longtext: 3K6K-pdb

content
HEADER    HYDROLASE                               09-OCT-09   3K6K
TITLE     CRYSTAL STRUCTURE AT 2.2 ANGSTROM OF HSL-HOMOLOG ESTE7 FROM
TITLE    2 A METAGENOME LIBRARY
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ESTERASE/LIPASE;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 SYNONYM: HORMONE SENSITIVE LIPASE HOMOLOG PROTEIN;
COMPND   5 EC: 3.1.1.-;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: UNCULTURED BACTERIUM;
SOURCE   3 ORGANISM_TAXID: 77133;
SOURCE   4 GENE: ESTE7;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 511693;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PET21A
KEYWDS    ALPHA/BETA HYDROLASE FOLD
EXPDTA    X-RAY DIFFRACTION
AUTHOR    K.Y.HWANG,K.H.NAM
REVDAT   1   27-OCT-09 3K6K    0
JRNL        AUTH   K.H.NAM,K.H.RHEE,W.H.LEE,B.-C.JEONG,H.K.SONG,
JRNL        AUTH 2 K.Y.HWANG
JRNL        TITL   BIOCHEMICAL AND STRUCTURAL ANALYSIS OF
JRNL        TITL 2 HORMONE-SENSITIVE LIPASE HOMOLOG ESTE7; INSIGHT
JRNL        TITL 3 INTO THE STABILIZED DIMERIZATION OF HSL-HOMOLOG
JRNL        TITL 4 PROTEINS
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   1
REMARK   2
REMARK   2 RESOLUTION.    2.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.5.0072
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 48.27
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 95.7
REMARK   3   NUMBER OF REFLECTIONS             : 84704
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.226
REMARK   3   R VALUE            (WORKING SET) : 0.223
REMARK   3   FREE R VALUE                     : 0.282
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 4253
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.20
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.26
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 4950
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 81.09
REMARK   3   BIN R VALUE           (WORKING SET) : 0.3060
REMARK   3   BIN FREE R VALUE SET COUNT          : 257
REMARK   3   BIN FREE R VALUE                    : 0.3750
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 8853
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 49
REMARK   3   SOLVENT ATOMS            : 433
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 27.31
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -0.05000
REMARK   3    B22 (A**2) : 0.37000
REMARK   3    B33 (A**2) : -0.32000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.236
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.219
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.168
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 6.723
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.938
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.900
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  9075 ; 0.020 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 12328 ; 1.976 ; 1.989
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  1172 ; 7.372 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   352 ;39.697 ;24.205
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1501 ;19.700 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    52 ;22.529 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  1406 ; 0.148 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  6772 ; 0.009 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  5852 ; 0.944 ; 1.500
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  9347 ; 1.737 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  3223 ; 2.953 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  2981 ; 4.528 ; 4.500
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3  RIDING POSITIONS U VALUES; REFINED INDIVIDUALLY
REMARK   4
REMARK   4 3K6K COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 13-OCT-09.
REMARK 100 THE RCSB ID CODE IS RCSB055609.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 18-DEC-08
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PAL/PLS
REMARK 200  BEAMLINE                       : 6C1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.23
REMARK 200  MONOCHROMATOR                  : GRAPHITE
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 210
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 84721
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.200
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 95.6
REMARK 200  DATA REDUNDANCY                : 5.000
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.0900
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.28
REMARK 200  COMPLETENESS FOR SHELL     (%) : 82.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.820
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: PDB ENTRY 3DNM
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 60.85
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.14
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M BIS-TRIS PROPANE, PH 7.0, 0.2M
REMARK 280  AMMONIUM SULFATE, 1M LITHIUM SULFATE, VAPOR DIFFUSION, HANGING
REMARK 280  DROP, TEMPERATURE 295K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: I 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z+1/2
REMARK 290       6555   -X,-Y+1/2,Z
REMARK 290       7555   -X+1/2,Y,-Z
REMARK 290       8555   X,-Y,-Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       58.57850
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      116.43500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       63.84950
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      116.43500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       58.57850
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       63.84950
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       58.57850
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       63.84950
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000      116.43500
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       63.84950
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       58.57850
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000      116.43500
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3, 4, 5, 6
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 300 REMARK: RECENTLY, THE DEPOSITORS HAVE CHARACTERIZED THE
REMARK 300 OLIGOMERIC STATE OF THIS PROTEIN. THIS PROTEIN REVEALS THE
REMARK 300 MONOMER AND DIMER FORMATION. BOTH OLIGOMERIC STATE REVEALS THE
REMARK 300 ENZYME ACTIVITY. BASED ON OUR STUDY, BIOLOGICAL ASSEMBLY OF
REMARK 300 ESTE7 ARE FORMING THE MONOMER AND DIMER FORMATION. ESPECIALLY,
REMARK 300 DIMERIC ESTE7 HAVE HIGH ACTIVITY THAN MONOMER ESTE7.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2790 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 22780 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -78.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2770 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 22020 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -59.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 4
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 5
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 6
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     GLY A     2
REMARK 465     ALA A     3
REMARK 465     MET A     4
REMARK 465     ASP A     5
REMARK 465     GLN A     6
REMARK 465     GLU A     7
REMARK 465     ILE A     8
REMARK 465     GLY A     9
REMARK 465     THR A    10
REMARK 465     VAL A    11
REMARK 465     THR A    12
REMARK 465     LYS A   310
REMARK 465     LEU A   311
REMARK 465     ALA A   312
REMARK 465     ALA A   313
REMARK 465     ALA A   314
REMARK 465     LEU A   315
REMARK 465     GLU A   316
REMARK 465     HIS A   317
REMARK 465     HIS A   318
REMARK 465     HIS A   319
REMARK 465     HIS A   320
REMARK 465     HIS A   321
REMARK 465     HIS A   322
REMARK 465     MET B     1
REMARK 465     GLY B     2
REMARK 465     ALA B     3
REMARK 465     MET B     4
REMARK 465     ASP B     5
REMARK 465     GLN B     6
REMARK 465     GLU B     7
REMARK 465     ILE B     8
REMARK 465     GLY B     9
REMARK 465     THR B    10
REMARK 465     VAL B    11
REMARK 465     THR B    12
REMARK 465     LYS B   310
REMARK 465     LEU B   311
REMARK 465     ALA B   312
REMARK 465     ALA B   313
REMARK 465     ALA B   314
REMARK 465     LEU B   315
REMARK 465     GLU B   316
REMARK 465     HIS B   317
REMARK 465     HIS B   318
REMARK 465     HIS B   319
REMARK 465     HIS B   320
REMARK 465     HIS B   321
REMARK 465     HIS B   322
REMARK 465     MET C     1
REMARK 465     GLY C     2
REMARK 465     ALA C     3
REMARK 465     MET C     4
REMARK 465     ASP C     5
REMARK 465     GLN C     6
REMARK 465     GLU C     7
REMARK 465     ILE C     8
REMARK 465     GLY C     9
REMARK 465     THR C    10
REMARK 465     VAL C    11
REMARK 465     THR C    12
REMARK 465     ASP C    13
REMARK 465     GLU C    33
REMARK 465     LYS C    34
REMARK 465     ASN C    35
REMARK 465     LEU C    36
REMARK 465     PRO C    37
REMARK 465     LEU C    38
REMARK 465     LYS C   310
REMARK 465     LEU C   311
REMARK 465     ALA C   312
REMARK 465     ALA C   313
REMARK 465     ALA C   314
REMARK 465     LEU C   315
REMARK 465     GLU C   316
REMARK 465     HIS C   317
REMARK 465     HIS C   318
REMARK 465     HIS C   319
REMARK 465     HIS C   320
REMARK 465     HIS C   321
REMARK 465     HIS C   322
REMARK 465     MET D     1
REMARK 465     GLY D     2
REMARK 465     ALA D     3
REMARK 465     MET D     4
REMARK 465     ASP D     5
REMARK 465     GLN D     6
REMARK 465     GLU D     7
REMARK 465     ILE D     8
REMARK 465     GLY D     9
REMARK 465     THR D    10
REMARK 465     VAL D    11
REMARK 465     THR D    12
REMARK 465     ASP D    13
REMARK 465     THR D    14
REMARK 465     LYS D    15
REMARK 465     SER D   309
REMARK 465     LYS D   310
REMARK 465     LEU D   311
REMARK 465     ALA D   312
REMARK 465     ALA D   313
REMARK 465     ALA D   314
REMARK 465     LEU D   315
REMARK 465     GLU D   316
REMARK 465     HIS D   317
REMARK 465     HIS D   318
REMARK 465     HIS D   319
REMARK 465     HIS D   320
REMARK 465     HIS D   321
REMARK 465     HIS D   322
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OG   SER C   157     O    HOH C   810              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ARG A 194   NE  -  CZ  -  NH1 ANGL. DEV. =   3.3 DEGREES
REMARK 500    ARG A 194   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.1 DEGREES
REMARK 500    ARG A 203   NE  -  CZ  -  NH1 ANGL. DEV. =   3.6 DEGREES
REMARK 500    ARG A 307   NE  -  CZ  -  NH1 ANGL. DEV. =   3.3 DEGREES
REMARK 500    ASP B  17   CB  -  CG  -  OD2 ANGL. DEV. =  -5.4 DEGREES
REMARK 500    ARG B 120   NE  -  CZ  -  NH1 ANGL. DEV. =   3.2 DEGREES
REMARK 500    ARG B 120   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.8 DEGREES
REMARK 500    ARG B 194   NE  -  CZ  -  NH1 ANGL. DEV. =   3.3 DEGREES
REMARK 500    ARG B 194   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.0 DEGREES
REMARK 500    ARG B 203   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.9 DEGREES
REMARK 500    ASP B 224   CB  -  CG  -  OD1 ANGL. DEV. =   5.9 DEGREES
REMARK 500    LEU B 273   CA  -  CB  -  CG  ANGL. DEV. =  14.3 DEGREES
REMARK 500    VAL C 101   CB  -  CA  -  C   ANGL. DEV. = -12.5 DEGREES
REMARK 500    ARG C 225   NE  -  CZ  -  NH2 ANGL. DEV. =   3.2 DEGREES
REMARK 500    LEU C 273   CA  -  CB  -  CG  ANGL. DEV. =  15.1 DEGREES
REMARK 500    ARG D 194   NE  -  CZ  -  NH1 ANGL. DEV. =   3.3 DEGREES
REMARK 500    ARG D 194   NE  -  CZ  -  NH2 ANGL. DEV. =  -5.4 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    THR A  63     -159.94   -143.12
REMARK 500    SER A 157     -115.61     47.18
REMARK 500    LEU A 190        2.17    -68.50
REMARK 500    SER A 193      -12.78   -146.32
REMARK 500    LEU A 212      -39.08    -34.88
REMARK 500    ASP A 278       15.08     59.14
REMARK 500    VAL A 282       55.49     39.48
REMARK 500    ALA B  32      129.70    -35.36
REMARK 500    ASN B  35       62.34   -117.93
REMARK 500    THR B  63     -158.88   -164.59
REMARK 500    ASP B  76       38.31    -88.49
REMARK 500    HIS B  87      161.88    -47.02
REMARK 500    TYR B  91       -5.61     77.05
REMARK 500    ASN B 125       86.79   -151.25
REMARK 500    SER B 157     -129.34     59.60
REMARK 500    GLU B 208       48.77     72.42
REMARK 500    THR B 211      -14.33   -141.40
REMARK 500    GLU B 250       53.15    -99.71
REMARK 500    ILE B 308       76.84    -68.23
REMARK 500    GLU C  30       -6.00    -57.15
REMARK 500    THR C  63     -158.82   -150.54
REMARK 500    TYR C  91       -0.87     66.24
REMARK 500    SER C 157     -129.16     56.54
REMARK 500    SER C 185       56.40     35.29
REMARK 500    VAL C 282       58.62     30.92
REMARK 500    ALA D  32      138.29    -33.95
REMARK 500    ASN D  35       99.10   -160.46
REMARK 500    PRO D  37     -175.00    -65.99
REMARK 500    GLN D  40      -73.33    -71.72
REMARK 500    ARG D  51        8.47    -60.99
REMARK 500    THR D  63     -164.29   -166.58
REMARK 500    TYR D  91       -6.03     85.85
REMARK 500    ASP D 118       69.83   -101.05
REMARK 500    SER D 157     -127.39     48.76
REMARK 500    ALA D 207       69.51   -107.86
REMARK 500    VAL D 233      -35.07    -38.04
REMARK 500    GLU D 250       48.62    -83.34
REMARK 500    ASP D 278       -1.08     76.65
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (11X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500   M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500     ASP B 210        22.4      L          L   OUTSIDE RANGE
REMARK 500     ASN B 291         7.4      L          L   EXPECTING SP3
REMARK 500     SER B 309        24.6      L          L   OUTSIDE RANGE
REMARK 500     THR C 163        21.9      L          L   OUTSIDE RANGE
REMARK 500     LYS D  34        23.0      L          L   OUTSIDE RANGE
REMARK 500     GLU D 124        24.8      L          L   OUTSIDE RANGE
REMARK 500     THR D 163        23.8      L          L   OUTSIDE RANGE
REMARK 500     MET D 215        24.5      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A 323
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A 324
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 B 323
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 B 324
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 B 325
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 C 323
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 C 324
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 D 323
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 D 324
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE BME D 325
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3DNM   RELATED DB: PDB
REMARK 900 THE SAME PROTEIN AT 2.8 ANGSTROM
DBREF  3K6K A    1   309  UNP    Q0GMU1   Q0GMU1_9BACT     1    309
DBREF  3K6K B    1   309  UNP    Q0GMU1   Q0GMU1_9BACT     1    309
DBREF  3K6K C    1   309  UNP    Q0GMU1   Q0GMU1_9BACT     1    309
DBREF  3K6K D    1   309  UNP    Q0GMU1   Q0GMU1_9BACT     1    309
SEQADV 3K6K LYS A  310  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU A  311  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA A  312  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA A  313  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA A  314  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU A  315  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K GLU A  316  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS A  317  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS A  318  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS A  319  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS A  320  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS A  321  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS A  322  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LYS B  310  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU B  311  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA B  312  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA B  313  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA B  314  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU B  315  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K GLU B  316  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS B  317  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS B  318  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS B  319  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS B  320  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS B  321  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS B  322  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LYS C  310  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU C  311  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA C  312  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA C  313  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA C  314  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU C  315  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K GLU C  316  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS C  317  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS C  318  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS C  319  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS C  320  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS C  321  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS C  322  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LYS D  310  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU D  311  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA D  312  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA D  313  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K ALA D  314  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K LEU D  315  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K GLU D  316  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS D  317  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS D  318  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS D  319  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS D  320  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS D  321  UNP  Q0GMU1              EXPRESSION TAG
SEQADV 3K6K HIS D  322  UNP  Q0GMU1              EXPRESSION TAG
SEQRES   1 A  322  MET GLY ALA MET ASP GLN GLU ILE GLY THR VAL THR ASP
SEQRES   2 A  322  THR LYS MET ASP PRO ARG ASP PHE LEU GLN LEU LEU LYS
SEQRES   3 A  322  ILE ASN ALA GLU LYS ALA GLU LYS ASN LEU PRO LEU ASP
SEQRES   4 A  322  GLN LYS ARG ALA GLY MET GLU ALA LEU CYS GLU ARG PHE
SEQRES   5 A  322  PRO ARG ALA GLU GLY VAL GLU LEU THR LEU THR ASP LEU
SEQRES   6 A  322  GLY GLY VAL PRO CYS ILE ARG GLN ALA THR ASP GLY ALA
SEQRES   7 A  322  GLY ALA ALA HIS ILE LEU TYR PHE HIS GLY GLY GLY TYR
SEQRES   8 A  322  ILE SER GLY SER PRO SER THR HIS LEU VAL LEU THR THR
SEQRES   9 A  322  GLN LEU ALA LYS GLN SER SER ALA THR LEU TRP SER LEU
SEQRES  10 A  322  ASP TYR ARG LEU ALA PRO GLU ASN PRO PHE PRO ALA ALA
SEQRES  11 A  322  VAL ASP ASP CYS VAL ALA ALA TYR ARG ALA LEU LEU LYS
SEQRES  12 A  322  THR ALA GLY SER ALA ASP ARG ILE ILE ILE ALA GLY ASP
SEQRES  13 A  322  SER ALA GLY GLY GLY LEU THR THR ALA SER MET LEU LYS
SEQRES  14 A  322  ALA LYS GLU ASP GLY LEU PRO MET PRO ALA GLY LEU VAL
SEQRES  15 A  322  MET LEU SER PRO PHE VAL ASP LEU THR LEU SER ARG TRP
SEQRES  16 A  322  SER ASN SER ASN LEU ALA ASP ARG ASP PHE LEU ALA GLU
SEQRES  17 A  322  PRO ASP THR LEU GLY GLU MET SER GLU LEU TYR VAL GLY
SEQRES  18 A  322  GLY GLU ASP ARG LYS ASN PRO LEU ILE SER PRO VAL TYR
SEQRES  19 A  322  ALA ASP LEU SER GLY LEU PRO GLU MET LEU ILE HIS VAL
SEQRES  20 A  322  GLY SER GLU GLU ALA LEU LEU SER ASP SER THR THR LEU
SEQRES  21 A  322  ALA GLU ARG ALA GLY ALA ALA GLY VAL SER VAL GLU LEU
SEQRES  22 A  322  LYS ILE TRP PRO ASP MET PRO HIS VAL PHE GLN MET TYR
SEQRES  23 A  322  GLY LYS PHE VAL ASN ALA ALA ASP ILE SER ILE LYS GLU
SEQRES  24 A  322  ILE CYS HIS TRP ILE SER ALA ARG ILE SER LYS LEU ALA
SEQRES  25 A  322  ALA ALA LEU GLU HIS HIS HIS HIS HIS HIS
SEQRES   1 B  322  MET GLY ALA MET ASP GLN GLU ILE GLY THR VAL THR ASP
SEQRES   2 B  322  THR LYS MET ASP PRO ARG ASP PHE LEU GLN LEU LEU LYS
SEQRES   3 B  322  ILE ASN ALA GLU LYS ALA GLU LYS ASN LEU PRO LEU ASP
SEQRES   4 B  322  GLN LYS ARG ALA GLY MET GLU ALA LEU CYS GLU ARG PHE
SEQRES   5 B  322  PRO ARG ALA GLU GLY VAL GLU LEU THR LEU THR ASP LEU
SEQRES   6 B  322  GLY GLY VAL PRO CYS ILE ARG GLN ALA THR ASP GLY ALA
SEQRES   7 B  322  GLY ALA ALA HIS ILE LEU TYR PHE HIS GLY GLY GLY TYR
SEQRES   8 B  322  ILE SER GLY SER PRO SER THR HIS LEU VAL LEU THR THR
SEQRES   9 B  322  GLN LEU ALA LYS GLN SER SER ALA THR LEU TRP SER LEU
SEQRES  10 B  322  ASP TYR ARG LEU ALA PRO GLU ASN PRO PHE PRO ALA ALA
SEQRES  11 B  322  VAL ASP ASP CYS VAL ALA ALA TYR ARG ALA LEU LEU LYS
SEQRES  12 B  322  THR ALA GLY SER ALA ASP ARG ILE ILE ILE ALA GLY ASP
SEQRES  13 B  322  SER ALA GLY GLY GLY LEU THR THR ALA SER MET LEU LYS
SEQRES  14 B  322  ALA LYS GLU ASP GLY LEU PRO MET PRO ALA GLY LEU VAL
SEQRES  15 B  322  MET LEU SER PRO PHE VAL ASP LEU THR LEU SER ARG TRP
SEQRES  16 B  322  SER ASN SER ASN LEU ALA ASP ARG ASP PHE LEU ALA GLU
SEQRES  17 B  322  PRO ASP THR LEU GLY GLU MET SER GLU LEU TYR VAL GLY
SEQRES  18 B  322  GLY GLU ASP ARG LYS ASN PRO LEU ILE SER PRO VAL TYR
SEQRES  19 B  322  ALA ASP LEU SER GLY LEU PRO GLU MET LEU ILE HIS VAL
SEQRES  20 B  322  GLY SER GLU GLU ALA LEU LEU SER ASP SER THR THR LEU
SEQRES  21 B  322  ALA GLU ARG ALA GLY ALA ALA GLY VAL SER VAL GLU LEU
SEQRES  22 B  322  LYS ILE TRP PRO ASP MET PRO HIS VAL PHE GLN MET TYR
SEQRES  23 B  322  GLY LYS PHE VAL ASN ALA ALA ASP ILE SER ILE LYS GLU
SEQRES  24 B  322  ILE CYS HIS TRP ILE SER ALA ARG ILE SER LYS LEU ALA
SEQRES  25 B  322  ALA ALA LEU GLU HIS HIS HIS HIS HIS HIS
SEQRES   1 C  322  MET GLY ALA MET ASP GLN GLU ILE GLY THR VAL THR ASP
SEQRES   2 C  322  THR LYS MET ASP PRO ARG ASP PHE LEU GLN LEU LEU LYS
SEQRES   3 C  322  ILE ASN ALA GLU LYS ALA GLU LYS ASN LEU PRO LEU ASP
SEQRES   4 C  322  GLN LYS ARG ALA GLY MET GLU ALA LEU CYS GLU ARG PHE
SEQRES   5 C  322  PRO ARG ALA GLU GLY VAL GLU LEU THR LEU THR ASP LEU
SEQRES   6 C  322  GLY GLY VAL PRO CYS ILE ARG GLN ALA THR ASP GLY ALA
SEQRES   7 C  322  GLY ALA ALA HIS ILE LEU TYR PHE HIS GLY GLY GLY TYR
SEQRES   8 C  322  ILE SER GLY SER PRO SER THR HIS LEU VAL LEU THR THR
SEQRES   9 C  322  GLN LEU ALA LYS GLN SER SER ALA THR LEU TRP SER LEU
SEQRES  10 C  322  ASP TYR ARG LEU ALA PRO GLU ASN PRO PHE PRO ALA ALA
SEQRES  11 C  322  VAL ASP ASP CYS VAL ALA ALA TYR ARG ALA LEU LEU LYS
SEQRES  12 C  322  THR ALA GLY SER ALA ASP ARG ILE ILE ILE ALA GLY ASP
SEQRES  13 C  322  SER ALA GLY GLY GLY LEU THR THR ALA SER MET LEU LYS
SEQRES  14 C  322  ALA LYS GLU ASP GLY LEU PRO MET PRO ALA GLY LEU VAL
SEQRES  15 C  322  MET LEU SER PRO PHE VAL ASP LEU THR LEU SER ARG TRP
SEQRES  16 C  322  SER ASN SER ASN LEU ALA ASP ARG ASP PHE LEU ALA GLU
SEQRES  17 C  322  PRO ASP THR LEU GLY GLU MET SER GLU LEU TYR VAL GLY
SEQRES  18 C  322  GLY GLU ASP ARG LYS ASN PRO LEU ILE SER PRO VAL TYR
SEQRES  19 C  322  ALA ASP LEU SER GLY LEU PRO GLU MET LEU ILE HIS VAL
SEQRES  20 C  322  GLY SER GLU GLU ALA LEU LEU SER ASP SER THR THR LEU
SEQRES  21 C  322  ALA GLU ARG ALA GLY ALA ALA GLY VAL SER VAL GLU LEU
SEQRES  22 C  322  LYS ILE TRP PRO ASP MET PRO HIS VAL PHE GLN MET TYR
SEQRES  23 C  322  GLY LYS PHE VAL ASN ALA ALA ASP ILE SER ILE LYS GLU
SEQRES  24 C  322  ILE CYS HIS TRP ILE SER ALA ARG ILE SER LYS LEU ALA
SEQRES  25 C  322  ALA ALA LEU GLU HIS HIS HIS HIS HIS HIS
SEQRES   1 D  322  MET GLY ALA MET ASP GLN GLU ILE GLY THR VAL THR ASP
SEQRES   2 D  322  THR LYS MET ASP PRO ARG ASP PHE LEU GLN LEU LEU LYS
SEQRES   3 D  322  ILE ASN ALA GLU LYS ALA GLU LYS ASN LEU PRO LEU ASP
SEQRES   4 D  322  GLN LYS ARG ALA GLY MET GLU ALA LEU CYS GLU ARG PHE
SEQRES   5 D  322  PRO ARG ALA GLU GLY VAL GLU LEU THR LEU THR ASP LEU
SEQRES   6 D  322  GLY GLY VAL PRO CYS ILE ARG GLN ALA THR ASP GLY ALA
SEQRES   7 D  322  GLY ALA ALA HIS ILE LEU TYR PHE HIS GLY GLY GLY TYR
SEQRES   8 D  322  ILE SER GLY SER PRO SER THR HIS LEU VAL LEU THR THR
SEQRES   9 D  322  GLN LEU ALA LYS GLN SER SER ALA THR LEU TRP SER LEU
SEQRES  10 D  322  ASP TYR ARG LEU ALA PRO GLU ASN PRO PHE PRO ALA ALA
SEQRES  11 D  322  VAL ASP ASP CYS VAL ALA ALA TYR ARG ALA LEU LEU LYS
SEQRES  12 D  322  THR ALA GLY SER ALA ASP ARG ILE ILE ILE ALA GLY ASP
SEQRES  13 D  322  SER ALA GLY GLY GLY LEU THR THR ALA SER MET LEU LYS
SEQRES  14 D  322  ALA LYS GLU ASP GLY LEU PRO MET PRO ALA GLY LEU VAL
SEQRES  15 D  322  MET LEU SER PRO PHE VAL ASP LEU THR LEU SER ARG TRP
SEQRES  16 D  322  SER ASN SER ASN LEU ALA ASP ARG ASP PHE LEU ALA GLU
SEQRES  17 D  322  PRO ASP THR LEU GLY GLU MET SER GLU LEU TYR VAL GLY
SEQRES  18 D  322  GLY GLU ASP ARG LYS ASN PRO LEU ILE SER PRO VAL TYR
SEQRES  19 D  322  ALA ASP LEU SER GLY LEU PRO GLU MET LEU ILE HIS VAL
SEQRES  20 D  322  GLY SER GLU GLU ALA LEU LEU SER ASP SER THR THR LEU
SEQRES  21 D  322  ALA GLU ARG ALA GLY ALA ALA GLY VAL SER VAL GLU LEU
SEQRES  22 D  322  LYS ILE TRP PRO ASP MET PRO HIS VAL PHE GLN MET TYR
SEQRES  23 D  322  GLY LYS PHE VAL ASN ALA ALA ASP ILE SER ILE LYS GLU
SEQRES  24 D  322  ILE CYS HIS TRP ILE SER ALA ARG ILE SER LYS LEU ALA
SEQRES  25 D  322  ALA ALA LEU GLU HIS HIS HIS HIS HIS HIS
HET    SO4  A 323       5
HET    SO4  A 324       5
HET    SO4  B 323       5
HET    SO4  B 324       5
HET    SO4  B 325       5
HET    SO4  C 323       5
HET    SO4  C 324       5
HET    SO4  D 323       5
HET    SO4  D 324       5
HET    BME  D 325       4
HETNAM     SO4 SULFATE ION
HETNAM     BME BETA-MERCAPTOETHANOL
FORMUL   5  SO4    9(O4 S 2-)
FORMUL  14  BME    C2 H6 O S
FORMUL  15  HOH   *433(H2 O)
HELIX    1   1 ASP A   17  LYS A   31  1                                  15
HELIX    2   2 PRO A   37  GLU A   50  1                                  14
HELIX    3   3 SER A   95  SER A  111  1                                  17
HELIX    4   4 PRO A  128  GLY A  146  1                                  19
HELIX    5   5 SER A  147  ASP A  149  5                                   3
HELIX    6   6 SER A  157  ASP A  173  1                                  17
HELIX    7   7 ARG A  194  LEU A  200  1                                   7
HELIX    8   8 ALA A  201  ASP A  204  5                                   4
HELIX    9   9 GLU A  208  GLY A  221  1                                  14
HELIX   10  10 SER A  231  ALA A  235  5                                   5
HELIX   11  11 LEU A  253  ALA A  267  1                                  15
HELIX   12  12 VAL A  282  GLY A  287  5                                   6
HELIX   13  13 VAL A  290  ALA A  306  1                                  17
HELIX   14  14 ASP B   17  ALA B   32  1                                  16
HELIX   15  15 PRO B   37  GLU B   50  1                                  14
HELIX   16  16 HIS B   99  SER B  111  1                                  13
HELIX   17  17 PRO B  128  GLY B  146  1                                  19
HELIX   18  18 SER B  147  ASP B  149  5                                   3
HELIX   19  19 SER B  157  ASP B  173  1                                  17
HELIX   20  20 ARG B  194  LEU B  200  1                                   7
HELIX   21  21 ALA B  201  ASP B  204  5                                   4
HELIX   22  22 GLU B  208  GLY B  221  1                                  14
HELIX   23  23 SER B  231  ALA B  235  5                                   5
HELIX   24  24 LEU B  253  ALA B  267  1                                  15
HELIX   25  25 VAL B  282  GLY B  287  5                                   6
HELIX   26  26 VAL B  290  ILE B  308  1                                  19
HELIX   27  27 ASP C   17  GLU C   30  1                                  14
HELIX   28  28 GLN C   40  ARG C   51  1                                  12
HELIX   29  29 HIS C   99  SER C  111  1                                  13
HELIX   30  30 PRO C  128  GLY C  146  1                                  19
HELIX   31  31 SER C  147  ASP C  149  5                                   3
HELIX   32  32 SER C  157  ASP C  173  1                                  17
HELIX   33  33 ARG C  194  LEU C  200  1                                   7
HELIX   34  34 ALA C  201  ASP C  204  5                                   4
HELIX   35  35 GLU C  208  GLY C  221  1                                  14
HELIX   36  36 SER C  231  ALA C  235  5                                   5
HELIX   37  37 LEU C  253  ALA C  267  1                                  15
HELIX   38  38 VAL C  282  TYR C  286  5                                   5
HELIX   39  39 VAL C  290  SER C  309  1                                  20
HELIX   40  40 ASP D   17  LYS D   31  1                                  15
HELIX   41  41 PRO D   37  ARG D   51  1                                  15
HELIX   42  42 HIS D   99  SER D  111  1                                  13
HELIX   43  43 PRO D  128  GLY D  146  1                                  19
HELIX   44  44 SER D  147  ASP D  149  5                                   3
HELIX   45  45 ALA D  158  ASP D  173  1                                  16
HELIX   46  46 ARG D  194  LEU D  200  1                                   7
HELIX   47  47 GLU D  208  GLY D  221  1                                  14
HELIX   48  48 SER D  231  ALA D  235  5                                   5
HELIX   49  49 LEU D  253  ALA D  267  1                                  15
HELIX   50  50 VAL D  282  TYR D  286  5                                   5
HELIX   51  51 ASN D  291  ILE D  308  1                                  18
SHEET    1   A32 GLU A  59  LEU A  65  0
SHEET    2   A32 VAL A  68  ALA A  74 -1  O  CYS A  70   N  THR A  63
SHEET    3   A32 THR A 113  LEU A 117 -1  O  LEU A 114   N  GLN A  73
SHEET    4   A32 HIS A  82  PHE A  86  1  N  ILE A  83   O  TRP A 115
SHEET    5   A32 ILE A 151  ASP A 156  1  O  ILE A 152   N  LEU A  84
SHEET    6   A32 GLY A 180  LEU A 184  1  O  VAL A 182   N  ILE A 153
SHEET    7   A32 MET A 243  GLY A 248  1  O  HIS A 246   N  MET A 183
SHEET    8   A32 VAL A 271  TRP A 276  1  O  TRP A 276   N  VAL A 247
SHEET    9   A32 VAL B 271  PRO B 280 -1  O  LEU B 273   N  LEU A 273
SHEET   10   A32 MET B 243  GLU B 250  1  N  ILE B 245   O  GLU B 272
SHEET   11   A32 GLY B 180  LEU B 184  1  N  MET B 183   O  LEU B 244
SHEET   12   A32 ILE B 151  ASP B 156  1  N  GLY B 155   O  LEU B 184
SHEET   13   A32 ALA B  81  PHE B  86  1  N  LEU B  84   O  ALA B 154
SHEET   14   A32 ALA B 112  LEU B 117  1  O  TRP B 115   N  ILE B  83
SHEET   15   A32 VAL B  68  ALA B  74 -1  N  GLN B  73   O  LEU B 114
SHEET   16   A32 GLU B  59  LEU B  65 -1  N  GLU B  59   O  ALA B  74
SHEET   17   A32 VAL C  58  LEU C  65 -1  O  LEU C  62   N  LEU B  60
SHEET   18   A32 VAL C  68  THR C  75 -1  O  ALA C  74   N  GLU C  59
SHEET   19   A32 THR C 113  LEU C 117 -1  O  SER C 116   N  ILE C  71
SHEET   20   A32 HIS C  82  PHE C  86  1  N  ILE C  83   O  TRP C 115
SHEET   21   A32 ILE C 151  ASP C 156  1  O  ALA C 154   N  LEU C  84
SHEET   22   A32 GLY C 180  LEU C 184  1  O  VAL C 182   N  ILE C 153
SHEET   23   A32 MET C 243  GLY C 248  1  O  LEU C 244   N  MET C 183
SHEET   24   A32 VAL C 271  TRP C 276  1  O  TRP C 276   N  VAL C 247
SHEET   25   A32 VAL D 271  PRO D 280 -1  O  LEU D 273   N  LEU C 273
SHEET   26   A32 MET D 243  GLU D 250  1  N  ILE D 245   O  GLU D 272
SHEET   27   A32 GLY D 180  LEU D 184  1  N  MET D 183   O  LEU D 244
SHEET   28   A32 ILE D 151  ASP D 156  1  N  ILE D 153   O  VAL D 182
SHEET   29   A32 HIS D  82  PHE D  86  1  N  LEU D  84   O  ILE D 152
SHEET   30   A32 THR D 113  LEU D 117  1  O  TRP D 115   N  ILE D  83
SHEET   31   A32 VAL D  68  ALA D  74 -1  N  GLN D  73   O  LEU D 114
SHEET   32   A32 GLU D  59  LEU D  65 -1  N  GLU D  59   O  ALA D  74
CISPEP   1 ALA A  122    PRO A  123          0         4.84
CISPEP   2 PHE A  127    PRO A  128          0         8.45
CISPEP   3 ALA B  122    PRO B  123          0         3.08
CISPEP   4 PHE B  127    PRO B  128          0        10.53
CISPEP   5 ALA C  122    PRO C  123          0         0.07
CISPEP   6 PHE C  127    PRO C  128          0        10.74
CISPEP   7 ALA D  122    PRO D  123          0         4.37
CISPEP   8 PHE D  127    PRO D  128          0         9.44
SITE     1 AC1  6 ARG A  54  ALA A  55  GLU A  56  GLN A 105
SITE     2 AC1  6 LYS A 108  HOH A 411
SITE     1 AC2  3 ARG A 194  TYR A 234  ARG A 263
SITE     1 AC3  3 GLU B  56  GLN B 105  LYS B 108
SITE     1 AC4  2 SER B 305  ILE B 308
SITE     1 AC5  5 THR A 258  THR A 259  THR B 258  THR B 259
SITE     2 AC5  5 GLU B 262
SITE     1 AC6  3 GLU C  56  GLN C 105  LYS C 108
SITE     1 AC7  3 ARG C 194  TYR C 234  ARG C 263
SITE     1 AC8  4 ALA D  55  GLU D  56  GLN D 105  LYS D 108
SITE     1 AC9  4 ARG D 194  TYR D 234  ARG D 263  HOH D 338
SITE     1 BC1  3 SER D 157  LEU D 206  HIS D 281
CRYST1  117.157  127.699  232.870  90.00  90.00  90.00 I 21 21 21   32
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008536  0.000000  0.000000        0.00000
SCALE2      0.000000  0.007831  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004294        0.00000
TER    2236      SER A 309
TER    4472      SER B 309
TER    6651      SER C 309
TER    8857      ILE D 308
MASTER      574    0   10   51   32    0   12    6 9335    4   49  100
END