longtext: 3K9B-pdb

content
HEADER    HYDROLASE                               15-OCT-09   3K9B
TITLE     CRYSTAL STRUCTURE OF HUMAN LIVER CARBOXYLESTERASE 1 (HCE1)
TITLE    2 IN COVALENT COMPLEX WITH THE NERVE AGENT CYCLOSARIN (GF)
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIVER CARBOXYLESTERASE 1;
COMPND   3 CHAIN: A, B, C;
COMPND   4 SYNONYM: ACYL COENZYME A:CHOLESTEROL ACYLTRANSFERASE, ACAT,
COMPND   5 MONOCYTE/MACROPHAGE SERINE ESTERASE, HMSE, SERINE ESTERASE
COMPND   6 1, BRAIN CARBOXYLESTERASE HBR1, TRIACYLGLYCEROL HYDROLASE,
COMPND   7 TGH, EGASYN, RETINYL ESTER HYDROLASE, REH;
COMPND   8 EC: 3.1.1.1;
COMPND   9 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: CES1, CES2, SES1;
SOURCE   6 EXPRESSION_SYSTEM: SPODOPTERA FRUGIPERDA;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 7108;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: SF21;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PBAK
KEYWDS    HYDROLASE, ORGANOPHOSPHORUS NERVE AGENT, ALTERNATIVE
KEYWDS   2 SPLICING, DISULFIDE BOND, ENDOPLASMIC RETICULUM,
KEYWDS   3 GLYCOPROTEIN, POLYMORPHISM, SERINE ESTERASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    A.C.HEMMERT,M.R.REDINBO
REVDAT   1   19-JAN-10 3K9B    0
JRNL        AUTH   A.C.HEMMERT,T.C.OTTO,M.WEIRDL,P.M.POTTER,
JRNL        AUTH 2 D.M.CERASOLI,M.R.REDINBO
JRNL        TITL   STEREOSPECIFIC REACTIONS OF HCE1 WITH NERVE AGENTS
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   1
REMARK   2
REMARK   2 RESOLUTION.    3.10 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 3.10
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 48.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 97.5
REMARK   3   NUMBER OF REFLECTIONS             : 36488
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.266
REMARK   3   FREE R VALUE                     : 0.299
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 9.700
REMARK   3   FREE R VALUE TEST SET COUNT      : 3643
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 3.10
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 3.30
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 94.00
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2680
REMARK   3   BIN FREE R VALUE                    : 0.2960
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 11155
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 30
REMARK   3   SOLVENT ATOMS            : 66
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 53.94
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 49.19
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -9.64200
REMARK   3    B22 (A**2) : 11.39900
REMARK   3    B33 (A**2) : -1.75800
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.014
REMARK   3   BOND ANGLES            (DEGREES) : 1.70
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : ANISOTROPIC OVERALL TEMPERATURE
REMARK   3                            FACTORS
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : NULL
REMARK   3   KSOL        : NULL
REMARK   3   BSOL        : 53.94
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : CNS_TOPPAR:PROTEIN_REP.PARAM
REMARK   3  PARAMETER FILE  2  : GFC.PAR
REMARK   3  PARAMETER FILE  3  : CNS_TOPPAR:WATER_REP.PARAM
REMARK   3  PARAMETER FILE  4  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3  TOPOLOGY FILE  2   : NULL
REMARK   3  TOPOLOGY FILE  3   : NULL
REMARK   3  TOPOLOGY FILE  4   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3K9B COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 22-OCT-09.
REMARK 100 THE RCSB ID CODE IS RCSB055708.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 24-MAR-08
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 22-ID
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0000
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MAR CCD 165 MM
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 36488
REMARK 200  RESOLUTION RANGE HIGH      (A) : 3.100
REMARK 200  RESOLUTION RANGE LOW       (A) : 48.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.100
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.5
REMARK 200  DATA REDUNDANCY                : 7.000
REMARK 200  R MERGE                    (I) : 0.13100
REMARK 200  R SYM                      (I) : 0.13100
REMARK 200   FOR THE DATA SET  : 13.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 3.10
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 3.20
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.48100
REMARK 200  R SYM FOR SHELL            (I) : 0.48100
REMARK 200   FOR SHELL         : 4.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: PDB ENTRY 1MX1
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 56.93
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.86
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 10% PEG 3350, 0.3 M LI2SO4, 0.1 M
REMARK 280  CITRATE PH 5.5, 0.1 M NACL, 0.1 M LICL, 5% GLYCEROL, VAPOR
REMARK 280  DIFFUSION, SITTING DROP, TEMPERATURE 295K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       27.80500
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      100.03000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       89.94000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      100.03000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       27.80500
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       89.94000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3, 4
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: TRIMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B, C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 4
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     LEU A  1304
REMARK 465     SER A  1305
REMARK 465     LEU A  1306
REMARK 465     ASP A  1307
REMARK 465     LEU A  1308
REMARK 465     GLN A  1309
REMARK 465     GLY A  1310
REMARK 465     ASP A  1311
REMARK 465     PRO A  1312
REMARK 465     ARG A  1313
REMARK 465     GLU A  1314
REMARK 465     SER A  1315
REMARK 465     GLN A  1316
REMARK 465     PRO A  1317
REMARK 465     LEU A  1318
REMARK 465     ASN A  1340
REMARK 465     PHE A  1341
REMARK 465     HIS A  1342
REMARK 465     SER A  1369
REMARK 465     GLU A  1370
REMARK 465     GLY A  1371
REMARK 465     GLN A  1372
REMARK 465     LEU A  1373
REMARK 465     ASP A  1374
REMARK 465     GLY A  1406
REMARK 465     THR A  1407
REMARK 465     ASP A  1408
REMARK 465     ASP A  1409
REMARK 465     THR A  1410
REMARK 465     GLN A  1450
REMARK 465     TYR A  1451
REMARK 465     ARG A  1452
REMARK 465     PRO A  1453
REMARK 465     SER A  1454
REMARK 465     GLU A  1483
REMARK 465     GLY A  1484
REMARK 465     ALA A  1485
REMARK 465     SER A  1486
REMARK 465     GLU A  1487
REMARK 465     GLU A  1488
REMARK 465     LEU A  1550
REMARK 465     PHE A  1551
REMARK 465     ALA A  1552
REMARK 465     LYS A  1553
REMARK 465     ASP B  2307
REMARK 465     LEU B  2308
REMARK 465     GLN B  2309
REMARK 465     GLY B  2310
REMARK 465     ASP B  2311
REMARK 465     PRO B  2312
REMARK 465     ARG B  2313
REMARK 465     GLU B  2314
REMARK 465     SER B  2315
REMARK 465     GLN B  2316
REMARK 465     PRO B  2317
REMARK 465     LEU B  2318
REMARK 465     ALA B  2337
REMARK 465     GLU B  2338
REMARK 465     ARG B  2339
REMARK 465     ASN B  2340
REMARK 465     SER B  2369
REMARK 465     GLU B  2370
REMARK 465     GLY B  2371
REMARK 465     GLN B  2372
REMARK 465     LEU B  2373
REMARK 465     ASP B  2374
REMARK 465     GLN B  2375
REMARK 465     LYS B  2376
REMARK 465     THR B  2377
REMARK 465     CYS B  2390
REMARK 465     ILE B  2391
REMARK 465     ALA B  2392
REMARK 465     LYS B  2393
REMARK 465     GLU B  2394
REMARK 465     GLU C  3041
REMARK 465     GLY C  3042
REMARK 465     PHE C  3043
REMARK 465     ALA C  3044
REMARK 465     GLN C  3045
REMARK 465     SER C  3305
REMARK 465     LEU C  3306
REMARK 465     ASP C  3307
REMARK 465     LEU C  3308
REMARK 465     GLN C  3309
REMARK 465     GLY C  3310
REMARK 465     ASP C  3311
REMARK 465     PRO C  3312
REMARK 465     ARG C  3313
REMARK 465     GLU C  3314
REMARK 465     SER C  3315
REMARK 465     GLN C  3316
REMARK 465     PRO C  3317
REMARK 465     LEU C  3318
REMARK 465     GLU C  3338
REMARK 465     ARG C  3339
REMARK 465     ASN C  3340
REMARK 465     PHE C  3341
REMARK 465     MET C  3364
REMARK 465     SER C  3365
REMARK 465     TYR C  3366
REMARK 465     PRO C  3367
REMARK 465     LEU C  3368
REMARK 465     SER C  3369
REMARK 465     GLU C  3370
REMARK 465     GLY C  3371
REMARK 465     GLN C  3372
REMARK 465     LEU C  3373
REMARK 465     ASP C  3374
REMARK 465     GLN C  3375
REMARK 465     LYS C  3376
REMARK 465     THR C  3377
REMARK 465     ALA C  3378
REMARK 465     MET C  3379
REMARK 465     SER C  3380
REMARK 465     LEU C  3381
REMARK 465     LEU C  3382
REMARK 465     TRP C  3383
REMARK 465     LYS C  3384
REMARK 465     SER C  3385
REMARK 465     TYR C  3386
REMARK 465     PRO C  3387
REMARK 465     LEU C  3388
REMARK 465     VAL C  3389
REMARK 465     CYS C  3390
REMARK 465     ILE C  3391
REMARK 465     ALA C  3392
REMARK 465     LYS C  3393
REMARK 465     GLU C  3394
REMARK 465     LEU C  3395
REMARK 465     THR C  3410
REMARK 465     VAL C  3411
REMARK 465     LYS C  3412
REMARK 465     LYS C  3413
REMARK 465     LYS C  3414
REMARK 465     SER C  3456
REMARK 465     SER C  3457
REMARK 465     ASP C  3458
REMARK 465     MET C  3459
REMARK 465     LYS C  3460
REMARK 465     PRO C  3461
REMARK 465     LYS C  3462
REMARK 465     THR C  3463
REMARK 465     LYS C  3482
REMARK 465     GLU C  3483
REMARK 465     GLY C  3484
REMARK 465     ASN C  3549
REMARK 465     LEU C  3550
REMARK 465     PHE C  3551
REMARK 465     ALA C  3552
REMARK 465     LYS C  3553
REMARK 475
REMARK 475 ZERO OCCUPANCY RESIDUES
REMARK 475 THE FOLLOWING RESIDUES WERE MODELED WITH ZERO OCCUPANCY.
REMARK 475 THE LOCATION AND PROPERTIES OF THESE RESIDUES MAY NOT
REMARK 475 BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 475 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE)
REMARK 475   M RES C SSEQI
REMARK 475     ALA A 1337
REMARK 475     GLY A 1356
REMARK 475     LYS A 1376
REMARK 475     GLN A 1534
REMARK 475     PHE B 2303
REMARK 475     HIS B 2342
REMARK 475     LEU B 2368
REMARK 475     TRP B 2383
REMARK 475     LYS B 2384
REMARK 475     TYR B 2386
REMARK 475     LEU B 2404
REMARK 475     GLY B 2405
REMARK 475     GLY B 2406
REMARK 475     SER B 2456
REMARK 475     ALA B 2545
REMARK 475     THR C 3252
REMARK 475     SER C 3253
REMARK 475     PHE C 3303
REMARK 475     GLY C 3405
REMARK 475     GLY C 3511
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     GLU A 1041   CB    CG    CD    OE1   OE2
REMARK 480     GLU A 1106   CB    CG    CD    OE1   OE2
REMARK 480     LYS A 1237   CB    CG    CD    CE    NZ
REMARK 480     LYS A 1258   CB    CG    CD    CE    NZ
REMARK 480     LYS A 1352   CB    CG    CD    CE    NZ
REMARK 480     GLN A 1375   CB    CG    CD    OE1   NE2
REMARK 480     LYS A 1384   CB    CG    CD    CE    NZ
REMARK 480     GLU A 1394   CB    CG    CD    OE1   OE2
REMARK 480     SER A 1457   C     CB    OG
REMARK 480     ASP A 1458   CB    CG    OD1   OD2
REMARK 480     MET A 1459   CB    CG    SD    CE
REMARK 480     LYS A 1462   CB    CG    CD    CE    NZ
REMARK 480     LYS A 1482   CB    CG    CD    CE    NZ
REMARK 480     ARG A 1491   CB    CG    CD    NE    CZ    NH1   NH2
REMARK 480     LYS A 1494   CB    CG    CD    CE    NZ
REMARK 480     LEU A 1514   CB    CG    CD1   CD2
REMARK 480     LYS A 1523   CB    CG    CD    CE    NZ
REMARK 480     ASN A 1532   CB    CG    OD1   ND2
REMARK 480     GLN A 1537   CB    CG    CD    OE1   NE2
REMARK 480     LYS A 1538   CB    CG    CD    CE    NZ
REMARK 480     LYS A 1542   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2036   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2092   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2105   CB    CG    CD    CE    NZ
REMARK 480     GLU B 2106   CB    CG    CD    OE1   OE2
REMARK 480     LYS B 2111   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2130   CB    CG    CD    CE    NZ
REMARK 480     THR B 2252   CB    OG1   CG2
REMARK 480     SER B 2253   CB    OG
REMARK 480     VAL B 2254   CB    CG1   CG2
REMARK 480     LYS B 2258   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2262   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2302   CB    CG    CD    CE    NZ
REMARK 480     LEU B 2306   CB    CG    CD1   CD2
REMARK 480     LYS B 2330   CB    CG    CD    CE    NZ
REMARK 480     GLN B 2336   CB    CG    CD    OE1   NE2
REMARK 480     THR B 2343   CB    OG1   CG2
REMARK 480     LEU B 2358   CB    CG    CD1   CD2
REMARK 480     MET B 2379   CB    CG    SD    CE
REMARK 480     SER B 2380   CA    CB    OG
REMARK 480     LEU B 2395   CB    CG    CD1   CD2
REMARK 480     TYR B 2403   CB    CG    CD1   CD2   CE1   CE2   CZ
REMARK 480     TYR B 2403   OH
REMARK 480     ASP B 2409   CB    CG    OD1   OD2
REMARK 480     LYS B 2412   CB    CG    CD    CE    NZ
REMARK 480     LYS B 2413   CB    CG    CD    CE    NZ
REMARK 480     ASP B 2415   CB    CG    OD1   OD2
REMARK 480     ARG B 2452   CB    CG    CD    NE    CZ    NH1   NH2
REMARK 480     SER B 2457   CB    OG
REMARK 480     ASP B 2458   CB    CG    OD1   OD2
REMARK 480     MET B 2459   CB    CG    SD    CE
REMARK 480     LYS B 2462   CB    CG    CD    CE    NZ
REMARK 480     ASN B 2532   CB    CG    OD1   ND2
REMARK 480     LEU C 3040   CB    CG    CD1   CD2
REMARK 480     PHE C 3076   CB    CG    CD1   CD2   CE1   CE2   CZ
REMARK 480     LYS C 3092   CB    CG    CD    CE    NZ
REMARK 480     LYS C 3105   CB    CG    CD    CE    NZ
REMARK 480     GLU C 3106   CB    CG    CD    OE1   OE2
REMARK 480     LYS C 3129   CB    CG    CD    CE    NZ
REMARK 480     ASP C 3260   CB    CG    OD1   OD2
REMARK 480     LEU C 3264   CB    CG    CD1   CD2
REMARK 480     GLN C 3267   CB    CG    CD    OE1   NE2
REMARK 480     LYS C 3275   CB    CG    CD    CE    NZ
REMARK 480     LYS C 3330   N     CA    C     O
REMARK 480     GLN C 3336   CB    CG    CD    OE1   NE2
REMARK 480     LYS C 3352   CB    CG    CD    CE    NZ
REMARK 480     GLU C 3401   CB    CG    CD    OE1   OE2
REMARK 480     LEU C 3404   CB    CG    CD1   CD2
REMARK 480     ASN C 3436   CB    CG    OD1   ND2
REMARK 480     ASP C 3439   CB    CG    OD1   OD2
REMARK 480     ARG C 3452   CB    CG    CD    NE    CZ    NH1   NH2
REMARK 480     ILE C 3465   CB    CG1   CG2   CD1
REMARK 480     LYS C 3494   CB    CG    CD    CE    NZ
REMARK 480     GLU C 3512   CB    CG    CD    OE1   OE2
REMARK 480     LYS C 3523   CB    CG    CD    CE    NZ
REMARK 480     GLN C 3537   CB    CG    CD    OE1   NE2
REMARK 480     LYS C 3538   CB    CG    CD    CE    NZ
REMARK 480     PHE C 3546   CB    CG    CD1   CD2   CE1   CE2   CZ
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   CG   MET A  1086     O    HOH A    75              1.22
REMARK 500   CG2  THR C  3088     OE2  GLU C  3291              1.87
REMARK 500   CG2  THR B  2088     OE2  GLU B  2291              1.91
REMARK 500   OG   SER B  2245     OH   TYR B  2346              1.97
REMARK 500   OG   SER A  1221     O1   WW2 A   193              1.98
REMARK 500   SG   CYS C  3087     CB   CYS C  3116              2.06
REMARK 500   O    ASN B  2532     OE1  GLN B  2534              2.09
REMARK 500   SG   CYS A  1087     CB   CYS A  1116              2.09
REMARK 500   SG   CYS B  2087     CB   CYS B  2116              2.10
REMARK 500   N    HIS B  2030     O    HOH B    28              2.11
REMARK 500   O    ASN A  1532     OE1  GLN A  1534              2.11
REMARK 500   N    CYS B  2087     O    HOH B    76              2.15
REMARK 500   ND1  HIS B  2030     O    HOH B    28              2.17
REMARK 500   OG   SER B  2221     O1   WW2 B   194              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   OE1  GLN A  1045     O    HOH C    23     2465     1.42
REMARK 500   OG   SER A  1039     NH1  ARG C  3491     2465     1.98
REMARK 500   CD   ARG B  2132     CG   LYS B  2275     1455     2.00
REMARK 500   NE2  GLN A  1095     ND2  ASN B  2238     1655     2.08
REMARK 500   CG2  THR A  1128     OE2  GLU C  3487     2465     2.14
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    PRO A1024   CD    PRO A1024   N      -0.098
REMARK 500    PRO A1046   CD    PRO A1046   N      -0.189
REMARK 500    PRO A1054   CD    PRO A1054   N      -0.102
REMARK 500    PRO A1058   CD    PRO A1058   N      -0.136
REMARK 500    PRO A1059   CD    PRO A1059   N      -0.147
REMARK 500    PRO A1062   CD    PRO A1062   N      -0.157
REMARK 500    PRO A1067   CD    PRO A1067   N      -0.135
REMARK 500    PRO A1068   CD    PRO A1068   N      -0.100
REMARK 500    PRO A1070   CD    PRO A1070   N      -0.109
REMARK 500    PRO A1084   CD    PRO A1084   N      -0.122
REMARK 500    PRO A1085   CB    PRO A1085   CG     -0.356
REMARK 500    PRO A1085   C     PRO A1085   O      -0.150
REMARK 500    CYS A1087   CA    CYS A1087   C       0.182
REMARK 500    CYS A1087   C     CYS A1087   O      -0.118
REMARK 500    PRO A1091   CD    PRO A1091   N      -0.097
REMARK 500    PRO A1109   CD    PRO A1109   N      -0.141
REMARK 500    PRO A1124   CD    PRO A1124   N      -0.117
REMARK 500    PRO A1212   CD    PRO A1212   N      -0.099
REMARK 500    PRO A1263   CD    PRO A1263   N      -0.123
REMARK 500    PRO A1332   CD    PRO A1332   N      -0.133
REMARK 500    PRO A1345   CB    PRO A1345   CG     -0.404
REMARK 500    PRO A1345   CG    PRO A1345   CD     -0.231
REMARK 500    PRO A1345   C     PRO A1345   O      -0.209
REMARK 500    TYR A1346   CB    TYR A1346   CG     -0.104
REMARK 500    TYR A1346   CG    TYR A1346   CD2    -0.157
REMARK 500    TYR A1346   CG    TYR A1346   CD1    -0.198
REMARK 500    TYR A1346   CD1   TYR A1346   CE1    -0.202
REMARK 500    TYR A1346   CE1   TYR A1346   CZ     -0.285
REMARK 500    TYR A1346   CZ    TYR A1346   OH     -0.162
REMARK 500    TYR A1346   CZ    TYR A1346   CE2    -0.256
REMARK 500    TYR A1346   CE2   TYR A1346   CD2    -0.196
REMARK 500    TYR A1346   C     TYR A1346   O      -0.180
REMARK 500    MET A1347   CG    MET A1347   SD     -0.194
REMARK 500    MET A1347   C     MET A1347   O      -0.218
REMARK 500    PRO A1387   CD    PRO A1387   N      -0.108
REMARK 500    PRO A1397   CD    PRO A1397   N      -0.110
REMARK 500    PRO A1429   CD    PRO A1429   N      -0.128
REMARK 500    PRO A1479   CD    PRO A1479   N      -0.141
REMARK 500    PRO A1509   CD    PRO A1509   N      -0.122
REMARK 500    PRO B2024   CD    PRO B2024   N      -0.104
REMARK 500    PRO B2046   CD    PRO B2046   N      -0.161
REMARK 500    PRO B2054   CD    PRO B2054   N      -0.101
REMARK 500    PRO B2058   CD    PRO B2058   N      -0.147
REMARK 500    PRO B2059   CD    PRO B2059   N      -0.137
REMARK 500    PRO B2062   CD    PRO B2062   N      -0.138
REMARK 500    PRO B2067   CD    PRO B2067   N      -0.137
REMARK 500    PRO B2068   CD    PRO B2068   N      -0.099
REMARK 500    PRO B2070   CD    PRO B2070   N      -0.121
REMARK 500    PRO B2073   CD    PRO B2073   N      -0.115
REMARK 500    PRO B2084   CD    PRO B2084   N      -0.139
REMARK 500
REMARK 500 THIS ENTRY HAS     117 BOND DEVIATIONS.
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    PRO A1024   CA  -  N   -  CD  ANGL. DEV. = -12.8 DEGREES
REMARK 500    PRO A1046   CA  -  N   -  CD  ANGL. DEV. = -13.0 DEGREES
REMARK 500    PRO A1054   CA  -  N   -  CD  ANGL. DEV. = -14.0 DEGREES
REMARK 500    PRO A1058   CA  -  N   -  CD  ANGL. DEV. = -13.8 DEGREES
REMARK 500    PRO A1058   C   -  N   -  CD  ANGL. DEV. = -19.1 DEGREES
REMARK 500    PRO A1059   CA  -  N   -  CD  ANGL. DEV. = -14.0 DEGREES
REMARK 500    PRO A1062   CA  -  N   -  CD  ANGL. DEV. = -13.8 DEGREES
REMARK 500    PRO A1067   CA  -  N   -  CD  ANGL. DEV. = -13.3 DEGREES
REMARK 500    PRO A1068   CA  -  N   -  CD  ANGL. DEV. = -14.3 DEGREES
REMARK 500    PRO A1070   CA  -  N   -  CD  ANGL. DEV. = -12.6 DEGREES
REMARK 500    PRO A1084   CA  -  N   -  CD  ANGL. DEV. = -13.4 DEGREES
REMARK 500    CYS A1087   CB  -  CA  -  C   ANGL. DEV. =  15.5 DEGREES
REMARK 500    CYS A1087   O   -  C   -  N   ANGL. DEV. = -11.5 DEGREES
REMARK 500    PRO A1091   CA  -  N   -  CD  ANGL. DEV. = -13.8 DEGREES
REMARK 500    PRO A1109   CA  -  N   -  CD  ANGL. DEV. = -13.1 DEGREES
REMARK 500    PRO A1124   CA  -  N   -  CD  ANGL. DEV. = -13.2 DEGREES
REMARK 500    PRO A1134   CA  -  N   -  CD  ANGL. DEV. = -12.7 DEGREES
REMARK 500    PRO A1212   CA  -  N   -  CD  ANGL. DEV. = -13.6 DEGREES
REMARK 500    PRO A1263   CA  -  N   -  CD  ANGL. DEV. = -13.0 DEGREES
REMARK 500    PRO A1263   C   -  N   -  CD  ANGL. DEV. = -14.1 DEGREES
REMARK 500    PRO A1332   CA  -  N   -  CD  ANGL. DEV. = -13.1 DEGREES
REMARK 500    PRO A1345   CA  -  N   -  CD  ANGL. DEV. =  -8.6 DEGREES
REMARK 500    PRO A1345   N   -  CA  -  CB  ANGL. DEV. =  -8.3 DEGREES
REMARK 500    PRO A1345   CA  -  C   -  O   ANGL. DEV. = -20.9 DEGREES
REMARK 500    PRO A1345   CA  -  C   -  N   ANGL. DEV. =  15.9 DEGREES
REMARK 500    MET A1347   CB  -  CA  -  C   ANGL. DEV. = -15.7 DEGREES
REMARK 500    PRO A1360   CA  -  N   -  CD  ANGL. DEV. = -13.6 DEGREES
REMARK 500    PRO A1360   C   -  N   -  CD  ANGL. DEV. = -16.8 DEGREES
REMARK 500    PRO A1387   CA  -  N   -  CD  ANGL. DEV. = -12.8 DEGREES
REMARK 500    PRO A1387   C   -  N   -  CD  ANGL. DEV. = -15.5 DEGREES
REMARK 500    PRO A1397   CA  -  N   -  CD  ANGL. DEV. = -13.5 DEGREES
REMARK 500    PRO A1397   C   -  N   -  CD  ANGL. DEV. = -17.9 DEGREES
REMARK 500    PRO A1429   CA  -  N   -  CD  ANGL. DEV. = -13.3 DEGREES
REMARK 500    PRO A1443   CA  -  N   -  CD  ANGL. DEV. = -12.6 DEGREES
REMARK 500    PRO A1479   CA  -  N   -  CD  ANGL. DEV. = -13.5 DEGREES
REMARK 500    PRO A1479   C   -  N   -  CD  ANGL. DEV. = -13.6 DEGREES
REMARK 500    PRO A1509   CA  -  N   -  CD  ANGL. DEV. = -13.7 DEGREES
REMARK 500    PRO A1518   CA  -  N   -  CD  ANGL. DEV. = -14.6 DEGREES
REMARK 500    PRO B2024   CA  -  N   -  CD  ANGL. DEV. = -13.2 DEGREES
REMARK 500    PRO B2046   CA  -  N   -  CD  ANGL. DEV. = -13.3 DEGREES
REMARK 500    PRO B2054   CA  -  N   -  CD  ANGL. DEV. = -13.7 DEGREES
REMARK 500    PRO B2058   CA  -  N   -  CD  ANGL. DEV. = -13.4 DEGREES
REMARK 500    PRO B2059   CA  -  N   -  CD  ANGL. DEV. = -13.6 DEGREES
REMARK 500    PRO B2062   CA  -  N   -  CD  ANGL. DEV. = -13.3 DEGREES
REMARK 500    PRO B2067   CA  -  N   -  CD  ANGL. DEV. = -13.2 DEGREES
REMARK 500    PRO B2068   CA  -  N   -  CD  ANGL. DEV. = -14.0 DEGREES
REMARK 500    PRO B2070   CA  -  N   -  CD  ANGL. DEV. = -12.6 DEGREES
REMARK 500    PRO B2073   CA  -  N   -  CD  ANGL. DEV. = -13.0 DEGREES
REMARK 500    PRO B2073   C   -  N   -  CD  ANGL. DEV. = -20.6 DEGREES
REMARK 500    PRO B2084   CA  -  N   -  CD  ANGL. DEV. = -13.6 DEGREES
REMARK 500
REMARK 500 THIS ENTRY HAS     106 ANGLE DEVIATIONS.
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    GLU A1041      127.91    -38.75
REMARK 500    PHE A1076     -133.15     83.44
REMARK 500    CYS A1087      131.92    -38.33
REMARK 500    LEU A1127      -18.11    -45.61
REMARK 500    MET A1145      -35.72   -131.59
REMARK 500    GLU A1183        7.00    -61.18
REMARK 500    SER A1185       75.70    174.90
REMARK 500    SER A1221     -130.40     62.03
REMARK 500    THR A1252       81.08    -65.21
REMARK 500    SER A1253      -39.76    -38.83
REMARK 500    TRP A1357      -55.31   -161.37
REMARK 500    ILE A1396      -61.16    -25.26
REMARK 500    PHE A1426      -52.14   -128.64
REMARK 500    ASN A1521     -140.27   -106.35
REMARK 500    LYS A1540       16.16     47.82
REMARK 500    GLU B2041      126.56    -36.53
REMARK 500    PRO B2059       55.38    -69.77
REMARK 500    PHE B2076     -163.22     68.57
REMARK 500    ALA B2080       60.77   -115.34
REMARK 500    LEU B2127      -18.15    -47.59
REMARK 500    MET B2145      -34.86   -130.96
REMARK 500    GLU B2183        8.08    -61.59
REMARK 500    SER B2185       75.82    175.57
REMARK 500    SER B2221     -129.63     62.35
REMARK 500    THR B2252       82.15    -64.64
REMARK 500    LEU B2304        6.80     57.06
REMARK 500    HIS B2342      109.81    -52.54
REMARK 500    TRP B2357      -54.48   -161.74
REMARK 500    ILE B2396      -60.82    -26.83
REMARK 500    PHE B2426      -53.02   -129.88
REMARK 500    TYR B2451      141.35   -172.74
REMARK 500    ASN B2521     -139.11   -105.22
REMARK 500    LYS B2538       71.09     44.22
REMARK 500    LYS B2540       15.33     47.78
REMARK 500    PRO C3059       54.58    -69.21
REMARK 500    PHE C3076     -162.73     68.71
REMARK 500    ALA C3080       61.72   -113.97
REMARK 500    MET C3086     -174.97    -56.14
REMARK 500    LEU C3127      -17.53    -46.76
REMARK 500    MET C3145      -35.07   -131.13
REMARK 500    GLU C3183        6.91    -61.50
REMARK 500    SER C3185       75.57    176.50
REMARK 500    SER C3221     -129.69     61.71
REMARK 500    THR C3252       81.09    -65.36
REMARK 500    TRP C3357      -54.45   -161.87
REMARK 500    PHE C3426      -52.52   -130.15
REMARK 500    TYR C3451      141.11   -172.65
REMARK 500    ASN C3521     -140.11   -107.04
REMARK 500    LYS C3540       14.88     48.33
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: MAIN CHAIN PLANARITY
REMARK 500
REMARK 500 THE FOLLOWING RESIDUES HAVE A PSEUDO PLANARITY
REMARK 500 TORSION ANGLE, C(I) - CA(I) - N(I+1) - O(I), GREATER
REMARK 500 10.0 DEGREES. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 500 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 500 I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        ANGLE
REMARK 500    CYS B2087         10.53
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    CYS A1087        16.3      L          L   OUTSIDE RANGE
REMARK 500    CYS B2087        22.5      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE WW2 A 193
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE WW2 B 194
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE WW2 C 195
DBREF  3K9B A 1024  1553  UNP    P23141   EST1_HUMAN      24    553
DBREF  3K9B B 2024  2553  UNP    P23141   EST1_HUMAN      24    553
DBREF  3K9B C 3024  3553  UNP    P23141   EST1_HUMAN      24    553
SEQADV 3K9B     A       UNP  P23141    GLN   362 DELETION
SEQADV 3K9B     B       UNP  P23141    GLN   362 DELETION
SEQADV 3K9B     C       UNP  P23141    GLN   362 DELETION
SEQRES   1 A  529  PRO VAL VAL ASP THR VAL HIS GLY LYS VAL LEU GLY LYS
SEQRES   2 A  529  PHE VAL SER LEU GLU GLY PHE ALA GLN PRO VAL ALA ILE
SEQRES   3 A  529  PHE LEU GLY ILE PRO PHE ALA LYS PRO PRO LEU GLY PRO
SEQRES   4 A  529  LEU ARG PHE THR PRO PRO GLN PRO ALA GLU PRO TRP SER
SEQRES   5 A  529  PHE VAL LYS ASN ALA THR SER TYR PRO PRO MET CYS THR
SEQRES   6 A  529  GLN ASP PRO LYS ALA GLY GLN LEU LEU SER GLU LEU PHE
SEQRES   7 A  529  THR ASN ARG LYS GLU ASN ILE PRO LEU LYS LEU SER GLU
SEQRES   8 A  529  ASP CYS LEU TYR LEU ASN ILE TYR THR PRO ALA ASP LEU
SEQRES   9 A  529  THR LYS LYS ASN ARG LEU PRO VAL MET VAL TRP ILE HIS
SEQRES  10 A  529  GLY GLY GLY LEU MET VAL GLY ALA ALA SER THR TYR ASP
SEQRES  11 A  529  GLY LEU ALA LEU ALA ALA HIS GLU ASN VAL VAL VAL VAL
SEQRES  12 A  529  THR ILE GLN TYR ARG LEU GLY ILE TRP GLY PHE PHE SER
SEQRES  13 A  529  THR GLY ASP GLU HIS SER ARG GLY ASN TRP GLY HIS LEU
SEQRES  14 A  529  ASP GLN VAL ALA ALA LEU ARG TRP VAL GLN ASP ASN ILE
SEQRES  15 A  529  ALA SER PHE GLY GLY ASN PRO GLY SER VAL THR ILE PHE
SEQRES  16 A  529  GLY GLU SER ALA GLY GLY GLU SER VAL SER VAL LEU VAL
SEQRES  17 A  529  LEU SER PRO LEU ALA LYS ASN LEU PHE HIS ARG ALA ILE
SEQRES  18 A  529  SER GLU SER GLY VAL ALA LEU THR SER VAL LEU VAL LYS
SEQRES  19 A  529  LYS GLY ASP VAL LYS PRO LEU ALA GLU GLN ILE ALA ILE
SEQRES  20 A  529  THR ALA GLY CYS LYS THR THR THR SER ALA VAL MET VAL
SEQRES  21 A  529  HIS CYS LEU ARG GLN LYS THR GLU GLU GLU LEU LEU GLU
SEQRES  22 A  529  THR THR LEU LYS MET LYS PHE LEU SER LEU ASP LEU GLN
SEQRES  23 A  529  GLY ASP PRO ARG GLU SER GLN PRO LEU LEU GLY THR VAL
SEQRES  24 A  529  ILE ASP GLY MET LEU LEU LEU LYS THR PRO GLU GLU LEU
SEQRES  25 A  529  GLN ALA GLU ARG ASN PHE HIS THR VAL PRO TYR MET VAL
SEQRES  26 A  529  GLY ILE ASN LYS GLN GLU PHE GLY TRP LEU ILE PRO MET
SEQRES  27 A  529  LEU MET SER TYR PRO LEU SER GLU GLY GLN LEU ASP GLN
SEQRES  28 A  529  LYS THR ALA MET SER LEU LEU TRP LYS SER TYR PRO LEU
SEQRES  29 A  529  VAL CYS ILE ALA LYS GLU LEU ILE PRO GLU ALA THR GLU
SEQRES  30 A  529  LYS TYR LEU GLY GLY THR ASP ASP THR VAL LYS LYS LYS
SEQRES  31 A  529  ASP LEU PHE LEU ASP LEU ILE ALA ASP VAL MET PHE GLY
SEQRES  32 A  529  VAL PRO SER VAL ILE VAL ALA ARG ASN HIS ARG ASP ALA
SEQRES  33 A  529  GLY ALA PRO THR TYR MET TYR GLU PHE GLN TYR ARG PRO
SEQRES  34 A  529  SER PHE SER SER ASP MET LYS PRO LYS THR VAL ILE GLY
SEQRES  35 A  529  ASP HIS GLY ASP GLU LEU PHE SER VAL PHE GLY ALA PRO
SEQRES  36 A  529  PHE LEU LYS GLU GLY ALA SER GLU GLU GLU ILE ARG LEU
SEQRES  37 A  529  SER LYS MET VAL MET LYS PHE TRP ALA ASN PHE ALA ARG
SEQRES  38 A  529  ASN GLY ASN PRO ASN GLY GLU GLY LEU PRO HIS TRP PRO
SEQRES  39 A  529  GLU TYR ASN GLN LYS GLU GLY TYR LEU GLN ILE GLY ALA
SEQRES  40 A  529  ASN THR GLN ALA ALA GLN LYS LEU LYS ASP LYS GLU VAL
SEQRES  41 A  529  ALA PHE TRP THR ASN LEU PHE ALA LYS
SEQRES   1 B  529  PRO VAL VAL ASP THR VAL HIS GLY LYS VAL LEU GLY LYS
SEQRES   2 B  529  PHE VAL SER LEU GLU GLY PHE ALA GLN PRO VAL ALA ILE
SEQRES   3 B  529  PHE LEU GLY ILE PRO PHE ALA LYS PRO PRO LEU GLY PRO
SEQRES   4 B  529  LEU ARG PHE THR PRO PRO GLN PRO ALA GLU PRO TRP SER
SEQRES   5 B  529  PHE VAL LYS ASN ALA THR SER TYR PRO PRO MET CYS THR
SEQRES   6 B  529  GLN ASP PRO LYS ALA GLY GLN LEU LEU SER GLU LEU PHE
SEQRES   7 B  529  THR ASN ARG LYS GLU ASN ILE PRO LEU LYS LEU SER GLU
SEQRES   8 B  529  ASP CYS LEU TYR LEU ASN ILE TYR THR PRO ALA ASP LEU
SEQRES   9 B  529  THR LYS LYS ASN ARG LEU PRO VAL MET VAL TRP ILE HIS
SEQRES  10 B  529  GLY GLY GLY LEU MET VAL GLY ALA ALA SER THR TYR ASP
SEQRES  11 B  529  GLY LEU ALA LEU ALA ALA HIS GLU ASN VAL VAL VAL VAL
SEQRES  12 B  529  THR ILE GLN TYR ARG LEU GLY ILE TRP GLY PHE PHE SER
SEQRES  13 B  529  THR GLY ASP GLU HIS SER ARG GLY ASN TRP GLY HIS LEU
SEQRES  14 B  529  ASP GLN VAL ALA ALA LEU ARG TRP VAL GLN ASP ASN ILE
SEQRES  15 B  529  ALA SER PHE GLY GLY ASN PRO GLY SER VAL THR ILE PHE
SEQRES  16 B  529  GLY GLU SER ALA GLY GLY GLU SER VAL SER VAL LEU VAL
SEQRES  17 B  529  LEU SER PRO LEU ALA LYS ASN LEU PHE HIS ARG ALA ILE
SEQRES  18 B  529  SER GLU SER GLY VAL ALA LEU THR SER VAL LEU VAL LYS
SEQRES  19 B  529  LYS GLY ASP VAL LYS PRO LEU ALA GLU GLN ILE ALA ILE
SEQRES  20 B  529  THR ALA GLY CYS LYS THR THR THR SER ALA VAL MET VAL
SEQRES  21 B  529  HIS CYS LEU ARG GLN LYS THR GLU GLU GLU LEU LEU GLU
SEQRES  22 B  529  THR THR LEU LYS MET LYS PHE LEU SER LEU ASP LEU GLN
SEQRES  23 B  529  GLY ASP PRO ARG GLU SER GLN PRO LEU LEU GLY THR VAL
SEQRES  24 B  529  ILE ASP GLY MET LEU LEU LEU LYS THR PRO GLU GLU LEU
SEQRES  25 B  529  GLN ALA GLU ARG ASN PHE HIS THR VAL PRO TYR MET VAL
SEQRES  26 B  529  GLY ILE ASN LYS GLN GLU PHE GLY TRP LEU ILE PRO MET
SEQRES  27 B  529  LEU MET SER TYR PRO LEU SER GLU GLY GLN LEU ASP GLN
SEQRES  28 B  529  LYS THR ALA MET SER LEU LEU TRP LYS SER TYR PRO LEU
SEQRES  29 B  529  VAL CYS ILE ALA LYS GLU LEU ILE PRO GLU ALA THR GLU
SEQRES  30 B  529  LYS TYR LEU GLY GLY THR ASP ASP THR VAL LYS LYS LYS
SEQRES  31 B  529  ASP LEU PHE LEU ASP LEU ILE ALA ASP VAL MET PHE GLY
SEQRES  32 B  529  VAL PRO SER VAL ILE VAL ALA ARG ASN HIS ARG ASP ALA
SEQRES  33 B  529  GLY ALA PRO THR TYR MET TYR GLU PHE GLN TYR ARG PRO
SEQRES  34 B  529  SER PHE SER SER ASP MET LYS PRO LYS THR VAL ILE GLY
SEQRES  35 B  529  ASP HIS GLY ASP GLU LEU PHE SER VAL PHE GLY ALA PRO
SEQRES  36 B  529  PHE LEU LYS GLU GLY ALA SER GLU GLU GLU ILE ARG LEU
SEQRES  37 B  529  SER LYS MET VAL MET LYS PHE TRP ALA ASN PHE ALA ARG
SEQRES  38 B  529  ASN GLY ASN PRO ASN GLY GLU GLY LEU PRO HIS TRP PRO
SEQRES  39 B  529  GLU TYR ASN GLN LYS GLU GLY TYR LEU GLN ILE GLY ALA
SEQRES  40 B  529  ASN THR GLN ALA ALA GLN LYS LEU LYS ASP LYS GLU VAL
SEQRES  41 B  529  ALA PHE TRP THR ASN LEU PHE ALA LYS
SEQRES   1 C  529  PRO VAL VAL ASP THR VAL HIS GLY LYS VAL LEU GLY LYS
SEQRES   2 C  529  PHE VAL SER LEU GLU GLY PHE ALA GLN PRO VAL ALA ILE
SEQRES   3 C  529  PHE LEU GLY ILE PRO PHE ALA LYS PRO PRO LEU GLY PRO
SEQRES   4 C  529  LEU ARG PHE THR PRO PRO GLN PRO ALA GLU PRO TRP SER
SEQRES   5 C  529  PHE VAL LYS ASN ALA THR SER TYR PRO PRO MET CYS THR
SEQRES   6 C  529  GLN ASP PRO LYS ALA GLY GLN LEU LEU SER GLU LEU PHE
SEQRES   7 C  529  THR ASN ARG LYS GLU ASN ILE PRO LEU LYS LEU SER GLU
SEQRES   8 C  529  ASP CYS LEU TYR LEU ASN ILE TYR THR PRO ALA ASP LEU
SEQRES   9 C  529  THR LYS LYS ASN ARG LEU PRO VAL MET VAL TRP ILE HIS
SEQRES  10 C  529  GLY GLY GLY LEU MET VAL GLY ALA ALA SER THR TYR ASP
SEQRES  11 C  529  GLY LEU ALA LEU ALA ALA HIS GLU ASN VAL VAL VAL VAL
SEQRES  12 C  529  THR ILE GLN TYR ARG LEU GLY ILE TRP GLY PHE PHE SER
SEQRES  13 C  529  THR GLY ASP GLU HIS SER ARG GLY ASN TRP GLY HIS LEU
SEQRES  14 C  529  ASP GLN VAL ALA ALA LEU ARG TRP VAL GLN ASP ASN ILE
SEQRES  15 C  529  ALA SER PHE GLY GLY ASN PRO GLY SER VAL THR ILE PHE
SEQRES  16 C  529  GLY GLU SER ALA GLY GLY GLU SER VAL SER VAL LEU VAL
SEQRES  17 C  529  LEU SER PRO LEU ALA LYS ASN LEU PHE HIS ARG ALA ILE
SEQRES  18 C  529  SER GLU SER GLY VAL ALA LEU THR SER VAL LEU VAL LYS
SEQRES  19 C  529  LYS GLY ASP VAL LYS PRO LEU ALA GLU GLN ILE ALA ILE
SEQRES  20 C  529  THR ALA GLY CYS LYS THR THR THR SER ALA VAL MET VAL
SEQRES  21 C  529  HIS CYS LEU ARG GLN LYS THR GLU GLU GLU LEU LEU GLU
SEQRES  22 C  529  THR THR LEU LYS MET LYS PHE LEU SER LEU ASP LEU GLN
SEQRES  23 C  529  GLY ASP PRO ARG GLU SER GLN PRO LEU LEU GLY THR VAL
SEQRES  24 C  529  ILE ASP GLY MET LEU LEU LEU LYS THR PRO GLU GLU LEU
SEQRES  25 C  529  GLN ALA GLU ARG ASN PHE HIS THR VAL PRO TYR MET VAL
SEQRES  26 C  529  GLY ILE ASN LYS GLN GLU PHE GLY TRP LEU ILE PRO MET
SEQRES  27 C  529  LEU MET SER TYR PRO LEU SER GLU GLY GLN LEU ASP GLN
SEQRES  28 C  529  LYS THR ALA MET SER LEU LEU TRP LYS SER TYR PRO LEU
SEQRES  29 C  529  VAL CYS ILE ALA LYS GLU LEU ILE PRO GLU ALA THR GLU
SEQRES  30 C  529  LYS TYR LEU GLY GLY THR ASP ASP THR VAL LYS LYS LYS
SEQRES  31 C  529  ASP LEU PHE LEU ASP LEU ILE ALA ASP VAL MET PHE GLY
SEQRES  32 C  529  VAL PRO SER VAL ILE VAL ALA ARG ASN HIS ARG ASP ALA
SEQRES  33 C  529  GLY ALA PRO THR TYR MET TYR GLU PHE GLN TYR ARG PRO
SEQRES  34 C  529  SER PHE SER SER ASP MET LYS PRO LYS THR VAL ILE GLY
SEQRES  35 C  529  ASP HIS GLY ASP GLU LEU PHE SER VAL PHE GLY ALA PRO
SEQRES  36 C  529  PHE LEU LYS GLU GLY ALA SER GLU GLU GLU ILE ARG LEU
SEQRES  37 C  529  SER LYS MET VAL MET LYS PHE TRP ALA ASN PHE ALA ARG
SEQRES  38 C  529  ASN GLY ASN PRO ASN GLY GLU GLY LEU PRO HIS TRP PRO
SEQRES  39 C  529  GLU TYR ASN GLN LYS GLU GLY TYR LEU GLN ILE GLY ALA
SEQRES  40 C  529  ASN THR GLN ALA ALA GLN LYS LEU LYS ASP LYS GLU VAL
SEQRES  41 C  529  ALA PHE TRP THR ASN LEU PHE ALA LYS
HET    WW2  A 193      10
HET    WW2  B 194      10
HET    WW2  C 195      10
HETNAM     WW2 CYCLOHEXYL (S)-METHYLPHOSPHONOFLUORIDOATE
HETSYN     WW2 CYCLOSARIN
FORMUL   4  WW2    3(C7 H14 F O2 P)
FORMUL   7  HOH   *66(H2 O)
HELIX    1   1 LEU A 1060  ARG A 1064  5                                   5
HELIX    2   2 ASP A 1090  THR A 1102  1                                  13
HELIX    3   3 ALA A 1148  TYR A 1152  5                                   5
HELIX    4   4 GLY A 1154  ASN A 1162  1                                   9
HELIX    5   5 LEU A 1172  PHE A 1178  1                                   7
HELIX    6   6 ASN A 1188  ILE A 1205  1                                  18
HELIX    7   7 ALA A 1206  PHE A 1208  5                                   3
HELIX    8   8 SER A 1221  SER A 1233  1                                  13
HELIX    9   9 PRO A 1234  LYS A 1237  5                                   4
HELIX   10  10 THR A 1252  VAL A 1254  5                                   3
HELIX   11  11 VAL A 1261  ALA A 1272  1                                  12
HELIX   12  12 THR A 1278  LYS A 1289  1                                  12
HELIX   13  13 THR A 1290  LYS A 1302  1                                  13
HELIX   14  14 THR A 1331  GLU A 1338  1                                   8
HELIX   15  15 TRP A 1357  MET A 1364  1                                   7
HELIX   16  16 GLN A 1375  SER A 1385  1                                  11
HELIX   17  17 SER A 1385  CYS A 1390  1                                   6
HELIX   18  18 LEU A 1395  GLY A 1405  1                                  11
HELIX   19  19 VAL A 1411  PHE A 1426  1                                  16
HELIX   20  20 PHE A 1426  ALA A 1440  1                                  15
HELIX   21  21 GLU A 1471  PHE A 1476  1                                   6
HELIX   22  22 GLY A 1477  LEU A 1481  5                                   5
HELIX   23  23 GLU A 1489  GLY A 1507  1                                  19
HELIX   24  24 LYS A 1540  ASN A 1549  1                                  10
HELIX   25  25 LEU B 2060  ARG B 2064  5                                   5
HELIX   26  26 ASP B 2090  THR B 2102  1                                  13
HELIX   27  27 ALA B 2148  TYR B 2152  5                                   5
HELIX   28  28 GLY B 2154  ASN B 2162  1                                   9
HELIX   29  29 LEU B 2172  PHE B 2178  1                                   7
HELIX   30  30 ASN B 2188  ILE B 2205  1                                  18
HELIX   31  31 ALA B 2206  PHE B 2208  5                                   3
HELIX   32  32 SER B 2221  SER B 2233  1                                  13
HELIX   33  33 PRO B 2234  LYS B 2237  5                                   4
HELIX   34  34 THR B 2252  VAL B 2254  5                                   3
HELIX   35  35 VAL B 2261  ALA B 2272  1                                  12
HELIX   36  36 THR B 2278  LYS B 2289  1                                  12
HELIX   37  37 THR B 2290  LYS B 2302  1                                  13
HELIX   38  38 THR B 2331  GLN B 2336  1                                   6
HELIX   39  39 TRP B 2357  MET B 2364  1                                   7
HELIX   40  40 ALA B 2378  SER B 2385  1                                   8
HELIX   41  41 LEU B 2395  GLY B 2405  1                                  11
HELIX   42  42 ASP B 2409  PHE B 2426  1                                  18
HELIX   43  43 PHE B 2426  ALA B 2440  1                                  15
HELIX   44  44 GLU B 2471  PHE B 2476  1                                   6
HELIX   45  45 GLY B 2477  LEU B 2481  5                                   5
HELIX   46  46 SER B 2486  GLY B 2507  1                                  22
HELIX   47  47 LYS B 2540  PHE B 2551  1                                  12
HELIX   48  48 LEU C 3060  ARG C 3064  5                                   5
HELIX   49  49 ASP C 3090  THR C 3102  1                                  13
HELIX   50  50 ALA C 3148  TYR C 3152  5                                   5
HELIX   51  51 GLY C 3154  ASN C 3162  1                                   9
HELIX   52  52 LEU C 3172  PHE C 3178  1                                   7
HELIX   53  53 ASN C 3188  ILE C 3205  1                                  18
HELIX   54  54 ALA C 3206  PHE C 3208  5                                   3
HELIX   55  55 SER C 3221  SER C 3233  1                                  13
HELIX   56  56 PRO C 3234  LYS C 3237  5                                   4
HELIX   57  57 THR C 3252  VAL C 3254  5                                   3
HELIX   58  58 VAL C 3261  ALA C 3272  1                                  12
HELIX   59  59 THR C 3278  LYS C 3289  1                                  12
HELIX   60  60 THR C 3290  LYS C 3302  1                                  13
HELIX   61  61 THR C 3331  ALA C 3337  1                                   7
HELIX   62  62 TRP C 3357  LEU C 3363  1                                   6
HELIX   63  63 ILE C 3396  GLY C 3405  1                                  10
HELIX   64  64 ASP C 3415  PHE C 3426  1                                  12
HELIX   65  65 PHE C 3426  ALA C 3440  1                                  15
HELIX   66  66 GLU C 3471  PHE C 3476  1                                   6
HELIX   67  67 GLY C 3477  LEU C 3481  5                                   5
HELIX   68  68 SER C 3486  GLY C 3507  1                                  22
HELIX   69  69 LYS C 3540  THR C 3548  1                                   9
SHEET    1   A 3 VAL A1025  THR A1028  0
SHEET    2   A 3 GLY A1031  LEU A1034 -1  O  VAL A1033   N  VAL A1026
SHEET    3   A 3 VAL A1077  ASN A1079  1  O  LYS A1078   N  LYS A1032
SHEET    1   B11 LYS A1036  VAL A1038  0
SHEET    2   B11 VAL A1047  PRO A1054 -1  O  VAL A1047   N  VAL A1038
SHEET    3   B11 TYR A1118  THR A1123 -1  O  ILE A1121   N  PHE A1050
SHEET    4   B11 VAL A1164  ILE A1168 -1  O  VAL A1165   N  TYR A1122
SHEET    5   B11 LEU A1133  ILE A1139  1  N  TRP A1138   O  VAL A1166
SHEET    6   B11 GLY A1210  GLU A1220  1  O  THR A1216   N  VAL A1135
SHEET    7   B11 ARG A1242  GLU A1246  1  O  GLU A1246   N  GLY A1219
SHEET    8   B11 TYR A1346  ASN A1351  1  O  MET A1347   N  ALA A1243
SHEET    9   B11 THR A1444  PHE A1449  1  O  TYR A1445   N  VAL A1348
SHEET   10   B11 GLY A1525  GLN A1528  1  O  LEU A1527   N  GLU A1448
SHEET   11   B11 GLN A1534  GLN A1537 -1  O  ALA A1536   N  TYR A1526
SHEET    1   C 2 MET A1086  CYS A1087  0
SHEET    2   C 2 LEU A1112  SER A1113  1  O  SER A1113   N  MET A1086
SHEET    1   D 2 VAL A1256  LYS A1257  0
SHEET    2   D 2 THR A1321  VAL A1322  1  O  THR A1321   N  LYS A1257
SHEET    1   E 3 VAL B2025  ASP B2027  0
SHEET    2   E 3 LYS B2032  LEU B2034 -1  O  VAL B2033   N  VAL B2026
SHEET    3   E 3 VAL B2077  ASN B2079  1  O  LYS B2078   N  LYS B2032
SHEET    1   F11 LYS B2036  VAL B2038  0
SHEET    2   F11 VAL B2047  PRO B2054 -1  O  VAL B2047   N  VAL B2038
SHEET    3   F11 TYR B2118  THR B2123 -1  O  ILE B2121   N  PHE B2050
SHEET    4   F11 VAL B2164  ILE B2168 -1  O  VAL B2165   N  TYR B2122
SHEET    5   F11 LEU B2133  ILE B2139  1  N  TRP B2138   O  VAL B2166
SHEET    6   F11 GLY B2210  GLU B2220  1  O  THR B2216   N  VAL B2135
SHEET    7   F11 ARG B2242  GLU B2246  1  O  GLU B2246   N  GLY B2219
SHEET    8   F11 TYR B2346  ASN B2351  1  O  GLY B2349   N  SER B2245
SHEET    9   F11 THR B2444  GLN B2450  1  O  TYR B2445   N  VAL B2348
SHEET   10   F11 GLY B2525  GLY B2530  1  O  ILE B2529   N  GLN B2450
SHEET   11   F11 GLN B2534  GLN B2537 -1  O  ALA B2536   N  TYR B2526
SHEET    1   G 2 MET B2086  CYS B2087  0
SHEET    2   G 2 LEU B2112  SER B2113  1  O  SER B2113   N  MET B2086
SHEET    1   H 2 VAL B2256  LYS B2257  0
SHEET    2   H 2 THR B2321  VAL B2322  1  O  THR B2321   N  LYS B2257
SHEET    1   I 3 VAL C3025  THR C3028  0
SHEET    2   I 3 GLY C3031  LEU C3034 -1  O  VAL C3033   N  VAL C3026
SHEET    3   I 3 VAL C3077  ASN C3079  1  O  LYS C3078   N  LYS C3032
SHEET    1   J11 LYS C3036  VAL C3038  0
SHEET    2   J11 VAL C3047  PRO C3054 -1  O  VAL C3047   N  VAL C3038
SHEET    3   J11 TYR C3118  THR C3123 -1  O  ILE C3121   N  PHE C3050
SHEET    4   J11 VAL C3164  ILE C3168 -1  O  VAL C3165   N  TYR C3122
SHEET    5   J11 LEU C3133  ILE C3139  1  N  TRP C3138   O  VAL C3166
SHEET    6   J11 GLY C3210  GLU C3220  1  O  THR C3216   N  VAL C3135
SHEET    7   J11 ARG C3242  GLU C3246  1  O  GLU C3246   N  GLY C3219
SHEET    8   J11 TYR C3346  ASN C3351  1  O  MET C3347   N  ALA C3243
SHEET    9   J11 THR C3444  GLN C3450  1  O  TYR C3445   N  VAL C3348
SHEET   10   J11 GLY C3525  GLY C3530  1  O  ILE C3529   N  GLN C3450
SHEET   11   J11 GLN C3534  GLN C3537 -1  O  ALA C3536   N  TYR C3526
SHEET    1   K 2 MET C3086  CYS C3087  0
SHEET    2   K 2 LEU C3112  SER C3113  1  O  SER C3113   N  MET C3086
SHEET    1   L 2 VAL C3256  LYS C3257  0
SHEET    2   L 2 THR C3321  VAL C3322  1  O  THR C3321   N  LYS C3257
SSBOND   1 CYS A 1087    CYS A 1116                          1555   1555  1.15
SSBOND   2 CYS A 1274    CYS A 1285                          1555   1555  2.04
SSBOND   3 CYS B 2087    CYS B 2116                          1555   1555  0.87
SSBOND   4 CYS B 2274    CYS B 2285                          1555   1555  2.05
SSBOND   5 CYS C 3087    CYS C 3116                          1555   1555  0.83
SSBOND   6 CYS C 3274    CYS C 3285                          1555   1555  2.05
LINK         OG  SER A1221                 P1  WW2 A 193     1555   1555  1.45
LINK         OG  SER B2221                 P1  WW2 B 194     1555   1555  1.48
LINK         OG  SER C3221                 P1  WW2 C 195     1555   1555  1.43
SITE     1 AC1  6 GLY A1141  GLY A1142  GLY A1143  SER A1221
SITE     2 AC1  6 ALA A1222  LEU A1363
SITE     1 AC2  7 GLY B2141  GLY B2142  GLY B2143  SER B2221
SITE     2 AC2  7 ALA B2222  LEU B2363  HIS B2468
SITE     1 AC3  6 GLY C3141  GLY C3142  GLY C3143  SER C3221
SITE     2 AC3  6 ALA C3222  HIS C3468
CRYST1   55.610  179.880  200.060  90.00  90.00  90.00 P 21 21 21   12
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017982  0.000000  0.000000        0.00000
SCALE2      0.000000  0.005559  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004999        0.00000
TER    3769      ASN A1549
TER    7648      LYS B2553
TER   11158      THR C3548
MASTER      806    0    3   69   54    0    6    611251    3   45  123
END