longtext: 3N2Z-pdb

content
HEADER    HYDROLASE                               19-MAY-10   3N2Z
TITLE     THE STRUCTURE OF HUMAN PROLYLCARBOXYPEPTIDASE AT 2.80 ANGSTROMS
TITLE    2 RESOLUTION
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LYSOSOMAL PRO-X CARBOXYPEPTIDASE;
COMPND   3 CHAIN: B;
COMPND   4 SYNONYM: PROLYLCARBOXYPEPTIDASE, PRCP, PROLINE CARBOXYPEPTIDASE,
COMPND   5 ANGIOTENSINASE C, LYSOSOMAL CARBOXYPEPTIDASE C;
COMPND   6 EC: 3.4.16.2;
COMPND   7 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: PRCP, PCP;
SOURCE   6 EXPRESSION_SYSTEM: CRICETULUS GRISEUS;
SOURCE   7 EXPRESSION_SYSTEM_COMMON: CHINESE HAMSTER;
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 10029;
SOURCE   9 EXPRESSION_SYSTEM_CELL: CHO CELLS
KEYWDS    ALPHA/BETA HYDROLASE, PRCP, SERINE CARBOXYPEPTIDASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.M.SOISSON,S.B.PATEL,K.J.LUMB,S.SHARMA
REVDAT   1   07-JUL-10 3N2Z    0
JRNL        AUTH   S.M.SOISSON,S.B.PATEL,P.D.ABEYWICKREMA,N.J.BRYNE,R.E.DIEHL,
JRNL        AUTH 2 D.L.HALL,R.E.FORD,J.C.REID,K.W.RICKERT,J.M.SHIPMAN,S.SHARMA,
JRNL        AUTH 3 K.J.LUMB
JRNL        TITL   STRUCTURAL DEFINITION AND SUBSTRATE SPECIFICITY OF THE S28
JRNL        TITL 2 PROTEASE FAMILY: THE CRYSTAL STRUCTURE OF HUMAN
JRNL        TITL 3 PROLYLCARBOXYPEPTIDASE.
JRNL        REF    BMC STRUCT.BIOL.              V.  10    16 2010
JRNL        REFN                   ESSN 1472-6807
JRNL        PMID   20540760
JRNL        DOI    10.1186/1472-6807-10-16
REMARK   2
REMARK   2 RESOLUTION.    2.79 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : BUSTER 2.9.4
REMARK   3   AUTHORS     : BRICOGNE,BLANC,BRANDL,FLENSBURG,KELLER,
REMARK   3               : PACIOREK,ROVERSI,SMART,VONRHEIN,WOMACK,
REMARK   3               : MATTHEWS,TEN EYCK,TRONRUD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.79
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 47.67
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.2
REMARK   3   NUMBER OF REFLECTIONS             : 37540
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD           : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION   : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET)  : 0.218
REMARK   3   R VALUE            (WORKING SET)  : 0.216
REMARK   3   FREE R VALUE                      : 0.244
REMARK   3   FREE R VALUE TEST SET SIZE   (%)  : 4.990
REMARK   3   FREE R VALUE TEST SET COUNT       : 1875
REMARK   3   ESTIMATED ERROR OF FREE R VALUE   : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED               : 19
REMARK   3   BIN RESOLUTION RANGE HIGH   (ANGSTROMS) : 2.79
REMARK   3   BIN RESOLUTION RANGE LOW    (ANGSTROMS) : 2.87
REMARK   3   BIN COMPLETENESS (WORKING+TEST)     (%) : 99.19
REMARK   3   REFLECTIONS IN BIN (WORKING + TEST SET) : 2839
REMARK   3   BIN R VALUE        (WORKING + TEST SET) : 0.3170
REMARK   3   REFLECTIONS IN BIN        (WORKING SET) : 2703
REMARK   3   BIN R VALUE               (WORKING SET) : 0.3150
REMARK   3   BIN FREE R VALUE                        : 0.3520
REMARK   3   BIN FREE R VALUE TEST SET SIZE      (%) : 4.79
REMARK   3   BIN FREE R VALUE TEST SET COUNT         : 136
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE     : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 3539
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 89
REMARK   3   SOLVENT ATOMS            : 76
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 64.57
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -16.02700
REMARK   3    B22 (A**2) : -16.02700
REMARK   3    B33 (A**2) : 32.05500
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT                    (A) : 0.43
REMARK   3   DPI (BLOW EQ-10) BASED ON R VALUE        (A) : NULL
REMARK   3   DPI (BLOW EQ-9) BASED ON FREE R VALUE    (A) : NULL
REMARK   3   DPI (CRUICKSHANK) BASED ON R VALUE       (A) : NULL
REMARK   3   DPI (CRUICKSHANK) BASED ON FREE R VALUE  (A) : NULL
REMARK   3
REMARK   3   REFERENCES: BLOW, D. (2002) ACTA CRYST D58, 792-797
REMARK   3               CRUICKSHANK, D.W.J. (1999) ACTA CRYST D55, 583-601
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.894
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.878
REMARK   3
REMARK   3   NUMBER OF GEOMETRIC FUNCTION TERMS DEFINED : 15
REMARK   3   TERM                          COUNT    WEIGHT   FUNCTION.
REMARK   3    BOND LENGTHS              : NULL   ; NULL   ; NULL
REMARK   3    BOND ANGLES               : NULL   ; NULL   ; NULL
REMARK   3    TORSION ANGLES            : NULL   ; NULL   ; NULL
REMARK   3    TRIGONAL CARBON PLANES    : NULL   ; NULL   ; NULL
REMARK   3    GENERAL PLANES            : NULL   ; NULL   ; NULL
REMARK   3    ISOTROPIC THERMAL FACTORS : NULL   ; NULL   ; NULL
REMARK   3    BAD NON-BONDED CONTACTS   : NULL   ; NULL   ; NULL
REMARK   3    IMPROPER TORSIONS         : NULL   ; NULL   ; NULL
REMARK   3    PSEUDOROTATION ANGLES     : NULL   ; NULL   ; NULL
REMARK   3    CHIRAL IMPROPER TORSION   : NULL   ; NULL   ; NULL
REMARK   3    SUM OF OCCUPANCIES        : NULL   ; NULL   ; NULL
REMARK   3    UTILITY DISTANCES         : NULL   ; NULL   ; NULL
REMARK   3    UTILITY ANGLES            : NULL   ; NULL   ; NULL
REMARK   3    UTILITY TORSION           : NULL   ; NULL   ; NULL
REMARK   3    IDEAL-DIST CONTACT TERM   : NULL   ; NULL   ; NULL
REMARK   3
REMARK   3   RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3    BOND LENGTHS                       (A) : NULL
REMARK   3    BOND ANGLES                  (DEGREES) : NULL
REMARK   3    PEPTIDE OMEGA TORSION ANGLES (DEGREES) : NULL
REMARK   3    OTHER TORSION ANGLES         (DEGREES) : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3N2Z COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 07-JUN-10.
REMARK 100 THE RCSB ID CODE IS RCSB059314.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 08-MAY-09
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ALS
REMARK 200  BEAMLINE                       : 5.0.2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.00
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL
REMARK 200  DATA SCALING SOFTWARE          : HKL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 37814
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.790
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : 10.900
REMARK 200  R MERGE                    (I) : 0.09800
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 9.9000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.79
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.87
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 10.90
REMARK 200  R MERGE FOR SHELL          (I) : 0.59400
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MIRAS
REMARK 200 SOFTWARE USED: SHARP
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): NULL
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): NULL
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 1.8M AMMONIUM SULFATE, 0.1M HEPES, 1-
REMARK 280  2% PEG400 MIXED IN A 2:1 RATIO, PH 7.5, VAPOR DIFFUSION, HANGING
REMARK 280  DROP, TEMPERATURE 277K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: H 3 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z
REMARK 290       3555   -X+Y,-X,Z
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z
REMARK 290       6555   -X,-X+Y,-Z
REMARK 290       7555   X+2/3,Y+1/3,Z+1/3
REMARK 290       8555   -Y+2/3,X-Y+1/3,Z+1/3
REMARK 290       9555   -X+Y+2/3,-X+1/3,Z+1/3
REMARK 290      10555   Y+2/3,X+1/3,-Z+1/3
REMARK 290      11555   X-Y+2/3,-Y+1/3,-Z+1/3
REMARK 290      12555   -X+2/3,-X+Y+1/3,-Z+1/3
REMARK 290      13555   X+1/3,Y+2/3,Z+2/3
REMARK 290      14555   -Y+1/3,X-Y+2/3,Z+2/3
REMARK 290      15555   -X+Y+1/3,-X+2/3,Z+2/3
REMARK 290      16555   Y+1/3,X+2/3,-Z+2/3
REMARK 290      17555   X-Y+1/3,-Y+2/3,-Z+2/3
REMARK 290      18555   -X+1/3,-X+Y+2/3,-Z+2/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   7  1.000000  0.000000  0.000000       90.57000
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       52.29061
REMARK 290   SMTRY3   7  0.000000  0.000000  1.000000       80.04333
REMARK 290   SMTRY1   8 -0.500000 -0.866025  0.000000       90.57000
REMARK 290   SMTRY2   8  0.866025 -0.500000  0.000000       52.29061
REMARK 290   SMTRY3   8  0.000000  0.000000  1.000000       80.04333
REMARK 290   SMTRY1   9 -0.500000  0.866025  0.000000       90.57000
REMARK 290   SMTRY2   9 -0.866025 -0.500000  0.000000       52.29061
REMARK 290   SMTRY3   9  0.000000  0.000000  1.000000       80.04333
REMARK 290   SMTRY1  10 -0.500000  0.866025  0.000000       90.57000
REMARK 290   SMTRY2  10  0.866025  0.500000  0.000000       52.29061
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000       80.04333
REMARK 290   SMTRY1  11  1.000000  0.000000  0.000000       90.57000
REMARK 290   SMTRY2  11  0.000000 -1.000000  0.000000       52.29061
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000       80.04333
REMARK 290   SMTRY1  12 -0.500000 -0.866025  0.000000       90.57000
REMARK 290   SMTRY2  12 -0.866025  0.500000  0.000000       52.29061
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000       80.04333
REMARK 290   SMTRY1  13  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  13  0.000000  1.000000  0.000000      104.58123
REMARK 290   SMTRY3  13  0.000000  0.000000  1.000000      160.08667
REMARK 290   SMTRY1  14 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  14  0.866025 -0.500000  0.000000      104.58123
REMARK 290   SMTRY3  14  0.000000  0.000000  1.000000      160.08667
REMARK 290   SMTRY1  15 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  15 -0.866025 -0.500000  0.000000      104.58123
REMARK 290   SMTRY3  15  0.000000  0.000000  1.000000      160.08667
REMARK 290   SMTRY1  16 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  16  0.866025  0.500000  0.000000      104.58123
REMARK 290   SMTRY3  16  0.000000  0.000000 -1.000000      160.08667
REMARK 290   SMTRY1  17  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  17  0.000000 -1.000000  0.000000      104.58123
REMARK 290   SMTRY3  17  0.000000  0.000000 -1.000000      160.08667
REMARK 290   SMTRY1  18 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  18 -0.866025  0.500000  0.000000      104.58123
REMARK 290   SMTRY3  18  0.000000  0.000000 -1.000000      160.08667
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 6950 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 37140 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -3.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   2  0.000000 -1.000000  0.000000      104.58123
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      160.08667
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     THR B   349
REMARK 465     ALA B   350
REMARK 465     THR B   351
REMARK 465     SER B   352
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   ND2  ASN B   101     O5   NAG B     1              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    TYR B  78      -19.27     85.39
REMARK 500    ASN B  92     -163.45   -117.99
REMARK 500    GLU B  93       58.06    -55.62
REMARK 500    TYR B 124        4.99     81.03
REMARK 500    SER B 128       55.91    -96.89
REMARK 500    LYS B 137      -70.17    -51.79
REMARK 500    GLU B 169      -48.96    -22.55
REMARK 500    SER B 179     -111.74     59.48
REMARK 500    TYR B 192       62.44   -117.24
REMARK 500    GLN B 208       49.48    -74.73
REMARK 500    GLU B 210     -127.72     45.25
REMARK 500    VAL B 217      -54.71    -29.81
REMARK 500    PHE B 297      -78.46    -74.88
REMARK 500    LEU B 313     -101.06    -87.04
REMARK 500    LYS B 314      -41.20     50.99
REMARK 500    ASN B 317       72.11    -67.71
REMARK 500    ASN B 336       26.41   -140.94
REMARK 500    THR B 365      -79.54   -109.26
REMARK 500    GLU B 452       10.00     80.77
REMARK 500    ALA B 454     -132.73    -87.98
REMARK 500    HIS B 456       58.56     39.81
REMARK 500    ALA B 464      -40.66    125.65
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    ASP B  76        23.8      L          L   OUTSIDE RANGE
REMARK 500    GLU B  93        24.5      L          L   OUTSIDE RANGE
REMARK 500    ASN B 102        24.0      L          L   OUTSIDE RANGE
REMARK 500    VAL B 217        22.5      L          L   OUTSIDE RANGE
REMARK 500    ILE B 272        24.6      L          L   OUTSIDE RANGE
REMARK 500    LYS B 314        23.1      L          L   OUTSIDE RANGE
REMARK 500    VAL B 333        23.2      L          L   OUTSIDE RANGE
REMARK 500    TRP B 406        25.0      L          L   OUTSIDE RANGE
REMARK 500    VAL B 437        24.7      L          L   OUTSIDE RANGE
REMARK 500    HIS B 456        24.8      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH B 495        DISTANCE =  5.92 ANGSTROMS
REMARK 525    HOH B 500        DISTANCE =  6.34 ANGSTROMS
REMARK 525    HOH B 519        DISTANCE =  5.14 ANGSTROMS
REMARK 525    HOH B 521        DISTANCE =  5.08 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 2
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 3
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 4
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 5
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 6
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 B 492
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3JYH   RELATED DB: PDB
DBREF  3N2Z B   46   491  UNP    P42785   PCP_HUMAN       46    491
SEQRES   1 B  446  LYS ASN TYR SER VAL LEU TYR PHE GLN GLN LYS VAL ASP
SEQRES   2 B  446  HIS PHE GLY PHE ASN THR VAL LYS THR PHE ASN GLN ARG
SEQRES   3 B  446  TYR LEU VAL ALA ASP LYS TYR TRP LYS LYS ASN GLY GLY
SEQRES   4 B  446  SER ILE LEU PHE TYR THR GLY ASN GLU GLY ASP ILE ILE
SEQRES   5 B  446  TRP PHE CYS ASN ASN THR GLY PHE MET TRP ASP VAL ALA
SEQRES   6 B  446  GLU GLU LEU LYS ALA MET LEU VAL PHE ALA GLU HIS ARG
SEQRES   7 B  446  TYR TYR GLY GLU SER LEU PRO PHE GLY ASP ASN SER PHE
SEQRES   8 B  446  LYS ASP SER ARG HIS LEU ASN PHE LEU THR SER GLU GLN
SEQRES   9 B  446  ALA LEU ALA ASP PHE ALA GLU LEU ILE LYS HIS LEU LYS
SEQRES  10 B  446  ARG THR ILE PRO GLY ALA GLU ASN GLN PRO VAL ILE ALA
SEQRES  11 B  446  ILE GLY GLY SER TYR GLY GLY MET LEU ALA ALA TRP PHE
SEQRES  12 B  446  ARG MET LYS TYR PRO HIS MET VAL VAL GLY ALA LEU ALA
SEQRES  13 B  446  ALA SER ALA PRO ILE TRP GLN PHE GLU ASP LEU VAL PRO
SEQRES  14 B  446  CYS GLY VAL PHE MET LYS ILE VAL THR THR ASP PHE ARG
SEQRES  15 B  446  LYS SER GLY PRO HIS CYS SER GLU SER ILE HIS ARG SER
SEQRES  16 B  446  TRP ASP ALA ILE ASN ARG LEU SER ASN THR GLY SER GLY
SEQRES  17 B  446  LEU GLN TRP LEU THR GLY ALA LEU HIS LEU CYS SER PRO
SEQRES  18 B  446  LEU THR SER GLN ASP ILE GLN HIS LEU LYS ASP TRP ILE
SEQRES  19 B  446  SER GLU THR TRP VAL ASN LEU ALA MET VAL ASP TYR PRO
SEQRES  20 B  446  TYR ALA SER ASN PHE LEU GLN PRO LEU PRO ALA TRP PRO
SEQRES  21 B  446  ILE LYS VAL VAL CYS GLN TYR LEU LYS ASN PRO ASN VAL
SEQRES  22 B  446  SER ASP SER LEU LEU LEU GLN ASN ILE PHE GLN ALA LEU
SEQRES  23 B  446  ASN VAL TYR TYR ASN TYR SER GLY GLN VAL LYS CYS LEU
SEQRES  24 B  446  ASN ILE SER GLU THR ALA THR SER SER LEU GLY THR LEU
SEQRES  25 B  446  GLY TRP SER TYR GLN ALA CYS THR GLU VAL VAL MET PRO
SEQRES  26 B  446  PHE CYS THR ASN GLY VAL ASP ASP MET PHE GLU PRO HIS
SEQRES  27 B  446  SER TRP ASN LEU LYS GLU LEU SER ASP ASP CYS PHE GLN
SEQRES  28 B  446  GLN TRP GLY VAL ARG PRO ARG PRO SER TRP ILE THR THR
SEQRES  29 B  446  MET TYR GLY GLY LYS ASN ILE SER SER HIS THR ASN ILE
SEQRES  30 B  446  VAL PHE SER ASN GLY GLU LEU ASP PRO TRP SER GLY GLY
SEQRES  31 B  446  GLY VAL THR LYS ASP ILE THR ASP THR LEU VAL ALA VAL
SEQRES  32 B  446  THR ILE SER GLU GLY ALA HIS HIS LEU ASP LEU ARG THR
SEQRES  33 B  446  LYS ASN ALA LEU ASP PRO MET SER VAL LEU LEU ALA ARG
SEQRES  34 B  446  SER LEU GLU VAL ARG HIS MET LYS ASN TRP ILE ARG ASP
SEQRES  35 B  446  PHE TYR ASP SER
MODRES 3N2Z ASN B  336  ASN  GLYCOSYLATION SITE
MODRES 3N2Z ASN B  415  ASN  GLYCOSYLATION SITE
MODRES 3N2Z ASN B  101  ASN  GLYCOSYLATION SITE
MODRES 3N2Z ASN B  345  ASN  GLYCOSYLATION SITE
MODRES 3N2Z ASN B  317  ASN  GLYCOSYLATION SITE
HET    NAG  B   1      14
HET    NAG  B   2      14
HET    NAG  B   3      14
HET    NAG  B   4      14
HET    NAG  B   5      14
HET    NAG  B   6      14
HET    SO4  B 492       5
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     SO4 SULFATE ION
FORMUL   2  NAG    6(C8 H15 N O6)
FORMUL   7  SO4    O4 S 2-
FORMUL   8  HOH   *76(H2 O)
HELIX    1   1 ASP B   95  THR B  103  1                                   9
HELIX    2   2 THR B  103  LYS B  114  1                                  12
HELIX    3   3 PHE B  131  LYS B  137  5                                   7
HELIX    4   4 THR B  146  ILE B  165  1                                  20
HELIX    5   5 GLY B  167  GLN B  171  5                                   5
HELIX    6   6 SER B  179  TYR B  192  1                                  14
HELIX    7   7 GLY B  216  LYS B  228  1                                  13
HELIX    8   8 HIS B  232  SER B  248  1                                  17
HELIX    9   9 THR B  250  LEU B  261  1                                  12
HELIX   10  10 ASP B  271  VAL B  289  1                                  19
HELIX   11  11 TRP B  304  LEU B  313  1                                  10
HELIX   12  12 SER B  319  TYR B  337  1                                  19
HELIX   13  13 SER B  353  GLU B  366  1                                  14
HELIX   14  14 ASN B  386  GLY B  399  1                                  14
HELIX   15  15 SER B  405  GLY B  412  1                                   8
HELIX   16  16 ASP B  430  GLY B  436  5                                   7
HELIX   17  17 HIS B  456  ARG B  460  5                                   5
HELIX   18  18 PRO B  467  ASP B  490  1                                  24
SHEET    1   A 8 SER B  49  LYS B  56  0
SHEET    2   A 8 THR B  67  ALA B  75 -1  O  TYR B  72   N  LEU B  51
SHEET    3   A 8 ALA B 115  ALA B 120 -1  O  LEU B 117   N  ALA B  75
SHEET    4   A 8 SER B  85  THR B  90  1  N  TYR B  89   O  VAL B 118
SHEET    5   A 8 VAL B 173  GLY B 178  1  O  ILE B 174   N  ILE B  86
SHEET    6   A 8 GLY B 198  ALA B 202  1  O  LEU B 200   N  ALA B 175
SHEET    7   A 8 ILE B 422  GLY B 427  1  O  VAL B 423   N  ALA B 199
SHEET    8   A 8 LEU B 445  ILE B 450  1  O  VAL B 446   N  PHE B 424
SHEET    1   B 2 SER B 295  ASN B 296  0
SHEET    2   B 2 PRO B 300  LEU B 301 -1  O  LEU B 301   N  SER B 295
SSBOND   1 CYS B  215    CYS B  372                          1555   1555  2.09
SSBOND   2 CYS B  233    CYS B  310                          1555   1555  2.05
SSBOND   3 CYS B  264    CYS B  343                          1555   1555  2.06
SSBOND   4 CYS B  364    CYS B  394                          1555   1555  2.05
LINK         ND2 ASN B 336                 C1  NAG B   3     1555   1555  1.42
LINK         ND2 ASN B 415                 C1  NAG B   6     1555   1555  1.44
LINK         ND2 ASN B 101                 C1  NAG B   1     1555   1555  1.45
LINK         O4  NAG B   3                 C1  NAG B   4     1555   1555  1.45
LINK         ND2 ASN B 345                 C1  NAG B   5     1555   1555  1.49
LINK         ND2 ASN B 317                 C1  NAG B   2     1555   1555  1.49
SITE     1 AC1  3 HOH B  21  ASN B 101  HOH B 517
SITE     1 AC2  3 ASN B 315  PRO B 316  ASN B 317
SITE     1 AC3  8 NAG B   4  HIS B 262  PHE B 328  ASN B 332
SITE     2 AC3  8 ASN B 336  GLY B 339  VAL B 341  LYS B 342
SITE     1 AC4  1 NAG B   3
SITE     1 AC5  3 ASN B 345  SER B 347  HOH B 520
SITE     1 AC6  4 HOH B   9  LYS B 162  ASN B 170  ASN B 415
SITE     1 AC7  4 LYS B  56  ARG B 403  PRO B 404  SER B 405
CRYST1  181.140  181.140  240.130  90.00  90.00 120.00 H 3 2        18
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.005521  0.003187  0.000000        0.00000
SCALE2      0.000000  0.006375  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004164        0.00000
TER    3540      SER B 491
MASTER      411    0    7   18   10    0    8    6 3704    1  102   35
END