longtext: 3LS2-pdb

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HEADER    HYDROLASE                               12-FEB-10   3LS2
TITLE     CRYSTAL STRUCTURE OF AN S-FORMYLGLUTATHIONE HYDROLASE FROM
TITLE    2 PSEUDOALTEROMONAS HALOPLANKTIS TAC125
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: S-FORMYLGLUTATHIONE HYDROLASE;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 EC: 3.1.1.-;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOALTEROMONAS HALOPLANKTIS;
SOURCE   3 ORGANISM_TAXID: 326442;
SOURCE   4 STRAIN: TAC125;
SOURCE   5 GENE: PSHAA1385;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PET28A
KEYWDS    S-FORMYLGLUTATHIONE HYDROLASE, PSYCHROPHILIC ORGANISM,
KEYWDS   2 PSEUDOALTEROMONAS HALOPLANKTIS, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    V.ALTERIO,G.DE SIMONE
REVDAT   1   23-MAR-10 3LS2    0
JRNL        AUTH   V.ALTERIO,V.AURILIA,A.ROMANELLI,A.PARRACINO,
JRNL        AUTH 2 M.SAVIANO,S.D'AURIA,G.DE SIMONE
JRNL        TITL   CRYSTAL STRUCTURE OF AN S-FORMYLGLUTATHIONE
JRNL        TITL 2 HYDROLASE FROM PSEUDOALTEROMONAS HALOPLANKTIS
JRNL        TITL 3 TAC125.
JRNL        REF    BIOPOLYMERS                                2010
JRNL        REFN                   ESSN 1079-0282
JRNL        PMID   20209484
JRNL        DOI    10.1002/BIP.21420
REMARK   2
REMARK   2 RESOLUTION.    2.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.1
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES, PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 95.9
REMARK   3   NUMBER OF REFLECTIONS             : 48748
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.161
REMARK   3   FREE R VALUE                     : 0.205
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : 2434
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : NULL
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : NULL
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : NULL
REMARK   3   BIN FREE R VALUE                    : NULL
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : NULL
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 8656
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 4
REMARK   3   SOLVENT ATOMS            : 1054
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 15.60
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.007
REMARK   3   BOND ANGLES            (DEGREES) : 1.40
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : ISOTROPIC
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : NULL
REMARK   3   KSOL        : NULL
REMARK   3   BSOL        : NULL
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3LS2 COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 23-FEB-10.
REMARK 100 THE RCSB ID CODE IS RCSB057658.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 10-JUL-09
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : ROTATING ANODE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU MICROMAX-007 HF
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.54178
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU SATURN 944
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 49990
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.200
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.7
REMARK 200  DATA REDUNDANCY                : 3.900
REMARK 200  R MERGE                    (I) : 0.05300
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 19.5000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.28
REMARK 200  COMPLETENESS FOR SHELL     (%) : 93.6
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.60
REMARK 200  R MERGE FOR SHELL          (I) : 0.16600
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 5.900
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: AMORE
REMARK 200 STARTING MODEL: PDB ENTRY 1PV1
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 37.94
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.98
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 20% (W/V) PEG MME 5000, 0.2 M
REMARK 280  SODIUM ACETATE, 0.1 M TRIS-HCL, PH 7.0, VAPOR DIFFUSION,
REMARK 280  HANGING DROP, TEMPERATURE 293 K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       24.74500
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       76.33500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       64.87500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       76.33500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       24.74500
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       64.87500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ALA A   280
REMARK 465     MET B     1
REMARK 465     ALA B   280
REMARK 465     MET C     1
REMARK 465     ALA C   280
REMARK 465     MET D     1
REMARK 465     ALA D   280
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     GLU A   41   CG    CD    OE1   OE2
REMARK 480     ASP A   88   CG    OD1   OD2
REMARK 480     GLU A  210   CD    OE1   OE2
REMARK 480     ASP A  229   CG    OD1   OD2
REMARK 480     GLU B   41   CG    CD    OE1   OE2
REMARK 480     LYS B   69   CD    CE    NZ
REMARK 480     GLN B  133   CD    OE1   NE2
REMARK 480     LYS C   69   CD    CE    NZ
REMARK 480     LYS C  208   CD    CE    NZ
REMARK 480     ASN C  226   CG    OD1   ND2
REMARK 480     LYS C  242   CD    CE    NZ
REMARK 480     ASN D  226   CG    OD1   ND2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    THR A  55       -9.91     71.28
REMARK 500    THR A  57     -151.98   -129.47
REMARK 500    SER A  84      174.62    177.59
REMARK 500    ALA A  99     -141.06     66.62
REMARK 500    ALA A 102       55.20   -152.16
REMARK 500    SER A 147     -110.05     55.87
REMARK 500    ASP A 257     -158.96   -107.50
REMARK 500    THR B  55       -5.16     70.38
REMARK 500    SER B  84      171.06    179.70
REMARK 500    ALA B  99     -141.27     66.67
REMARK 500    SER B 147     -108.41     52.66
REMARK 500    ASP B 257     -157.47   -111.92
REMARK 500    THR C  55       -8.17     72.47
REMARK 500    THR C  57     -156.90   -123.19
REMARK 500    SER C  84      176.19    174.71
REMARK 500    ASP C  94       46.76    -72.15
REMARK 500    ALA C  99     -135.62     69.21
REMARK 500    ALA C 102       57.81   -153.38
REMARK 500    PHE C 104       18.30     53.68
REMARK 500    SER C 147     -106.57     51.96
REMARK 500    PRO C 247       62.16    -69.73
REMARK 500    ASP C 257     -156.79   -106.34
REMARK 500    THR D  55       -4.21     74.42
REMARK 500    SER D  84      174.38    178.82
REMARK 500    ALA D  99     -138.70     68.52
REMARK 500    ALA D 102       56.61   -145.37
REMARK 500    PHE D 104       19.86     54.06
REMARK 500    SER D 147     -108.75     54.11
REMARK 500    LYS D 194       -6.65    -59.95
REMARK 500    PRO D 247       62.58    -69.66
REMARK 500    ASP D 257     -152.19   -107.11
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL B 281
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL C 281
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL A 281
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL D 281
DBREF  3LS2 A    1   280  UNP    Q3IL66   Q3IL66_PSEHT     1    280
DBREF  3LS2 B    1   280  UNP    Q3IL66   Q3IL66_PSEHT     1    280
DBREF  3LS2 C    1   280  UNP    Q3IL66   Q3IL66_PSEHT     1    280
DBREF  3LS2 D    1   280  UNP    Q3IL66   Q3IL66_PSEHT     1    280
SEQRES   1 A  280  MET LEU GLU ASN ILE SER SER VAL LYS VAL SER GLY GLY
SEQRES   2 A  280  TRP HIS LYS GLN TYR THR HIS SER ALA VAL SER THR HIS
SEQRES   3 A  280  CYS THR MET ARG PHE ALA VAL PHE LEU PRO PRO GLY ALA
SEQRES   4 A  280  SER GLU SER ASN LYS VAL PRO VAL LEU TYR TRP LEU SER
SEQRES   5 A  280  GLY LEU THR CYS THR ASP GLU ASN PHE MET GLN LYS ALA
SEQRES   6 A  280  GLY ALA PHE LYS LYS ALA ALA GLU LEU GLY ILE ALA ILE
SEQRES   7 A  280  VAL ALA PRO ASP THR SER PRO ARG GLY ASP ASN VAL PRO
SEQRES   8 A  280  ASN GLU ASP SER TYR ASP PHE ALA GLN GLY ALA GLY PHE
SEQRES   9 A  280  TYR VAL ASN ALA THR GLN ALA PRO TYR ASN THR HIS PHE
SEQRES  10 A  280  ASN MET TYR ASP TYR VAL VAL ASN GLU LEU PRO ALA LEU
SEQRES  11 A  280  ILE GLU GLN HIS PHE PRO VAL THR SER THR LYS ALA ILE
SEQRES  12 A  280  SER GLY HIS SER MET GLY GLY HIS GLY ALA LEU MET ILE
SEQRES  13 A  280  ALA LEU LYS ASN PRO GLN ASP TYR VAL SER ALA SER ALA
SEQRES  14 A  280  PHE SER PRO ILE VAL ASN PRO ILE ASN CYS PRO TRP GLY
SEQRES  15 A  280  VAL LYS ALA PHE THR GLY TYR LEU GLY ALA ASP LYS THR
SEQRES  16 A  280  THR TRP ALA GLN TYR ASP SER CYS LYS LEU MET ALA LYS
SEQRES  17 A  280  ALA GLU GLN SER ASN TYR LEU PRO MET LEU VAL SER GLN
SEQRES  18 A  280  GLY ASP ALA ASP ASN PHE LEU ASP GLU GLN LEU LYS PRO
SEQRES  19 A  280  GLN ASN LEU VAL ALA VAL ALA LYS GLN LYS ASP TYR PRO
SEQRES  20 A  280  LEU THR LEU GLU MET GLN THR GLY TYR ASP HIS SER TYR
SEQRES  21 A  280  PHE PHE ILE SER SER PHE ILE ASP GLN HIS LEU VAL PHE
SEQRES  22 A  280  HIS HIS GLN TYR LEU SER ALA
SEQRES   1 B  280  MET LEU GLU ASN ILE SER SER VAL LYS VAL SER GLY GLY
SEQRES   2 B  280  TRP HIS LYS GLN TYR THR HIS SER ALA VAL SER THR HIS
SEQRES   3 B  280  CYS THR MET ARG PHE ALA VAL PHE LEU PRO PRO GLY ALA
SEQRES   4 B  280  SER GLU SER ASN LYS VAL PRO VAL LEU TYR TRP LEU SER
SEQRES   5 B  280  GLY LEU THR CYS THR ASP GLU ASN PHE MET GLN LYS ALA
SEQRES   6 B  280  GLY ALA PHE LYS LYS ALA ALA GLU LEU GLY ILE ALA ILE
SEQRES   7 B  280  VAL ALA PRO ASP THR SER PRO ARG GLY ASP ASN VAL PRO
SEQRES   8 B  280  ASN GLU ASP SER TYR ASP PHE ALA GLN GLY ALA GLY PHE
SEQRES   9 B  280  TYR VAL ASN ALA THR GLN ALA PRO TYR ASN THR HIS PHE
SEQRES  10 B  280  ASN MET TYR ASP TYR VAL VAL ASN GLU LEU PRO ALA LEU
SEQRES  11 B  280  ILE GLU GLN HIS PHE PRO VAL THR SER THR LYS ALA ILE
SEQRES  12 B  280  SER GLY HIS SER MET GLY GLY HIS GLY ALA LEU MET ILE
SEQRES  13 B  280  ALA LEU LYS ASN PRO GLN ASP TYR VAL SER ALA SER ALA
SEQRES  14 B  280  PHE SER PRO ILE VAL ASN PRO ILE ASN CYS PRO TRP GLY
SEQRES  15 B  280  VAL LYS ALA PHE THR GLY TYR LEU GLY ALA ASP LYS THR
SEQRES  16 B  280  THR TRP ALA GLN TYR ASP SER CYS LYS LEU MET ALA LYS
SEQRES  17 B  280  ALA GLU GLN SER ASN TYR LEU PRO MET LEU VAL SER GLN
SEQRES  18 B  280  GLY ASP ALA ASP ASN PHE LEU ASP GLU GLN LEU LYS PRO
SEQRES  19 B  280  GLN ASN LEU VAL ALA VAL ALA LYS GLN LYS ASP TYR PRO
SEQRES  20 B  280  LEU THR LEU GLU MET GLN THR GLY TYR ASP HIS SER TYR
SEQRES  21 B  280  PHE PHE ILE SER SER PHE ILE ASP GLN HIS LEU VAL PHE
SEQRES  22 B  280  HIS HIS GLN TYR LEU SER ALA
SEQRES   1 C  280  MET LEU GLU ASN ILE SER SER VAL LYS VAL SER GLY GLY
SEQRES   2 C  280  TRP HIS LYS GLN TYR THR HIS SER ALA VAL SER THR HIS
SEQRES   3 C  280  CYS THR MET ARG PHE ALA VAL PHE LEU PRO PRO GLY ALA
SEQRES   4 C  280  SER GLU SER ASN LYS VAL PRO VAL LEU TYR TRP LEU SER
SEQRES   5 C  280  GLY LEU THR CYS THR ASP GLU ASN PHE MET GLN LYS ALA
SEQRES   6 C  280  GLY ALA PHE LYS LYS ALA ALA GLU LEU GLY ILE ALA ILE
SEQRES   7 C  280  VAL ALA PRO ASP THR SER PRO ARG GLY ASP ASN VAL PRO
SEQRES   8 C  280  ASN GLU ASP SER TYR ASP PHE ALA GLN GLY ALA GLY PHE
SEQRES   9 C  280  TYR VAL ASN ALA THR GLN ALA PRO TYR ASN THR HIS PHE
SEQRES  10 C  280  ASN MET TYR ASP TYR VAL VAL ASN GLU LEU PRO ALA LEU
SEQRES  11 C  280  ILE GLU GLN HIS PHE PRO VAL THR SER THR LYS ALA ILE
SEQRES  12 C  280  SER GLY HIS SER MET GLY GLY HIS GLY ALA LEU MET ILE
SEQRES  13 C  280  ALA LEU LYS ASN PRO GLN ASP TYR VAL SER ALA SER ALA
SEQRES  14 C  280  PHE SER PRO ILE VAL ASN PRO ILE ASN CYS PRO TRP GLY
SEQRES  15 C  280  VAL LYS ALA PHE THR GLY TYR LEU GLY ALA ASP LYS THR
SEQRES  16 C  280  THR TRP ALA GLN TYR ASP SER CYS LYS LEU MET ALA LYS
SEQRES  17 C  280  ALA GLU GLN SER ASN TYR LEU PRO MET LEU VAL SER GLN
SEQRES  18 C  280  GLY ASP ALA ASP ASN PHE LEU ASP GLU GLN LEU LYS PRO
SEQRES  19 C  280  GLN ASN LEU VAL ALA VAL ALA LYS GLN LYS ASP TYR PRO
SEQRES  20 C  280  LEU THR LEU GLU MET GLN THR GLY TYR ASP HIS SER TYR
SEQRES  21 C  280  PHE PHE ILE SER SER PHE ILE ASP GLN HIS LEU VAL PHE
SEQRES  22 C  280  HIS HIS GLN TYR LEU SER ALA
SEQRES   1 D  280  MET LEU GLU ASN ILE SER SER VAL LYS VAL SER GLY GLY
SEQRES   2 D  280  TRP HIS LYS GLN TYR THR HIS SER ALA VAL SER THR HIS
SEQRES   3 D  280  CYS THR MET ARG PHE ALA VAL PHE LEU PRO PRO GLY ALA
SEQRES   4 D  280  SER GLU SER ASN LYS VAL PRO VAL LEU TYR TRP LEU SER
SEQRES   5 D  280  GLY LEU THR CYS THR ASP GLU ASN PHE MET GLN LYS ALA
SEQRES   6 D  280  GLY ALA PHE LYS LYS ALA ALA GLU LEU GLY ILE ALA ILE
SEQRES   7 D  280  VAL ALA PRO ASP THR SER PRO ARG GLY ASP ASN VAL PRO
SEQRES   8 D  280  ASN GLU ASP SER TYR ASP PHE ALA GLN GLY ALA GLY PHE
SEQRES   9 D  280  TYR VAL ASN ALA THR GLN ALA PRO TYR ASN THR HIS PHE
SEQRES  10 D  280  ASN MET TYR ASP TYR VAL VAL ASN GLU LEU PRO ALA LEU
SEQRES  11 D  280  ILE GLU GLN HIS PHE PRO VAL THR SER THR LYS ALA ILE
SEQRES  12 D  280  SER GLY HIS SER MET GLY GLY HIS GLY ALA LEU MET ILE
SEQRES  13 D  280  ALA LEU LYS ASN PRO GLN ASP TYR VAL SER ALA SER ALA
SEQRES  14 D  280  PHE SER PRO ILE VAL ASN PRO ILE ASN CYS PRO TRP GLY
SEQRES  15 D  280  VAL LYS ALA PHE THR GLY TYR LEU GLY ALA ASP LYS THR
SEQRES  16 D  280  THR TRP ALA GLN TYR ASP SER CYS LYS LEU MET ALA LYS
SEQRES  17 D  280  ALA GLU GLN SER ASN TYR LEU PRO MET LEU VAL SER GLN
SEQRES  18 D  280  GLY ASP ALA ASP ASN PHE LEU ASP GLU GLN LEU LYS PRO
SEQRES  19 D  280  GLN ASN LEU VAL ALA VAL ALA LYS GLN LYS ASP TYR PRO
SEQRES  20 D  280  LEU THR LEU GLU MET GLN THR GLY TYR ASP HIS SER TYR
SEQRES  21 D  280  PHE PHE ILE SER SER PHE ILE ASP GLN HIS LEU VAL PHE
SEQRES  22 D  280  HIS HIS GLN TYR LEU SER ALA
HET     CL  A 281       1
HET     CL  B 281       1
HET     CL  C 281       1
HET     CL  D 281       1
HETNAM      CL CHLORIDE ION
FORMUL   5   CL    4(CL 1-)
FORMUL   9  HOH   *1054(H2 O)
HELIX    1   1 ASP A   58  ALA A   65  1                                   8
HELIX    2   2 ALA A   67  GLY A   75  1                                   9
HELIX    3   3 ASN A  118  ASN A  125  1                                   8
HELIX    4   4 ASN A  125  PHE A  135  1                                  11
HELIX    5   5 SER A  147  ASN A  160  1                                  14
HELIX    6   6 ASN A  175  ASN A  178  5                                   4
HELIX    7   7 CYS A  179  GLY A  191  1                                  13
HELIX    8   8 ASP A  193  TYR A  200  5                                   8
HELIX    9   9 ASP A  201  LYS A  208  1                                   8
HELIX   10  10 ALA A  209  TYR A  214  5                                   6
HELIX   11  11 LYS A  233  ASP A  245  1                                  13
HELIX   12  12 SER A  259  SER A  279  1                                  21
HELIX   13  13 ASP B   58  ALA B   65  1                                   8
HELIX   14  14 ALA B   67  GLY B   75  1                                   9
HELIX   15  15 ASN B  118  ASN B  125  1                                   8
HELIX   16  16 ASN B  125  PHE B  135  1                                  11
HELIX   17  17 SER B  147  ASN B  160  1                                  14
HELIX   18  18 ASN B  175  ASN B  178  5                                   4
HELIX   19  19 CYS B  179  GLY B  191  1                                  13
HELIX   20  20 ASP B  193  TYR B  200  5                                   8
HELIX   21  21 ASP B  201  LYS B  208  1                                   8
HELIX   22  22 GLU B  210  TYR B  214  5                                   5
HELIX   23  23 LYS B  233  ASP B  245  1                                  13
HELIX   24  24 SER B  259  SER B  279  1                                  21
HELIX   25  25 ASP C   58  ALA C   65  1                                   8
HELIX   26  26 ALA C   67  GLY C   75  1                                   9
HELIX   27  27 ASN C  118  ASN C  125  1                                   8
HELIX   28  28 ASN C  125  PHE C  135  1                                  11
HELIX   29  29 SER C  147  ASN C  160  1                                  14
HELIX   30  30 ASN C  175  ASN C  178  5                                   4
HELIX   31  31 CYS C  179  GLY C  191  1                                  13
HELIX   32  32 ASP C  193  TYR C  200  5                                   8
HELIX   33  33 ASP C  201  LYS C  208  1                                   8
HELIX   34  34 ALA C  209  TYR C  214  5                                   6
HELIX   35  35 LYS C  233  ASP C  245  1                                  13
HELIX   36  36 SER C  259  SER C  279  1                                  21
HELIX   37  37 ASP D   58  ALA D   65  1                                   8
HELIX   38  38 ALA D   67  GLY D   75  1                                   9
HELIX   39  39 ASN D  118  ASN D  125  1                                   8
HELIX   40  40 ASN D  125  PHE D  135  1                                  11
HELIX   41  41 SER D  147  ASN D  160  1                                  14
HELIX   42  42 ASN D  175  ASN D  178  5                                   4
HELIX   43  43 CYS D  179  GLY D  191  1                                  13
HELIX   44  44 ASP D  193  GLN D  199  1                                   7
HELIX   45  45 ASP D  201  LYS D  208  1                                   8
HELIX   46  46 ALA D  209  TYR D  214  5                                   6
HELIX   47  47 LYS D  233  ASP D  245  1                                  13
HELIX   48  48 SER D  259  SER D  279  1                                  21
SHEET    1   A 6 GLU A   3  VAL A  10  0
SHEET    2   A 6 GLY A  13  ALA A  22 -1  O  HIS A  15   N  VAL A   8
SHEET    3   A 6 CYS A  27  LEU A  35 -1  O  PHE A  31   N  TYR A  18
SHEET    4   A 6 ALA A  77  ALA A  80 -1  O  ILE A  78   N  PHE A  34
SHEET    5   A 6 VAL A  45  LEU A  51  1  N  TRP A  50   O  VAL A  79
SHEET    6   A 6 VAL A 137  GLY A 145  1  O  SER A 144   N  TYR A  49
SHEET    1   B 3 ALA A 167  PHE A 170  0
SHEET    2   B 3 MET A 217  GLY A 222  1  O  SER A 220   N  ALA A 169
SHEET    3   B 3 LEU A 248  GLN A 253  1  O  GLU A 251   N  VAL A 219
SHEET    1   C 6 GLU B   3  VAL B  10  0
SHEET    2   C 6 GLY B  13  ALA B  22 -1  O  HIS B  15   N  VAL B   8
SHEET    3   C 6 CYS B  27  LEU B  35 -1  O  MET B  29   N  HIS B  20
SHEET    4   C 6 ALA B  77  ALA B  80 -1  O  ALA B  80   N  ALA B  32
SHEET    5   C 6 VAL B  45  LEU B  51  1  N  TRP B  50   O  VAL B  79
SHEET    6   C 6 VAL B 137  GLY B 145  1  O  ALA B 142   N  TYR B  49
SHEET    1   D 3 ALA B 167  PHE B 170  0
SHEET    2   D 3 MET B 217  GLY B 222  1  O  SER B 220   N  ALA B 169
SHEET    3   D 3 LEU B 248  GLN B 253  1  O  GLN B 253   N  GLN B 221
SHEET    1   E 6 GLU C   3  VAL C  10  0
SHEET    2   E 6 GLY C  13  ALA C  22 -1  O  THR C  19   N  GLU C   3
SHEET    3   E 6 CYS C  27  LEU C  35 -1  O  VAL C  33   N  LYS C  16
SHEET    4   E 6 ALA C  77  ALA C  80 -1  O  ALA C  80   N  ALA C  32
SHEET    5   E 6 VAL C  45  LEU C  51  1  N  LEU C  48   O  ALA C  77
SHEET    6   E 6 VAL C 137  GLY C 145  1  O  ALA C 142   N  TYR C  49
SHEET    1   F 3 ALA C 167  PHE C 170  0
SHEET    2   F 3 MET C 217  GLY C 222  1  O  SER C 220   N  ALA C 169
SHEET    3   F 3 LEU C 248  GLN C 253  1  O  GLN C 253   N  GLN C 221
SHEET    1   G 6 GLU D   3  VAL D  10  0
SHEET    2   G 6 GLY D  13  ALA D  22 -1  O  THR D  19   N  GLU D   3
SHEET    3   G 6 CYS D  27  LEU D  35 -1  O  PHE D  31   N  TYR D  18
SHEET    4   G 6 ALA D  77  ALA D  80 -1  O  ALA D  80   N  ALA D  32
SHEET    5   G 6 VAL D  45  LEU D  51  1  N  TRP D  50   O  VAL D  79
SHEET    6   G 6 VAL D 137  GLY D 145  1  O  SER D 144   N  TYR D  49
SHEET    1   H 3 ALA D 167  PHE D 170  0
SHEET    2   H 3 MET D 217  GLY D 222  1  O  SER D 220   N  ALA D 169
SHEET    3   H 3 LEU D 248  GLN D 253  1  O  GLU D 251   N  VAL D 219
CISPEP   1 ALA A  111    PRO A  112          0         0.09
CISPEP   2 ALA B  111    PRO B  112          0         1.09
CISPEP   3 ALA C  111    PRO C  112          0        -0.06
CISPEP   4 ALA D  111    PRO D  112          0        -1.30
SITE     1 AC1  6 PRO B  91  PRO B 112  TYR B 113  THR B 115
SITE     2 AC1  6 HIS B 116  PHE B 117
SITE     1 AC2  6 PRO C  91  PRO C 112  TYR C 113  THR C 115
SITE     2 AC2  6 HIS C 116  PHE C 117
SITE     1 AC3  6 PRO A  91  PRO A 112  TYR A 113  THR A 115
SITE     2 AC3  6 HIS A 116  PHE A 117
SITE     1 AC4  6 PRO D  91  PRO D 112  TYR D 113  THR D 115
SITE     2 AC4  6 HIS D 116  PHE D 117
CRYST1   49.490  129.750  152.670  90.00  90.00  90.00 P 21 21 21   16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.020206  0.000000  0.000000        0.00000
SCALE2      0.000000  0.007707  0.000000        0.00000
SCALE3      0.000000  0.000000  0.006550        0.00000
TER    2165      SER A 279
TER    4330      SER B 279
TER    6495      SER C 279
TER    8660      SER D 279
MASTER      315    0    4   48   36    0    8    6 9714    4    0   88
END