHE TYR GLY ILE ASN LYS GLY GLN
SEQRES  14 E  271  VAL VAL SER GLN GLY VAL GLN THR GLN THR LEU GLN ILE
SEQRES  15 E  271  ALA LEU LEU ALA SER LEU LYS ALA THR VAL ASP CYS VAL
SEQRES  16 E  271  THR ALA PHE ALA GLU THR ASP PHE ARG PRO ASP MET ALA
SEQRES  17 E  271  LYS ILE ASP VAL PRO THR LEU VAL ILE HIS GLY ASP GLY
SEQRES  18 E  271  ASP GLN ILE VAL PRO PHE GLU THR THR GLY LYS VAL ALA
SEQRES  19 E  271  ALA GLU LEU ILE LYS GLY ALA GLU LEU LYS VAL TYR LYS
SEQRES  20 E  271  ASP ALA PRO HIS GLY PHE ALA VAL THR HIS ALA GLN GLN
SEQRES  21 E  271  LEU ASN GLU ASP LEU LEU ALA PHE LEU LYS ARG
SEQRES   1 F  271  SER THR PHE VAL ALA LYS ASP GLY THR GLN ILE TYR PHE
SEQRES   2 F  271  LYS ASP TRP GLY SER GLY LYS PRO VAL LEU PHE SER HIS
SEQRES   3 F  271  GLY TRP PRO LEU ASP ALA ASP MET TRP GLU TYR GLN MET
SEQRES   4 F  271  GLU TYR LEU SER SER ARG GLY TYR ARG THR ILE ALA PHE
SEQRES   5 F  271  ASP ARG ARG GLY PHE GLY ARG SER ASP GLN PRO TRP THR
SEQRES   6 F  271  GLY ASN ASP TYR ASP THR PHE ALA ASP ASP ILE ALA GLN
SEQRES   7 F  271  LEU ILE GLU HIS LEU ASP LEU LYS GLU VAL THR LEU VAL
SEQRES   8 F  271  GLY PHE SER MET GLY GLY GLY ASP VAL ALA ARG TYR ILE
SEQRES   9 F  271  ALA ARG HIS GLY SER ALA ARG VAL ALA GLY LEU VAL LEU
SEQRES  10 F  271  LEU GLY ALA VAL THR PRO LEU PHE GLY GLN LYS PRO ASP
SEQRES  11 F  271  TYR PRO GLN GLY VAL PRO LEU ASP VAL PHE ALA ARG PHE
SEQRES  12 F  271  LYS THR GLU LEU LEU LYS ASP ARG ALA GLN PHE ILE SER
SEQRES  13 F  271  ASP PHE ASN ALA PRO PHE TYR GLY ILE ASN LYS GLY GLN
SEQRES  14 F  271  VAL VAL SER GLN GLY VAL GLN THR GLN THR LEU GLN ILE
SEQRES  15 F  271  ALA LEU LEU ALA SER LEU LYS ALA THR VAL ASP CYS VAL
SEQRES  16 F  271  THR ALA PHE ALA GLU THR ASP PHE ARG PRO ASP MET ALA
SEQRES  17 F  271  LYS ILE ASP VAL PRO THR LEU VAL ILE HIS GLY ASP GLY
SEQRES  18 F  271  ASP GLN ILE VAL PRO PHE GLU THR THR GLY LYS VAL ALA
SEQRES  19 F  271  ALA GLU LEU ILE LYS GLY ALA GLU LEU LYS VAL TYR LYS
SEQRES  20 F  271  ASP ALA PRO HIS GLY PHE ALA VAL THR HIS ALA GLN GLN
SEQRES  21 F  271  LEU ASN GLU ASP LEU LEU ALA PHE LEU LYS ARG
HET    ACT  A 272       4
HET    SO4  A 273       5
HET    GOL  A 274       6
HET    GOL  A 275       6
HET    GOL  A 276       6
HET    ACT  B 272       4
HET    SO4  B 273       5
HET    GOL  B 274       6
HET    GOL  B 275       6
HET    GOL  B 276       6
HET    ACT  C 272       4
HET    SO4  C 273       5
HET    ACT  C 274       4
HET    GOL  C 275       6
HET    GOL  C 276       6
HET    GOL  C 277       6
HET    GOL  C 278       6
HET    ACT  D 272       4
HET    SO4  D 273       5
HET    GOL  D 274       6
HET    GOL  D 275       6
HET    GOL  D 276       6
HET    ACT  E 272       4
HET    SO4  E 273       5
HET    ACT  E 274       4
HET    GOL  E 275       6
HET    GOL  E 276       6
HET    GOL  E 277       6
HET    ACT  F 272       4
HET    ACT  F 273       4
HET    ACT  F 274       4
HET    GOL  F 275       6
HET    GOL  F 276       6
HET    GOL  F 277       6
HET    GOL  F 278       6
HETNAM     ACT ACETATE ION
HETNAM     SO4 SULFATE ION
HETNAM     GOL GLYCEROL
FORMUL   7  ACT    10(C2 H3 O2 1-)
FORMUL   8  SO4    5(O4 S 2-)
FORMUL   9  GOL    20(C3 H8 O3)
FORMUL  42  HOH   *1036(H2 O)
HELIX    1   1 ASP A   31  MET A   34  5                                   4
HELIX    2   2 TRP A   35  SER A   44  1                                  10
HELIX    3   3 ASP A   68  ASP A   84  1                                  17
HELIX    4   4 MET A   95  GLY A  108  1                                  14
HELIX    5   5 PRO A  136  GLY A  164  1                                  29
HELIX    6   6 ILE A  165  GLY A  168  5                                   4
HELIX    7   7 SER A  172  ALA A  186  1                                  15
HELIX    8   8 SER A  187  THR A  201  1                                  15
HELIX    9   9 PHE A  203  ILE A  210  1                                   8
HELIX   10  10 PRO A  226  THR A  229  5                                   4
HELIX   11  11 THR A  230  ILE A  238  1                                   9
HELIX   12  12 GLY A  252  HIS A  257  1                                   6
HELIX   13  13 HIS A  257  ARG A  271  1                                  15
HELIX   14  14 ASP B   31  MET B   34  5                                   4
HELIX   15  15 TRP B   35  SER B   44  1                                  10
HELIX   16  16 ASP B   68  ASP B   84  1                                  17
HELIX   17  17 MET B   95  GLY B  108  1                                  14
HELIX   18  18 PRO B  136  GLY B  164  1                                  29
HELIX   19  19 ILE B  165  GLY B  168  5                                   4
HELIX   20  20 SER B  172  ALA B  186  1                                  15
HELIX   21  21 SER B  187  THR B  201  1                                  15
HELIX   22  22 PHE B  203  ILE B  210  1                                   8
HELIX   23  23 PRO B  226  THR B  229  5                                   4
HELIX   24  24 THR B  230  ILE B  238  1                                   9
HELIX   25  25 GLY B  252  HIS B  257  1                                   6
HELIX   26  26 HIS B  257  ARG B  271  1                                  15
HELIX   27  27 ASP C   31  MET C   34  5                                   4
HELIX   28  28 TRP C   35  SER C   44  1                                  10
HELIX   29  29 ASP C   68  ASP C   84  1                                  17
HELIX   30  30 MET C   95  GLY C  108  1                                  14
HELIX   31  31 PRO C  136  GLY C  164  1                                  29
HELIX   32  32 ILE C  165  GLY C  168  5                                   4
HELIX   33  33 SER C  172  ALA C  186  1                                  15
HELIX   34  34 SER C  187  THR C  201  1                                  15
HELIX   35  35 PHE C  203  ILE C  210  1                                   8
HELIX   36  36 PRO C  226  THR C  229  5                                   4
HELIX   37  37 THR C  230  ILE C  238  1                                   9
HELIX   38  38 GLY C  252  HIS C  257  1                                   6
HELIX   39  39 HIS C  257  LYS C  270  1                                  14
HELIX   40  40 ASP D   31  MET D   34  5                                   4
HELIX   41  41 TRP D   35  SER D   44  1                                  10
HELIX   42  42 ASP D   68  LEU D   83  1                                  16
HELIX   43  43 MET D   95  GLY D  108  1                                  14
HELIX   44  44 PRO D  136  TYR D  163  1                                  28
HELIX   45  45 GLY D  164  GLY D  168  5                                   5
HELIX   46  46 SER D  172  LEU D  185  1                                  14
HELIX   47  47 SER D  187  THR D  201  1                                  15
HELIX   48  48 PHE D  203  ILE D  210  1                                   8
HELIX   49  49 PRO D  226  THR D  229  5                                   4
HELIX   50  50 THR D  230  ILE D  238  1                                   9
HELIX   51  51 GLY D  252  HIS D  257  1                                   6
HELIX   52  52 HIS D  257  ARG D  271  1                                  15
HELIX   53  53 ASP E   31  MET E   34  5                                   4
HELIX   54  54 TRP E   35  SER E   44  1                                  10
HELIX   55  55 ASP E   68  LEU E   83  1                                  16
HELIX   56  56 MET E   95  GLY E  108  1                                  14
HELIX   57  57 PRO E  136  GLY E  164  1                                  29
HELIX   58  58 ILE E  165  GLY E  168  5                                   4
HELIX   59  59 SER E  172  ALA E  186  1                                  15
HELIX   60  60 SER E  187  THR E  201  1                                  15
HELIX   61  61 PHE E  203  LYS E  209  1                                   7
HELIX   62  62 PRO E  226  THR E  229  5                                   4
HELIX   63  63 THR E  230  ILE E  238  1                                   9
HELIX   64  64 GLY E  252  HIS E  257  1                                   6
HELIX   65  65 HIS E  257  ARG E  271  1                                  15
HELIX   66  66 ASP F   31  MET F   34  5                                   4
HELIX   67  67 TRP F   35  SER F   44  1                                  10
HELIX   68  68 ASP F   68  ASP F   84  1                                  17
HELIX   69  69 MET F   95  GLY F  108  1                                  14
HELIX   70  70 PRO F  136  TYR F  163  1                                  28
HELIX   71  71 GLY F  164  GLY F  168  5                                   5
HELIX   72  72 SER F  172  ALA F  186  1                                  15
HELIX   73  73 SER F  187  THR F  201  1                                  15
HELIX   74  74 PHE F  203  ALA F  208  1                                   6
HELIX   75  75 PRO F  226  THR F  229  5                                   4
HELIX   76  76 THR F  230  ILE F  238  1                                   9
HELIX   77  77 GLY F  252  HIS F  257  1                                   6
HELIX   78  78 HIS F  257  LYS F  270  1                                  14
SHEET    1   A 8 THR A   2  VAL A   4  0
SHEET    2   A 8 GLN A  10  TRP A  16 -1  O  ILE A  11   N  PHE A   3
SHEET    3   A 8 TYR A  47  PHE A  52 -1  O  THR A  49   N  TRP A  16
SHEET    4   A 8 LYS A  20  SER A  25  1  N  PHE A  24   O  ILE A  50
SHEET    5   A 8 VAL A  88  PHE A  93  1  O  THR A  89   N  LEU A  23
SHEET    6   A 8 VAL A 112  LEU A 118  1  O  LEU A 118   N  GLY A  92
SHEET    7   A 8 THR A 214  GLY A 219  1  O  ILE A 217   N  LEU A 117
SHEET    8   A 8 GLU A 242  TYR A 246  1  O  GLU A 242   N  VAL A 216
SHEET    1   B 8 THR B   2  VAL B   4  0
SHEET    2   B 8 GLN B  10  TRP B  16 -1  O  ILE B  11   N  PHE B   3
SHEET    3   B 8 ARG B  48  PHE B  52 -1  O  THR B  49   N  TRP B  16
SHEET    4   B 8 PRO B  21  SER B  25  1  N  PHE B  24   O  ILE B  50
SHEET    5   B 8 VAL B  88  PHE B  93  1  O  VAL B  91   N  LEU B  23
SHEET    6   B 8 VAL B 112  LEU B 118  1  O  VAL B 116   N  LEU B  90
SHEET    7   B 8 THR B 214  GLY B 219  1  O  ILE B 217   N  LEU B 117
SHEET    8   B 8 GLU B 242  TYR B 246  1  O  LYS B 244   N  VAL B 216
SHEET    1   C 8 THR C   2  VAL C   4  0
SHEET    2   C 8 GLN C  10  TRP C  16 -1  O  ILE C  11   N  PHE C   3
SHEET    3   C 8 TYR C  47  PHE C  52 -1  O  THR C  49   N  TRP C  16
SHEET    4   C 8 LYS C  20  SER C  25  1  N  PHE C  24   O  ILE C  50
SHEET    5   C 8 VAL C  88  PHE C  93  1  O  VAL C  91   N  SER C  25
SHEET    6   C 8 VAL C 112  LEU C 118  1  O  LEU C 118   N  GLY C  92
SHEET    7   C 8 THR C 214  GLY C 219  1  O  ILE C 217   N  LEU C 117
SHEET    8   C 8 GLU C 242  TYR C 246  1  O  GLU C 242   N  VAL C 216
SHEET    1   D 8 THR D   2  VAL D   4  0
SHEET    2   D 8 GLN D  10  TRP D  16 -1  O  ILE D  11   N  PHE D   3
SHEET    3   D 8 TYR D  47  PHE D  52 -1  O  THR D  49   N  TRP D  16
SHEET    4   D 8 LYS D  20  SER D  25  1  N  PHE D  24   O  ILE D  50
SHEET    5   D 8 VAL D  88  PHE D  93  1  O  VAL D  91   N  SER D  25
SHEET    6   D 8 VAL D 112  LEU D 118  1  O  LEU D 118   N  GLY D  92
SHEET    7   D 8 THR D 214  GLY D 219  1  O  ILE D 217   N  LEU D 117
SHEET    8   D 8 GLU D 242  TYR D 246  1  O  GLU D 242   N  VAL D 216
SHEET    1   E 8 THR E   2  VAL E   4  0
SHEET    2   E 8 GLN E  10  TRP E  16 -1  O  ILE E  11   N  PHE E   3
SHEET    3   E 8 ARG E  48  PHE E  52 -1  O  THR E  49   N  TRP E  16
SHEET    4   E 8 PRO E  21  SER E  25  1  N  PHE E  24   O  ILE E  50
SHEET    5   E 8 VAL E  88  PHE E  93  1  O  VAL E  91   N  SER E  25
SHEET    6   E 8 VAL E 112  LEU E 118  1  O  LEU E 118   N  GLY E  92
SHEET    7   E 8 THR E 214  GLY E 219  1  O  LEU E 215   N  LEU E 117
SHEET    8   E 8 GLU E 242  TYR E 246  1  O  LYS E 244   N  VAL E 216
SHEET    1   F 8 THR F   2  VAL F   4  0
SHEET    2   F 8 GLN F  10  GLY F  17 -1  O  ILE F  11   N  PHE F   3
SHEET    3   F 8 TYR F  47  PHE F  52 -1  O  THR F  49   N  TRP F  16
SHEET    4   F 8 LYS F  20  SER F  25  1  N  PHE F  24   O  ILE F  50
SHEET    5   F 8 VAL F  88  PHE F  93  1  O  VAL F  91   N  SER F  25
SHEET    6   F 8 VAL F 112  LEU F 118  1  O  LEU F 118   N  GLY F  92
SHEET    7   F 8 THR F 214  GLY F 219  1  O  LEU F 215   N  LEU F 117
SHEET    8   F 8 GLU F 242  TYR F 246  1  O  GLU F 242   N  THR F 214
CISPEP   1 TRP A   28    PRO A   29          0        -9.42
CISPEP   2 THR A  122    PRO A  123          0         6.96
CISPEP   3 TRP B   28    PRO B   29          0       -17.73
CISPEP   4 THR B  122    PRO B  123          0         3.03
CISPEP   5 TRP C   28    PRO C   29          0       -10.93
CISPEP   6 THR C  122    PRO C  123          0         6.27
CISPEP   7 TRP D   28    PRO D   29          0        -7.41
CISPEP   8 THR D  122    PRO D  123          0         8.80
CISPEP   9 TRP E   28    PRO E   29          0        -8.39
CISPEP  10 THR E  122    PRO E  123          0         4.91
CISPEP  11 TRP F   28    PRO F   29          0        -8.42
CISPEP  12 THR F  122    PRO F  123          0         8.68
SITE     1 AC1  5 TRP A  28  SER A  94  MET A  95  VAL A 121
SITE     2 AC1  5 HOH A 278
SITE     1 AC2  1 ARG A 106
SITE     1 AC3  8 SER A   1  TYR A  12  PHE A  13  LYS A  14
SITE     2 AC3  8 ASP A  15  ASP C  61  GLN C  62  HOH C 659
SITE     1 AC4  5 LEU A 137  ALA A 141  LYS A 144  GLU A 200
SITE     2 AC4  5 HOH A 289
SITE     1 AC5  6 LEU A  30  TYR A 163  VAL A 175  GLN A 178
SITE     2 AC5  6 THR A 179  HOH A 311
SITE     1 AC6  6 TRP B  28  SER B  94  MET B  95  VAL B 121
SITE     2 AC6  6 PHE B 198  HOH B 277
SITE     1 AC7  2 ARG B 106  HOH B 602
SITE     1 AC8  6 LEU B 137  LYS B 144  THR B 196  GLU B 200
SITE     2 AC8  6 HOH B 535  HOH B 548
SITE     1 AC9  6 ASP A  61  GLN A  62  SER B   1  TYR B  12
SITE     2 AC9  6 LYS B  14  ASP B  15
SITE     1 BC1  5 TYR B 163  GLN B 178  THR B 179  ILE B 182
SITE     2 BC1  5 HOH B 291
SITE     1 BC2  5 TRP C  28  SER C  94  MET C  95  VAL C 121
SITE     2 BC2  5 HOH C 279
SITE     1 BC3  1 ARG C 106
SITE     1 BC4  1 ILE C 224
SITE     1 BC5  8 ASP B  61  GLN B  62  SER C   1  TYR C  12
SITE     2 BC5  8 PHE C  13  LYS C  14  ASP C  15  HOH C 913
SITE     1 BC6  4 LEU C 137  LYS C 144  GLU C 200  HOH C 752
SITE     1 BC7  5 LEU C  30  TYR C 163  GLN C 178  THR C 179
SITE     2 BC7  5 ILE C 182
SITE     1 BC8  2 ASN A 166  GLY C  19
SITE     1 BC9  6 TRP D  28  SER D  94  MET D  95  VAL D 121
SITE     2 BC9  6 PHE D 198  HOH D 277
SITE     1 CC1  1 ARG D 106
SITE     1 CC2  5 LEU D 137  ALA D 141  LYS D 144  THR D 196
SITE     2 CC2  5 GLU D 200
SITE     1 CC3  6 SER D   1  TYR D  12  LYS D  14  ASP D  15
SITE     2 CC3  6 ASP E  61  GLN E  62
SITE     1 CC4  7 TYR D 163  VAL D 175  GLN D 178  THR D 179
SITE     2 CC4  7 ILE D 182  HOH D 293  HOH D 419
SITE     1 CC5  5 TRP E  28  SER E  94  MET E  95  VAL E 121
SITE     2 CC5  5 HOH E 278
SITE     1 CC6  1 ARG E 106
SITE     1 CC7  1 ILE E 224
SITE     1 CC8  5 SER E   1  TYR E  12  ASP E  15  ASP F  61
SITE     2 CC8  5 GLN F  62
SITE     1 CC9  6 LEU E 137  ALA E 141  LYS E 144  THR E 196
SITE     2 CC9  6 GLU E 200  HOH E 284
SITE     1 DC1  6 TYR E 163  GLN E 178  THR E 179  ILE E 182
SITE     2 DC1  6 HOH E 332  HOH E 481
SITE     1 DC2  6 TRP F  28  SER F  94  MET F  95  VAL F 121
SITE     2 DC2  6 PHE F 198  HOH F 279
SITE     1 DC3  4 VAL F 245  LYS F 247  GLN F 260  HOH F 590
SITE     1 DC4  2 GLN F 223  ILE F 224
SITE     1 DC5  8 ASP D  61  GLN D  62  SER F   1  TYR F  12
SITE     2 DC5  8 PHE F  13  LYS F  14  ASP F  15  HOH F 294
SITE     1 DC6  5 LEU F 137  ALA F 141  LYS F 144  THR F 196
SITE     2 DC6  5 GLU F 200
SITE     1 DC7  5 SER F 156  ASN F 159  GLN F 176  HOH F 418
SITE     2 DC7  5 HOH F1034
SITE     1 DC8  5 TYR F 163  GLN F 178  ILE F 182  HOH F 330
SITE     2 DC8  5 HOH F 982
CRYST1  145.490  145.490  129.990  90.00  90.00 120.00 P 32         18
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.006873  0.003968  0.000000        0.00000
SCALE2      0.000000  0.007937  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007693        0.00000
TER    2133      ARG A 271
TER    4266      ARG B 271
TER    6404      ARG C 271
TER    8537      ARG D 271
TER   10675      ARG E 271
TER   12808      ARG F 271
MASTER      506    0   35   78   48    0   59    614023    6  185  126
END