longtext: 3LIP-pdb

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HEADER    HYDROLASE                               18-APR-97   3LIP
TITLE     OPEN CONFORMATION OF PSEUDOMONAS CEPACIA LIPASE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: TRIACYL-GLYCEROL-HYDROLASE;
COMPND   3 CHAIN: NULL;
COMPND   4 SYNONYM: LIPASE;
COMPND   5 EC: 3.1.1.3;
COMPND   6 BIOLOGICAL_UNIT: MONOMER
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOMONAS CEPACIA
SOURCE   3 (BURKHOLDERIA CEPACIA)
KEYWDS    HYDROLASE, LIPASE, PSEUDOMONADACEAE, CATALYTIC TRIAD,
KEYWDS   2 OXYANION, OPEN, WITHOUT INHIBITOR
EXPDTA    X-RAY DIFFRACTION
AUTHOR    D.A.LANG,D.SCHOMBURG
REVDAT   1   16-JUN-97 3LIP    0
JRNL        AUTH   J.D.SCHRAG,Y.LI,M.CYGLER,D.LANG,T.BURGDORF,
JRNL        AUTH 2 H.J.HECHT,R.SCHMID,D.SCHOMBURG,T.J.RYDEL,
JRNL        AUTH 3 J.D.OLIVER,L.C.STRICKLAND,C.M.DUNAWAY,S.B.LARSON,
JRNL        AUTH 4 J.DAY,A.MCPHERSON
JRNL        TITL   THE OPEN CONFORMATION OF A PSEUDOMONAS LIPASE
JRNL        REF    STRUCTURE (LONDON)            V.   5   187 1997
JRNL        REFN   ASTM STRUE6  UK ISSN 0969-2126                 2005
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   U.BORNSCHEUER,O.W.REIF,R.LAUSCH,R.FREITAG,
REMARK   1  AUTH 2 T.SCHEPER,F.N.KOLISIS,U.MENGE
REMARK   1  TITL   LIPASE OF PSEUDOMONAS CEPACIA FOR BIOTECHNOLOGICAL
REMARK   1  TITL 2 PURPOSES: PURIFICATION, CRYSTALLIZATION AND
REMARK   1  TITL 3 CHARACTERIZATION
REMARK   1  REF    BIOCHIM.BIOPHYS.ACTA          V.1201    55 1994
REMARK   1  REFN   ASTM BBACAQ  NE ISSN 0006-3002                 0113
REMARK   1 REFERENCE 2
REMARK   1  AUTH   D.A.LANG,B.HOFMANN,T.BURGDORF,L.HAALCK,H.-J.HECHT,
REMARK   1  AUTH 2 F.SPENER,R.D.SCHMID,D.SCHOMBURG
REMARK   1  TITL   STRUCTURE ELUCIDATION OF TWO LIPASES OF THE
REMARK   1  TITL 2 PSEUDOMONAS FAMILY
REMARK   1  REF    CLOSING MEETING OF THE EC               26 1994
REMARK   1  REF  2 BRIDGE LIPASE T-PROJECT
REMARK   1  REF  3 "CHARACTERIZATION OF LIPASES
REMARK   1  REF  4 FOR INDUSTRIAL APPLICATIONS:
REMARK   1  REF  5 3D STRUCTURE AND CATALYTIC
REMARK   1  REF  6 MECHANISM", PROGRAMME,
REMARK   1  REF  7 ABSTRACTS & PARTICIPANTS,
REMARK   1  REF  8 INTERNATIONAL WORKSHOP,
REMARK   1  REF  9 BENDOR ISLAND, BANDOL,
REMARK   1  REF 10 FRANCE, SEPTEMBER 14-17,
REMARK   1  REF 11 1994
REMARK   1  REFN   ASTM         FR ISBN                           2138
REMARK   2
REMARK   2 RESOLUTION. 2.0  ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PROLSQ
REMARK   3   AUTHORS     : KONNERT,HENDRICKSON
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.0
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 8.0
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.0
REMARK   3   COMPLETENESS FOR RANGE        (%) : 83.
REMARK   3   NUMBER OF REFLECTIONS             : 17424
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : NULL
REMARK   3   R VALUE     (WORKING + TEST SET) : NULL
REMARK   3   R VALUE            (WORKING SET) : 0.188
REMARK   3   FREE R VALUE                     : NULL
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : NULL
REMARK   3
REMARK   3  FIT/AGREEMENT OF MODEL WITH ALL DATA.
REMARK   3   R VALUE   (WORKING + TEST SET, NO CUTOFF) : NULL
REMARK   3   R VALUE          (WORKING SET, NO CUTOFF) : 0.188
REMARK   3   FREE R VALUE                  (NO CUTOFF) : NULL
REMARK   3   FREE R VALUE TEST SET SIZE (%, NO CUTOFF) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT   (NO CUTOFF) : NULL
REMARK   3   TOTAL NUMBER OF REFLECTIONS   (NO CUTOFF) : 17424
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2338
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 1
REMARK   3   SOLVENT ATOMS            : 193
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 12.95
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.2
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 8.0
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   DISTANCE RESTRAINTS.                    RMS    SIGMA
REMARK   3    BOND LENGTH                     (A) : 0.02  ; 0.02
REMARK   3    ANGLE DISTANCE                  (A) : 0.039 ; 0.040
REMARK   3    INTRAPLANAR 1-4 DISTANCE        (A) : 0.052 ; 0.050
REMARK   3    H-BOND OR METAL COORDINATION    (A) : NULL  ; 0.050
REMARK   3
REMARK   3   PLANE RESTRAINT                  (A) : 0.015 ; 0.020
REMARK   3   CHIRAL-CENTER RESTRAINT       (A**3) : 0.086 ; 0.08
REMARK   3
REMARK   3   NON-BONDED CONTACT RESTRAINTS.
REMARK   3    SINGLE TORSION                  (A) : 0.174 ; 0.30
REMARK   3    MULTIPLE TORSION                (A) : 0.138 ; 0.30
REMARK   3    H-BOND (X...Y)                  (A) : NULL  ; 0.30
REMARK   3    H-BOND (X-H...Y)                (A) : NULL  ; 0.30
REMARK   3
REMARK   3   CONFORMATIONAL TORSION ANGLE RESTRAINTS.
REMARK   3    SPECIFIED                 (DEGREES) : 15.0  ; NULL
REMARK   3    PLANAR                    (DEGREES) : 2.387 ; 3.0
REMARK   3    STAGGERED                 (DEGREES) : 12.031; 15.0
REMARK   3    TRANSVERSE                (DEGREES) : 23.919; 15.0
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.512 ; 1.000
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 2.406 ; 1.500
REMARK   3   SIDE-CHAIN BOND              (A**2) : 1.474 ; 1.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 2.02  ; 1.50
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3LIP COMPLIES WITH FORMAT V. 2.2, 16-DEC-1996
REMARK   6
REMARK   6 DENSITY FOR RESIDUES 217 - 223 IS FRAGMENTED. THEREFORE
REMARK   6 THIS REGION WAS MODELED STEREOCHEMICALLY.
REMARK   7
REMARK   7 THE ELECTRON DENSITY FOR THE RESIDUES THR 18, SER 87, THR
REMARK   7 217 AND LEU 234 IS WELL DEFINED IN SPITE OF DIHEDRAL
REMARK   7 ANGLES OF THESE RESIDUES LYING OUTSIDE THEIR EXPECTED
REMARK   7 RANGE.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : MAY-1994
REMARK 200  TEMPERATURE           (KELVIN) : 283
REMARK 200  PH                             : 8.7
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : EMBL/DESY, HAMBURG
REMARK 200  BEAMLINE                       : BW6
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : GRAPHITE
REMARK 200  OPTICS                         : MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 17688
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.0
REMARK 200  RESOLUTION RANGE LOW       (A) : 18.7
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 83.3
REMARK 200  DATA REDUNDANCY                : 2.0
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : 0.095
REMARK 200  <I/SIGMA(I)> FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.0
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.05
REMARK 200  COMPLETENESS FOR SHELL     (%) : 83.1
REMARK 200  DATA REDUNDANCY IN SHELL       : 1.63
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : 0.183
REMARK 200  <I/SIGMA(I)> FOR SHELL         : NULL
REMARK 200
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR
REMARK 200                                         REPLACEMENT
REMARK 200 SOFTWARE USED: AMORE
REMARK 200 STARTING MODEL: PDB ENTRY 1CVL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 48.
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.38
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 29-30 % N-PROPANOL, 0.1 M TRIS
REMARK 280 HCL BUFFER, PH 8.4-8.7, AT 12 DEGREES IN FOUR WEEKS,
REMARK 280 PROTEIN CONCENTRATION 22 MG/ML
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y,-Z
REMARK 290       3555   1/2+X,1/2+Y,Z
REMARK 290       4555   1/2-X,1/2+Y,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       45.64959
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       23.64961
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       45.64959
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       23.64961
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  SOME OF THESE MAY BE ATOMS
REMARK 500 LOCATED ON SPECIAL POSITIONS IN THE CELL.  ATOMS WITH
REMARK 500 NON-BLANK ALTERNATE LOCATION INDICATORS ARE NOT INCLUDED
REMARK 500 IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH     533     O    HOH     583     4545     0.27
REMARK 500   O    HOH     583     O    HOH     533     4555     0.27
REMARK 500   O    HOH     579     O    HOH     478     4556     0.53
REMARK 500   O    HOH     478     O    HOH     579     4546     0.53
REMARK 500   O    HOH     547     O    HOH     586     4545     1.65
REMARK 500   O    HOH     586     O    HOH     547     4555     1.65
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: ACT
REMARK 800 SITE_DESCRIPTION: CATALYTIC TRIAD RESIDUES.
REMARK 800
REMARK 800 SITE_IDENTIFIER: OXY
REMARK 800 SITE_DESCRIPTION: OXYANION HOLE.
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 3LIP       SWS     P22088       1 -    44 NOT IN ATOMS LIST
REMARK 999
REMARK 999 REFERENCE: THIS SEQUENCE MATCHES THAT REFERENCED IN
REMARK 999 NAKANISHI ET AL. (1991): CLONING, SEQUENCING, REGULATION
REMARK 999 OF THE LIPASE GENE FROM PSEUDOMONAS SP. M 12-33. IN
REMARK 999 LIPASES: STRUCTURE, MECHANISM, GENETIC ENGINEERING. GBF
REMARK 999 MONOGRAPHS NO. 16 (ALBERGHINA, SCHMID, VERGER EDS.) VCH,
REMARK 999 WEINHEIM, PP. 263-266.
DBREF  3LIP      1   320  SWS    P22088   LIP_BURCE       45    364
SEQADV 3LIP ASP      2  SWS  P22088    ALA    46 CONFLICT
SEQADV 3LIP ASN      3  SWS  P22088    GLY    47 CONFLICT
SEQADV 3LIP THR     18  SWS  P22088    SER    62 CONFLICT
SEQADV 3LIP ARG     40  SWS  P22088    ASN    84 CONFLICT
SEQADV 3LIP THR     92  SWS  P22088    SER   136 CONFLICT
SEQADV 3LIP GLY    125  SWS  P22088    ASP   169 CONFLICT
SEQADV 3LIP THR    137  SWS  P22088    SER   181 CONFLICT
SEQADV 3LIP ASN    154  SWS  P22088    HIS   198 CONFLICT
SEQADV 3LIP LYS    165  SWS  P22088    GLN   209 CONFLICT
SEQADV 3LIP GLN    171  SWS  P22088    ARG   215 CONFLICT
SEQADV 3LIP ILE    218  SWS  P22088    LEU   262 CONFLICT
SEQADV 3LIP ILE    232  SWS  P22088    LEU   276 CONFLICT
SEQADV 3LIP ALA    240  SWS  P22088    VAL   284 CONFLICT
SEQADV 3LIP PRO    243  SWS  P22088    LEU   287 CONFLICT
SEQADV 3LIP VAL    256  SWS  P22088    ILE   300 CONFLICT
SEQADV 3LIP VAL    266  SWS  P22088    LEU   310 CONFLICT
SEQADV 3LIP GLN    276  SWS  P22088    LYS   320 CONFLICT
SEQADV 3LIP ASN    300  SWS  P22088    TYR   344 CONFLICT
SEQRES   1    320  ALA ASP ASN TYR ALA ALA THR ARG TYR PRO ILE ILE LEU
SEQRES   2    320  VAL HIS GLY LEU THR GLY THR ASP LYS TYR ALA GLY VAL
SEQRES   3    320  LEU GLU TYR TRP TYR GLY ILE GLN GLU ASP LEU GLN GLN
SEQRES   4    320  ARG GLY ALA THR VAL TYR VAL ALA ASN LEU SER GLY PHE
SEQRES   5    320  GLN SER ASP ASP GLY PRO ASN GLY ARG GLY GLU GLN LEU
SEQRES   6    320  LEU ALA TYR VAL LYS THR VAL LEU ALA ALA THR GLY ALA
SEQRES   7    320  THR LYS VAL ASN LEU VAL GLY HIS SER GLN GLY GLY LEU
SEQRES   8    320  THR SER ARG TYR VAL ALA ALA VAL ALA PRO ASP LEU VAL
SEQRES   9    320  ALA SER VAL THR THR ILE GLY THR PRO HIS ARG GLY SER
SEQRES  10    320  GLU PHE ALA ASP PHE VAL GLN GLY VAL LEU ALA TYR ASP
SEQRES  11    320  PRO THR GLY LEU SER SER THR VAL ILE ALA ALA PHE VAL
SEQRES  12    320  ASN VAL PHE GLY ILE LEU THR SER SER SER ASN ASN THR
SEQRES  13    320  ASN GLN ASP ALA LEU ALA ALA LEU LYS THR LEU THR THR
SEQRES  14    320  ALA GLN ALA ALA THR TYR ASN GLN ASN TYR PRO SER ALA
SEQRES  15    320  GLY LEU GLY ALA PRO GLY SER CYS GLN THR GLY ALA PRO
SEQRES  16    320  THR GLU THR VAL GLY GLY ASN THR HIS LEU LEU TYR SER
SEQRES  17    320  TRP ALA GLY THR ALA ILE GLN PRO THR ILE SER VAL PHE
SEQRES  18    320  GLY VAL THR GLY ALA THR ASP THR SER THR ILE PRO LEU
SEQRES  19    320  VAL ASP PRO ALA ASN ALA LEU ASP PRO SER THR LEU ALA
SEQRES  20    320  LEU PHE GLY THR GLY THR VAL MET VAL ASN ARG GLY SER
SEQRES  21    320  GLY GLN ASN ASP GLY VAL VAL SER LYS CYS SER ALA LEU
SEQRES  22    320  TYR GLY GLN VAL LEU SER THR SER TYR LYS TRP ASN HIS
SEQRES  23    320  LEU ASP GLU ILE ASN GLN LEU LEU GLY VAL ARG GLY ALA
SEQRES  24    320  ASN ALA GLU ASP PRO VAL ALA VAL ILE ARG THR HIS ALA
SEQRES  25    320  ASN ARG LEU LYS LEU ALA GLY VAL
HET     CA    321       1
HETNAM      CA CALCIUM ION
FORMUL   2   CA    CA1 2+
FORMUL   3  HOH   *193(H2 O1)
HELIX    1   1 ILE     33  GLN     39  1                                   7
HELIX    2   2 ARG     61  THR     76  1                                  16
HELIX    3   3 GLY     89  VAL     99  5                                  11
HELIX    4   4 GLU    118  LEU    127  1                                  10
HELIX    5   5 LEU    134  THR    150  1                                  17
HELIX    6   6 ALA    160  THR    166  1                                   7
HELIX    7   7 THR    169  ASN    178  1                                  10
HELIX    8   8 ALA    238  LEU    241  5                                   4
HELIX    9   9 SER    244  VAL    256  1                                  13
HELIX   10  10 LYS    269  ALA    272  1                                   4
HELIX   11  11 PRO    304  ALA    318  1                                  15
SHEET    1  A1 6 VAL    44  VAL    46  0
SHEET    2  A1 6 PRO    10  VAL    14  1  N  ILE    11   O  TYR    45
SHEET    3  A1 6 VAL    81  HIS    86  1  N  ASN    82   O  PRO    10
SHEET    4  A1 6 VAL   104  ILE   110  1  N  ALA   105   O  VAL    81
SHEET    5  A1 6 THR   203  GLY   211  1  N  LEU   205   O  VAL   107
SHEET    6  A1 6 THR   196  THR   198 -1  N  GLU   197   O  HIS   204
SHEET    1  A2 6 VAL    44  VAL    46  0
SHEET    2  A2 6 PRO    10  VAL    14  1  N  ILE    11   O  TYR    45
SHEET    3  A2 6 VAL    81  HIS    86  1  N  ASN    82   O  PRO    10
SHEET    4  A2 6 VAL   104  ILE   110  1  N  ALA   105   O  VAL    81
SHEET    5  A2 6 THR   203  GLY   211  1  N  LEU   205   O  VAL   107
SHEET    6  A2 6 GLN   276  TYR   282  1  N  SER   279   O  SER   208
SHEET    1   B 2 LYS    22  TYR    23  0
SHEET    2   B 2 LEU    27  GLU    28 -1  N  LEU    27   O  TYR    23
SHEET    1   C 2 ILE   214  VAL   220  0
SHEET    2   C 2 VAL   223  ASP   228 -1  N  THR   227   O  GLN   215
LINK        CA    CA   321                 OD1 ASP   288
LINK        CA    CA   321                 O   GLN   292
LINK        CA    CA   321                 O   VAL   296
LINK        CA    CA   321                 O   HOH   452
LINK        CA    CA   321                 OD1 ASP   242
LINK        CA    CA   321                 O   HOH   467
CISPEP   1 GLN    292    LEU    293          0        -1.88
SITE     1 ACT  3 SER    87  HIS   286  ASP   264
SITE     1 OXY  2 LEU    17  GLN    88
CRYST1   91.300   47.300   85.400  90.00 121.40  90.00 C 1 2 1       4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.010953  0.000000  0.006686        0.00000
SCALE2      0.000000  0.021142  0.000000        0.00000
SCALE3      0.000000  0.000000  0.013719        0.00000