longtext: 3ICV-pdb

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HEADER    HYDROLASE                               18-JUL-09   3ICV
TITLE     STRUCTURAL CONSEQUENCES OF A CIRCULAR PERMUTATION ON LIPASE
TITLE    2 B FROM CANDIDA ANTARTICA
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE B;
COMPND   3 CHAIN: A;
COMPND   4 SYNONYM: CALB;
COMPND   5 EC: 3.1.1.3;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: CANDIDA ANTARCTICA;
SOURCE   3 ORGANISM_COMMON: YEAST;
SOURCE   4 ORGANISM_TAXID: 34362;
SOURCE   5 EXPRESSION_SYSTEM: PICHIA PASTORIS;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 4922
KEYWDS    CIRCULAR PERMUTATION, CLEAVAGE ON PAIR OF BASIC RESIDUES,
KEYWDS   2 GLYCOPROTEIN, HYDROLASE, LIPID DEGRADATION, ZYMOGEN,
KEYWDS   3 DISULFIDE BOND
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.R.HORTON,Z.QIAN,D.JIA,S.LUTZ,X.CHENG
REVDAT   1   06-OCT-09 3ICV    0
JRNL        AUTH   Z.QIAN,J.R.HORTON,X.CHENG,S.LUTZ
JRNL        TITL   STRUCTURAL REDESIGN OF LIPASE B FROM CANDIDA
JRNL        TITL 2 ANTARCTICA BY CIRCULAR PERMUTATION AND INCREMENTAL
JRNL        TITL 3 TRUNCATION.
JRNL        REF    J.MOL.BIOL.                                2009
JRNL        REFN                   ESSN 1089-8638
JRNL        PMID   19683009
JRNL        DOI    10.1016/J.JMB.2009.08.008
REMARK   1
REMARK   2
REMARK   2 RESOLUTION.    1.49 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.1
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.49
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 27.81
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 97.4
REMARK   3   NUMBER OF REFLECTIONS             : 62943
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.177
REMARK   3   FREE R VALUE                     : 0.193
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : 3197
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.49
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.54
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 92.20
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2200
REMARK   3   BIN FREE R VALUE                    : 0.2250
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 280
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.013
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2202
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 44
REMARK   3   SOLVENT ATOMS            : 413
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 20.50
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 24.80
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -2.88000
REMARK   3    B22 (A**2) : -2.88000
REMARK   3    B33 (A**2) : 5.76000
REMARK   3    B12 (A**2) : -2.14000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.15
REMARK   3   ESD FROM SIGMAA              (A) : 0.09
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 27.81
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.17
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.09
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.008
REMARK   3   BOND ANGLES            (DEGREES) : 1.60
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 24.10
REMARK   3   IMPROPER ANGLES        (DEGREES) : 1.08
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : ISOTROPIC
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : NULL
REMARK   3   KSOL        : NULL
REMARK   3   BSOL        : NULL
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3ICV COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 22-JUL-09.
REMARK 100 THE RCSB ID CODE IS RCSB054252.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 20-OCT-07
REMARK 200  TEMPERATURE           (KELVIN) : 173.0
REMARK 200  PH                             : 5.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 22-ID
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0000
REMARK 200  MONOCHROMATOR                  : SI 111
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARMOSAIC 300 MM CCD
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 64040
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.490
REMARK 200  RESOLUTION RANGE LOW       (A) : 27.810
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.1
REMARK 200  DATA REDUNDANCY                : 10.500
REMARK 200  R MERGE                    (I) : 0.07700
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 23.3000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.49
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.54
REMARK 200  COMPLETENESS FOR SHELL     (%) : 98.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 9.30
REMARK 200  R MERGE FOR SHELL          (I) : 0.34500
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 7.200
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: GLRF
REMARK 200 STARTING MODEL: PDB ENTRY 1LBT
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 59.13
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.01
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 25% PEG 3350, 0.1M BIS-TRIS, 0.2M
REMARK 280  SODIUM CHLORIDE, PH 5.5, VAPOR DIFFUSION, HANGING DROP,
REMARK 280  TEMPERATURE 289K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 32 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+2/3
REMARK 290       3555   -X+Y,-X,Z+1/3
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z+1/3
REMARK 290       6555   -X,-X+Y,-Z+2/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       36.66467
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       18.33233
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       18.33233
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000       36.66467
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 5210 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 22070 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -23.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -0.500000 -0.866025  0.000000      167.52000
REMARK 350   BIOMT2   2 -0.866025  0.500000  0.000000       96.71772
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000       91.66167
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 762  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 773  LIES ON A SPECIAL POSITION.
REMARK 375      HOH A 786  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ALA A     1
REMARK 465     ALA A     2
REMARK 465     ILE A     3
REMARK 465     VAL A     4
REMARK 465     ALA A     5
REMARK 465     GLY A   176
REMARK 465     PRO A   177
REMARK 465     LEU A   178
REMARK 465     ASP A   179
REMARK 465     ALA A   180
REMARK 465     LEU A   181
REMARK 465     LEU A   300
REMARK 465     THR A   301
REMARK 465     PRO A   302
REMARK 465     GLU A   303
REMARK 465     GLN A   304
REMARK 465     LYS A   305
REMARK 465     VAL A   306
REMARK 465     ALA A   307
REMARK 465     ALA A   308
REMARK 465     ALA A   309
REMARK 465     ALA A   310
REMARK 465     LEU A   311
REMARK 465     LEU A   312
REMARK 465     ALA A   313
REMARK 465     PRO A   314
REMARK 465     ALA A   315
REMARK 465     ALA A   316
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A  63       83.18   -154.71
REMARK 500    ASN A  85      -92.96   -139.12
REMARK 500    ASP A 109      118.54    -35.48
REMARK 500    SER A 139     -131.61     56.53
REMARK 500    ASP A 168       67.13   -108.81
REMARK 500    GLN A 227       10.95   -140.06
REMARK 500    ASN A 240       -1.54     73.43
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 559        DISTANCE =  6.58 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 401
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 402
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE BTB A 403
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3ICW   RELATED DB: PDB
REMARK 900 STRUCTURE OF A CIRCULAR PERMUTATION ON LIPASE B FROM
REMARK 900 CANDIDA ANTARTICA WITH BOUND SUICIDE INHIBITOR
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 THE RECOMBINANT PROTEIN IS A "CIRCULAR PERMUTATION" OF LIPASE B
REMARK 999 IN WHICH THE C-TERMINAL SEQUENCE
REMARK 999 AAIVAGPKQNCEPDLMPYARPFAVGKRTCSGIVTP HAS BEEN MOVED TO THE N-
REMARK 999 TERMINUS WITH THE TP REPLACED WITH G. ALA82 AND THR114 REPRESENT
REMARK 999 VARIATIONS FROM THE PUBLISHED WILD TYPE SEQUENCE AND THEIR
REMARK 999 ACCURACY WAS CONFIRMED INDEPENDENTLY BY DNA SEQUENCING
DBREF  3ICV A    1    33  UNP    P41365   LIPB_CANAR     308    340
DBREF  3ICV A   35   316  UNP    P41365   LIPB_CANAR      26    307
SEQADV 3ICV ALA A   91  UNP  P41365    THR    82 SEE REMARK 999
SEQADV 3ICV THR A  123  UNP  P41365    ALA   114 SEE REMARK 999
SEQRES   1 A  316  ALA ALA ILE VAL ALA GLY PRO LYS GLN ASN CYS GLU PRO
SEQRES   2 A  316  ASP LEU MET PRO TYR ALA ARG PRO PHE ALA VAL GLY LYS
SEQRES   3 A  316  ARG THR CYS SER GLY ILE VAL GLY LEU PRO SER GLY SER
SEQRES   4 A  316  ASP PRO ALA PHE SER GLN PRO LYS SER VAL LEU ASP ALA
SEQRES   5 A  316  GLY LEU THR CYS GLN GLY ALA SER PRO SER SER VAL SER
SEQRES   6 A  316  LYS PRO ILE LEU LEU VAL PRO GLY THR GLY THR THR GLY
SEQRES   7 A  316  PRO GLN SER PHE ASP SER ASN TRP ILE PRO LEU SER ALA
SEQRES   8 A  316  GLN LEU GLY TYR THR PRO CYS TRP ILE SER PRO PRO PRO
SEQRES   9 A  316  PHE MET LEU ASN ASP THR GLN VAL ASN THR GLU TYR MET
SEQRES  10 A  316  VAL ASN ALA ILE THR THR LEU TYR ALA GLY SER GLY ASN
SEQRES  11 A  316  ASN LYS LEU PRO VAL LEU THR TRP SER GLN GLY GLY LEU
SEQRES  12 A  316  VAL ALA GLN TRP GLY LEU THR PHE PHE PRO SER ILE ARG
SEQRES  13 A  316  SER LYS VAL ASP ARG LEU MET ALA PHE ALA PRO ASP TYR
SEQRES  14 A  316  LYS GLY THR VAL LEU ALA GLY PRO LEU ASP ALA LEU ALA
SEQRES  15 A  316  VAL SER ALA PRO SER VAL TRP GLN GLN THR THR GLY SER
SEQRES  16 A  316  ALA LEU THR THR ALA LEU ARG ASN ALA GLY GLY LEU THR
SEQRES  17 A  316  GLN ILE VAL PRO THR THR ASN LEU TYR SER ALA THR ASP
SEQRES  18 A  316  GLU ILE VAL GLN PRO GLN VAL SER ASN SER PRO LEU ASP
SEQRES  19 A  316  SER SER TYR LEU PHE ASN GLY LYS ASN VAL GLN ALA GLN
SEQRES  20 A  316  ALA VAL CYS GLY PRO LEU PHE VAL ILE ASP HIS ALA GLY
SEQRES  21 A  316  SER LEU THR SER GLN PHE SER TYR VAL VAL GLY ARG SER
SEQRES  22 A  316  ALA LEU ARG SER THR THR GLY GLN ALA ARG SER ALA ASP
SEQRES  23 A  316  TYR GLY ILE THR ASP CYS ASN PRO LEU PRO ALA ASN ASP
SEQRES  24 A  316  LEU THR PRO GLU GLN LYS VAL ALA ALA ALA ALA LEU LEU
SEQRES  25 A  316  ALA PRO ALA ALA
HET    NAG  A 401      15
HET    NAG  A 402      15
HET    BTB  A 403      14
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     BTB 2-[BIS-(2-HYDROXY-ETHYL)-AMINO]-2-HYDROXYMETHYL-
HETNAM   2 BTB  PROPANE-1,3-DIOL
HETSYN     NAG NAG
HETSYN     BTB BIS-TRIS BUFFER
FORMUL   2  NAG    2(C8 H15 N O6)
FORMUL   4  BTB    C8 H19 N O5
FORMUL   5  HOH   *413(H2 O)
HELIX    1   1 ALA A   19  ALA A   23  5                                   5
HELIX    2   2 PRO A   46  GLY A   53  1                                   8
HELIX    3   3 THR A   77  ASP A   83  1                                   7
HELIX    4   4 ASN A   85  LEU A   93  1                                   9
HELIX    5   5 ASP A  109  SER A  128  1                                  20
HELIX    6   6 SER A  139  PHE A  152  1                                  14
HELIX    7   7 PRO A  153  ARG A  156  5                                   4
HELIX    8   8 ALA A  185  GLN A  191  1                                   7
HELIX    9   9 SER A  195  ALA A  204  1                                  10
HELIX   10  10 ALA A  246  GLY A  251  1                                   6
HELIX   11  11 ALA A  259  SER A  264  1                                   6
HELIX   12  12 SER A  264  SER A  277  1                                  14
HELIX   13  13 ARG A  283  TYR A  287  5                                   5
HELIX   14  14 GLY A  288  CYS A  292  5                                   5
SHEET    1   A 2 ARG A  27  THR A  28  0
SHEET    2   A 2 GLY A  31  ILE A  32 -1  O  GLY A  31   N  THR A  28
SHEET    1   B 7 LEU A  54  CYS A  56  0
SHEET    2   B 7 THR A  96  ILE A 100 -1  O  TRP A  99   N  THR A  55
SHEET    3   B 7 PRO A  67  VAL A  71  1  N  LEU A  70   O  CYS A  98
SHEET    4   B 7 LEU A 133  TRP A 138  1  O  LEU A 136   N  LEU A  69
SHEET    5   B 7 VAL A 159  PHE A 165  1  O  MET A 163   N  VAL A 135
SHEET    6   B 7 THR A 213  TYR A 217  1  O  THR A 214   N  ALA A 164
SHEET    7   B 7 LYS A 242  GLN A 245  1  O  VAL A 244   N  ASN A 215
SSBOND   1 CYS A   11    CYS A   29                          1555   1555  2.04
SSBOND   2 CYS A   56    CYS A   98                          1555   1555  2.06
SSBOND   3 CYS A  250    CYS A  292                          1555   1555  2.03
CISPEP   1 PRO A  103    PRO A  104          0        -0.99
CISPEP   2 GLN A  225    PRO A  226          0         0.22
SITE     1 AC1  4 VAL A 228  SER A 229  LEU A 233  HOH A 450
SITE     1 AC2  8 PRO A 104  ASN A 108  HOH A 567  HOH A 650
SITE     2 AC2  8 HOH A 676  HOH A 683  HOH A 696  HOH A 733
SITE     1 AC3  7 TRP A 138  GLU A 222  ALA A 259  HOH A 453
SITE     2 AC3  7 HOH A 672  HOH A 732  HOH A 801
CRYST1  111.680  111.680   54.997  90.00  90.00 120.00 P 32 2 1      6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008954  0.005169  0.000000        0.00000
SCALE2      0.000000  0.010339  0.000000        0.00000
SCALE3      0.000000  0.000000  0.018183        0.00000
TER    2203      ASP A 299
MASTER      332    0    3   14    9    0    5    6 2659    1   50   25
END