longtext: 3O4H-pdb

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HEADER    HYDROLASE                               27-JUL-10   3O4H
TITLE     STRUCTURE AND CATALYSIS OF ACYLAMINOACYL PEPTIDASE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACYLAMINO-ACID-RELEASING ENZYME;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 SYNONYM: AARE, ACYL-PEPTIDE HYDROLASE, APH, ACYLAMINOACYL-PEPTIDASE;
COMPND   5 EC: 3.4.19.1;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: AEROPYRUM PERNIX;
SOURCE   3 ORGANISM_TAXID: 56636;
SOURCE   4 GENE: APE_1547.1;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   8 EXPRESSION_SYSTEM_PLASMID: PET22B
KEYWDS    ALPHA/BETA HYDROLASE FOLD, BETA PROPELLER, HYDROLASE, OLIGOPEPTIDASE,
KEYWDS   2 SIZE SELECTIVITY
EXPDTA    X-RAY DIFFRACTION
AUTHOR    V.HARMAT,K.DOMOKOS,D.K.MENYHARD,A.PALLO,Z.SZELTNER,I.SZAMOSI,T.BEKE-
AUTHOR   2 SOMFAI,G.NARAY-SZABO,L.POLGAR
REVDAT   1   17-NOV-10 3O4H    0
JRNL        AUTH   V.HARMAT,K.DOMOKOS,D.K.MENYHARD,A.PALLO,Z.SZELTNER,
JRNL        AUTH 2 I.SZAMOSI,T.BEKE-SOMFAI,G.NARAY-SZABO,L.POLGAR
JRNL        TITL   STRUCTURE AND CATALYSIS OF ACYLAMINOACYL PEPTIDASE: CLOSED
JRNL        TITL 2 AND OPEN SUBUNITS OF A DIMER OLIGOPEPTIDASE
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    1.82 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.5.0102
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.82
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 24.62
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 95.3
REMARK   3   NUMBER OF REFLECTIONS             : 196431
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.205
REMARK   3   R VALUE            (WORKING SET) : 0.203
REMARK   3   FREE R VALUE                     : 0.234
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 10388
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.82
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.87
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 14326
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 94.08
REMARK   3   BIN R VALUE           (WORKING SET) : 0.3300
REMARK   3   BIN FREE R VALUE SET COUNT          : 737
REMARK   3   BIN FREE R VALUE                    : 0.3460
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 17118
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 32
REMARK   3   SOLVENT ATOMS            : 928
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 36.26
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.00000
REMARK   3    B22 (A**2) : 0.09000
REMARK   3    B33 (A**2) : -0.03000
REMARK   3    B12 (A**2) : 0.03000
REMARK   3    B13 (A**2) : 0.04000
REMARK   3    B23 (A**2) : 0.06000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): NULL
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.134
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.118
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 8.964
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.960
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.947
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A): 17614 ; 0.019 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A): 11984 ; 0.008 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 23937 ; 1.722 ; 1.974
REMARK   3   BOND ANGLES OTHERS          (DEGREES): 29051 ; 1.426 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  2322 ; 6.335 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   692 ;33.218 ;22.442
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  2720 ;15.420 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):   152 ;21.006 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  2694 ; 0.098 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A): 19995 ; 0.008 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  3712 ; 0.006 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2): 11413 ; 0.775 ; 1.500
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  4756 ; 0.483 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2): 18228 ; 1.157 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  6201 ; 2.026 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  5699 ; 2.924 ; 4.500
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : 4
REMARK   3
REMARK   3  NCS GROUP NUMBER               : 1
REMARK   3     CHAIN NAMES                    : A C
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 7
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE
REMARK   3           1     A      6       A      10      3
REMARK   3           1     C      6       C      10      3
REMARK   3           2     A     12       A      21      3
REMARK   3           2     C     12       C      21      3
REMARK   3           3     A    322       A     344      3
REMARK   3           3     C    322       C     344      3
REMARK   3           4     A    346       A     430      3
REMARK   3           4     C    346       C     430      3
REMARK   3           5     A    432       A     434      3
REMARK   3           5     C    432       C     434      3
REMARK   3           6     A    436       A     541      3
REMARK   3           6     C    436       C     541      3
REMARK   3           7     A    543       A     580      3
REMARK   3           7     C    543       C     580      3
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT
REMARK   3   TIGHT POSITIONAL   1    A    (A):   1553 ; 0.060 ; 0.050
REMARK   3   LOOSE POSITIONAL   1    A    (A):   1739 ; 0.160 ; 5.000
REMARK   3   TIGHT THERMAL      1    A (A**2):   1553 ; 0.260 ; 0.500
REMARK   3   LOOSE THERMAL      1    A (A**2):   1739 ; 0.220 ;10.000
REMARK   3
REMARK   3  NCS GROUP NUMBER               : 2
REMARK   3     CHAIN NAMES                    : B D
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 6
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE
REMARK   3           1     B      7       B      21      3
REMARK   3           1     D      7       D      21      3
REMARK   3           2     B    322       B     341      3
REMARK   3           2     D    322       D     341      3
REMARK   3           3     B    343       B     344      3
REMARK   3           3     D    343       D     344      3
REMARK   3           4     B    346       B     427      3
REMARK   3           4     D    346       D     427      3
REMARK   3           5     B    429       B     430      3
REMARK   3           5     D    429       D     430      3
REMARK   3           6     B    433       B     579      3
REMARK   3           6     D    433       D     579      3
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT
REMARK   3   TIGHT POSITIONAL   2    B    (A):   1566 ; 0.070 ; 0.050
REMARK   3   LOOSE POSITIONAL   2    B    (A):   1745 ; 0.170 ; 5.000
REMARK   3   TIGHT THERMAL      2    B (A**2):   1566 ; 0.230 ; 0.500
REMARK   3   LOOSE THERMAL      2    B (A**2):   1745 ; 0.190 ;10.000
REMARK   3
REMARK   3  NCS GROUP NUMBER               : 3
REMARK   3     CHAIN NAMES                    : A C
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 2
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE
REMARK   3           1     A     27       A      40      3
REMARK   3           1     C     27       C      40      3
REMARK   3           2     A     42       A     317      3
REMARK   3           2     C     42       C     317      3
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT
REMARK   3   TIGHT POSITIONAL   3    A    (A):   1680 ; 0.060 ; 0.050
REMARK   3   LOOSE POSITIONAL   3    A    (A):   1814 ; 0.140 ; 5.000
REMARK   3   TIGHT THERMAL      3    A (A**2):   1680 ; 0.200 ; 0.500
REMARK   3   LOOSE THERMAL      3    A (A**2):   1814 ; 0.190 ;10.000
REMARK   3
REMARK   3  NCS GROUP NUMBER               : 4
REMARK   3     CHAIN NAMES                    : B D
REMARK   3     NUMBER OF COMPONENTS NCS GROUP : 4
REMARK   3       COMPONENT C  SSSEQI  TO  C   SSSEQI   CODE
REMARK   3           1     B     25       B      71      3
REMARK   3           1     D     25       D      71      3
REMARK   3           2     B     73       B      97      3
REMARK   3           2     D     73       D      97      3
REMARK   3           3     B     99       B     158      3
REMARK   3           3     D     99       D     158      3
REMARK   3           4     B    160       B     317      3
REMARK   3           4     D    160       D     317      3
REMARK   3                   GROUP CHAIN        COUNT   RMS     WEIGHT
REMARK   3   TIGHT POSITIONAL   4    B    (A):   1680 ; 0.100 ; 0.050
REMARK   3   LOOSE POSITIONAL   4    B    (A):   1793 ; 0.170 ; 5.000
REMARK   3   TIGHT THERMAL      4    B (A**2):   1680 ; 0.230 ; 0.500
REMARK   3   LOOSE THERMAL      4    B (A**2):   1793 ; 0.190 ;10.000
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 8
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 2
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     6        A    23
REMARK   3    RESIDUE RANGE :   A   319        A   581
REMARK   3    ORIGIN FOR THE GROUP (A): -16.1077  -6.6514  -0.0263
REMARK   3    T TENSOR
REMARK   3      T11:   0.1881 T22:   0.0898
REMARK   3      T33:   0.1993 T12:   0.1188
REMARK   3      T13:  -0.0993 T23:  -0.0528
REMARK   3    L TENSOR
REMARK   3      L11:   0.8710 L22:   1.1408
REMARK   3      L33:   1.3810 L12:   0.4496
REMARK   3      L13:   0.4164 L23:   0.0098
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1367 S12:  -0.1196 S13:   0.1605
REMARK   3      S21:  -0.0977 S22:  -0.0923 S23:   0.1540
REMARK   3      S31:  -0.3306 S32:  -0.2206 S33:   0.2291
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 2
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B     7        B    23
REMARK   3    RESIDUE RANGE :   B   319        B   580
REMARK   3    ORIGIN FOR THE GROUP (A):  16.1847   6.0457   2.3542
REMARK   3    T TENSOR
REMARK   3      T11:   0.2311 T22:   0.2671
REMARK   3      T33:   0.2174 T12:   0.1552
REMARK   3      T13:  -0.0811 T23:  -0.1262
REMARK   3    L TENSOR
REMARK   3      L11:   1.6888 L22:   1.3908
REMARK   3      L33:   2.9427 L12:   1.3034
REMARK   3      L13:   0.8425 L23:   1.1548
REMARK   3    S TENSOR
REMARK   3      S11:   0.1373 S12:   0.3967 S13:  -0.1774
REMARK   3      S21:  -0.0747 S22:   0.3300 S23:  -0.1340
REMARK   3      S31:   0.3112 S32:   0.6555 S33:  -0.4673
REMARK   3
REMARK   3   TLS GROUP : 3
REMARK   3    NUMBER OF COMPONENTS GROUP : 2
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   C     5        C    23
REMARK   3    RESIDUE RANGE :   C   319        C   581
REMARK   3    ORIGIN FOR THE GROUP (A): -18.5147  31.2688 -29.4688
REMARK   3    T TENSOR
REMARK   3      T11:   0.0400 T22:   0.0698
REMARK   3      T33:   0.1201 T12:   0.0112
REMARK   3      T13:   0.0358 T23:  -0.0058
REMARK   3    L TENSOR
REMARK   3      L11:   0.9555 L22:   3.0101
REMARK   3      L33:   2.1834 L12:   0.6262
REMARK   3      L13:   0.8288 L23:   0.5827
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0007 S12:   0.1874 S13:  -0.2160
REMARK   3      S21:   0.0715 S22:   0.1576 S23:  -0.1798
REMARK   3      S31:   0.0083 S32:   0.2972 S33:  -0.1570
REMARK   3
REMARK   3   TLS GROUP : 4
REMARK   3    NUMBER OF COMPONENTS GROUP : 2
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   D     7        D    23
REMARK   3    RESIDUE RANGE :   D   319        D   581
REMARK   3    ORIGIN FOR THE GROUP (A): -23.8525   2.9426 -49.5696
REMARK   3    T TENSOR
REMARK   3      T11:   0.0629 T22:   0.1121
REMARK   3      T33:   0.1885 T12:  -0.0038
REMARK   3      T13:   0.0538 T23:  -0.0506
REMARK   3    L TENSOR
REMARK   3      L11:   1.0119 L22:   3.2599
REMARK   3      L33:   2.6193 L12:   0.1335
REMARK   3      L13:   0.8112 L23:   1.9555
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0723 S12:  -0.0019 S13:   0.0392
REMARK   3      S21:   0.0289 S22:   0.3581 S23:  -0.4862
REMARK   3      S31:   0.0498 S32:   0.3254 S33:  -0.2858
REMARK   3
REMARK   3   TLS GROUP : 5
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A    24        A   318
REMARK   3    ORIGIN FOR THE GROUP (A): -12.4343 -34.1199  -2.1774
REMARK   3    T TENSOR
REMARK   3      T11:   0.0762 T22:   0.0521
REMARK   3      T33:   0.0860 T12:  -0.0032
REMARK   3      T13:   0.0302 T23:   0.0261
REMARK   3    L TENSOR
REMARK   3      L11:   0.7919 L22:   0.8188
REMARK   3      L33:   1.4908 L12:   0.2050
REMARK   3      L13:   0.1902 L23:   0.2134
REMARK   3    S TENSOR
REMARK   3      S11:   0.0184 S12:  -0.0775 S13:  -0.0039
REMARK   3      S21:  -0.1318 S22:  -0.0825 S23:   0.0523
REMARK   3      S31:   0.0654 S32:  -0.1502 S33:   0.0642
REMARK   3
REMARK   3   TLS GROUP : 6
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B    24        B   318
REMARK   3    ORIGIN FOR THE GROUP (A):  16.6059  32.6061  21.4787
REMARK   3    T TENSOR
REMARK   3      T11:   0.0715 T22:   0.1563
REMARK   3      T33:   0.0834 T12:  -0.0194
REMARK   3      T13:   0.0623 T23:  -0.0116
REMARK   3    L TENSOR
REMARK   3      L11:   1.3053 L22:   1.5333
REMARK   3      L33:   0.9326 L12:  -0.1577
REMARK   3      L13:   0.3020 L23:  -0.3468
REMARK   3    S TENSOR
REMARK   3      S11:   0.0573 S12:   0.1708 S13:  -0.0306
REMARK   3      S21:   0.0105 S22:  -0.0403 S23:   0.1297
REMARK   3      S31:  -0.0806 S32:   0.0987 S33:  -0.0169
REMARK   3
REMARK   3   TLS GROUP : 7
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   C    24        C   318
REMARK   3    ORIGIN FOR THE GROUP (A):   0.8813  46.5323 -43.0351
REMARK   3    T TENSOR
REMARK   3      T11:   0.1450 T22:   0.1835
REMARK   3      T33:   0.2742 T12:  -0.0511
REMARK   3      T13:  -0.0593 T23:   0.0486
REMARK   3    L TENSOR
REMARK   3      L11:   2.2174 L22:   1.6436
REMARK   3      L33:   1.7293 L12:   0.2116
REMARK   3      L13:   0.5260 L23:   0.2123
REMARK   3    S TENSOR
REMARK   3      S11:   0.2139 S12:   0.1949 S13:  -0.3610
REMARK   3      S21:   0.2212 S22:  -0.0918 S23:  -0.4789
REMARK   3      S31:   0.0681 S32:   0.3851 S33:  -0.1221
REMARK   3
REMARK   3   TLS GROUP : 8
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   D    24        D   318
REMARK   3    ORIGIN FOR THE GROUP (A): -55.1609  -5.3036 -48.2023
REMARK   3    T TENSOR
REMARK   3      T11:   0.1277 T22:   0.3122
REMARK   3      T33:   0.0973 T12:  -0.0627
REMARK   3      T13:   0.0704 T23:   0.0376
REMARK   3    L TENSOR
REMARK   3      L11:   1.7861 L22:   2.1145
REMARK   3      L33:   1.5290 L12:   0.1057
REMARK   3      L13:   0.1661 L23:   0.1583
REMARK   3    S TENSOR
REMARK   3      S11:   0.0097 S12:  -0.5459 S13:   0.0153
REMARK   3      S21:   0.3101 S22:  -0.0051 S23:   0.1806
REMARK   3      S31:   0.1309 S32:  -0.2821 S33:  -0.0047
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : BABINET MODEL WITH MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 3O4H COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 29-JUL-10.
REMARK 100 THE RCSB ID CODE IS RCSB060653.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 11-DEC-09
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.5
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : EMBL/DESY, HAMBURG
REMARK 200  BEAMLINE                       : X11
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.8148
REMARK 200  MONOCHROMATOR                  : DOUBLE CRYSTAL SI(111),
REMARK 200                                   HORIZONTALLY FOCUSSING
REMARK 200  OPTICS                         : MIRROR
REMARK 200
REMARK 200  DETECTOR TYPE                  : FLAT PANEL
REMARK 200  DETECTOR MANUFACTURER          : MAR555 FLAT PANEL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 206854
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.820
REMARK 200  RESOLUTION RANGE LOW       (A) : 30.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 95.2
REMARK 200  DATA REDUNDANCY                : 1.880
REMARK 200  R MERGE                    (I) : 0.02800
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 22.7200
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.82
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.87
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.1
REMARK 200  DATA REDUNDANCY IN SHELL       : 1.88
REMARK 200  R MERGE FOR SHELL          (I) : 0.60600
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.260
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: HYDROLASE AND PROPELLER DOMAINS OF PDB ENTRY 2HU5.
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 50.34
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.48
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 78MM SODIUM ACETATE, 0.44MM EDTA,
REMARK 280  6.7MM DITHIOTHREITOL, 2.4% PEG 4000 , PH 5.5, VAPOR DIFFUSION,
REMARK 280  HANGING DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 300 REMARK: THE BIOLOGICAL ASSEMBLY IS HOMODIMER. ONE DIMER CONSISTS OF
REMARK 300 CHAINS A AND B, AND THE OTHER DIMER CONSISTS OF CHAINS C AND D.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 3450 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 39700 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -30.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 3100 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 39930 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -31.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C, D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     ARG A     2
REMARK 465     ILE A     3
REMARK 465     ILE A     4
REMARK 465     MET A     5
REMARK 465     ARG A   582
REMARK 465     MET B     1
REMARK 465     ARG B     2
REMARK 465     ILE B     3
REMARK 465     ILE B     4
REMARK 465     MET B     5
REMARK 465     PRO B     6
REMARK 465     ARG B   581
REMARK 465     ARG B   582
REMARK 465     MET C     1
REMARK 465     ARG C     2
REMARK 465     ILE C     3
REMARK 465     ILE C     4
REMARK 465     ARG C   582
REMARK 465     MET D     1
REMARK 465     ARG D     2
REMARK 465     ILE D     3
REMARK 465     ILE D     4
REMARK 465     MET D     5
REMARK 465     PRO D     6
REMARK 465     ARG D   582
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A   8    OE1  OE2
REMARK 470     GLU A  43    CG   CD   OE1  OE2
REMARK 470     LYS A  85    CE   NZ
REMARK 470     LYS A  94    CD   CE   NZ
REMARK 470     GLU A 131    CD   OE1  OE2
REMARK 470     ARG A 216    CZ   NH1  NH2
REMARK 470     GLU A 217    CG   CD   OE1  OE2
REMARK 470     ARG A 226    CD   NE   CZ   NH1  NH2
REMARK 470     LYS A 238    CG   CD   CE   NZ
REMARK 470     ARG A 244    NE   CZ   NH1  NH2
REMARK 470     LYS A 294    CD   CE   NZ
REMARK 470     ARG A 327    CZ   NH1  NH2
REMARK 470     PHE A 341    CG   CD1  CD2  CE1  CE2  CZ
REMARK 470     GLU A 352    CG   CD   OE1  OE2
REMARK 470     GLU A 479    CG   CD   OE1  OE2
REMARK 470     LYS A 530    NZ
REMARK 470     ARG A 581    CG   CD   NE   CZ   NH1  NH2
REMARK 470     VAL B   7    CG1  CG2
REMARK 470     ASP B  52    CG   OD1  OD2
REMARK 470     GLU B  55    CG   CD   OE1  OE2
REMARK 470     ARG B  61    CZ   NH1  NH2
REMARK 470     LYS B  85    CG   CD   CE   NZ
REMARK 470     GLU B  88    CG   CD   OE1  OE2
REMARK 470     GLN B 104    CD   OE1  NE2
REMARK 470     ARG B 113    NH1  NH2
REMARK 470     GLU B 131    CG   CD   OE1  OE2
REMARK 470     ARG B 149    NE   CZ   NH1  NH2
REMARK 470     ARG B 174    CZ   NH1  NH2
REMARK 470     GLU B 194    CG   CD   OE1  OE2
REMARK 470     ARG B 216    CZ   NH1  NH2
REMARK 470     GLU B 217    CD   OE1  OE2
REMARK 470     GLU B 234    CG   CD   OE1  OE2
REMARK 470     ASP B 325    CG   OD1  OD2
REMARK 470     GLU B 406    CG   CD   OE1  OE2
REMARK 470     LYS B 410    CE   NZ
REMARK 470     ARG B 431    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS B 463    NZ
REMARK 470     ARG B 486    CD   NE   CZ   NH1  NH2
REMARK 470     ASN B 487    CG   OD1  ND2
REMARK 470     GLU B 490    CG   CD   OE1  OE2
REMARK 470     GLN B 491    CG   CD   OE1  NE2
REMARK 470     SER B 496    OG
REMARK 470     GLU B 498    CG   CD   OE1  OE2
REMARK 470     ILE B 499    CD1
REMARK 470     ARG B 501    CD   NE   CZ   NH1  NH2
REMARK 470     ASP B 510    CG   OD1  OD2
REMARK 470     ARG B 511    NE   CZ   NH1  NH2
REMARK 470     LYS B 513    CD   CE   NZ
REMARK 470     ARG B 526    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS B 530    CD   CE   NZ
REMARK 470     GLU B 538    CD   OE1  OE2
REMARK 470     ARG B 542    CZ   NH1  NH2
REMARK 470     ILE B 558    CD1
REMARK 470     ILE B 567    CD1
REMARK 470     GLU B 580    CB   CG   CD   OE1  OE2
REMARK 470     MET C   5    SD   CE
REMARK 470     GLU C   8    CG   CD   OE1  OE2
REMARK 470     ASP C  32    CG   OD1  OD2
REMARK 470     GLU C  43    CG   CD   OE1  OE2
REMARK 470     ARG C  61    CZ   NH1  NH2
REMARK 470     LYS C  85    CE   NZ
REMARK 470     GLN C  89    CD   OE1  NE2
REMARK 470     LYS C  94    CD   CE   NZ
REMARK 470     ARG C  99    CD   NE   CZ   NH1  NH2
REMARK 470     GLN C 104    CG   CD   OE1  NE2
REMARK 470     GLU C 107    CD   OE1  OE2
REMARK 470     LYS C 110    CG   CD   CE   NZ
REMARK 470     GLU C 131    CG   CD   OE1  OE2
REMARK 470     ARG C 133    CZ   NH1  NH2
REMARK 470     ARG C 149    CD   NE   CZ   NH1  NH2
REMARK 470     ARG C 188    NE   CZ   NH1  NH2
REMARK 470     LYS C 207    CE   NZ
REMARK 470     GLU C 217    CG   CD   OE1  OE2
REMARK 470     GLU C 234    CG   CD   OE1  OE2
REMARK 470     ARG C 244    CD   NE   CZ   NH1  NH2
REMARK 470     GLU C 279    OE1  OE2
REMARK 470     LYS C 294    NZ
REMARK 470     ARG C 328    CZ   NH1  NH2
REMARK 470     GLU C 432    CG   CD   OE1  OE2
REMARK 470     GLU C 479    CD   OE1  OE2
REMARK 470     LEU C 480    CG   CD1  CD2
REMARK 470     ARG C 486    NE   CZ   NH1  NH2
REMARK 470     ARG C 497    CZ   NH1  NH2
REMARK 470     GLU C 498    CG   CD   OE1  OE2
REMARK 470     ARG C 501    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS C 530    CG   CD   CE   NZ
REMARK 470     ILE C 551    CD1
REMARK 470     THR C 560    CG2
REMARK 470     GLU C 562    CG   CD   OE1  OE2
REMARK 470     ARG C 581    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS D  35    CE   NZ
REMARK 470     GLU D  55    CG   CD   OE1  OE2
REMARK 470     VAL D  83    CG1  CG2
REMARK 470     SER D  84    OG
REMARK 470     LYS D  85    CG   CD   CE   NZ
REMARK 470     GLU D  88    CG   CD   OE1  OE2
REMARK 470     GLN D  89    CG   CD   OE1  NE2
REMARK 470     LYS D  94    NZ
REMARK 470     ARG D  99    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS D 110    CG   CD   CE   NZ
REMARK 470     GLU D 131    CB   CG   CD   OE1  OE2
REMARK 470     ASP D 132    CG   OD1  OD2
REMARK 470     ARG D 133    CG   CD   NE   CZ   NH1  NH2
REMARK 470     VAL D 134    CG1  CG2
REMARK 470     ARG D 149    CG   CD   NE   CZ   NH1  NH2
REMARK 470     ASN D 181    CG   OD1  ND2
REMARK 470     GLU D 194    OE1  OE2
REMARK 470     ARG D 216    NE   CZ   NH1  NH2
REMARK 470     GLU D 324    CG   CD   OE1  OE2
REMARK 470     ARG D 328    CG   CD   NE   CZ   NH1  NH2
REMARK 470     GLU D 490    CG   CD   OE1  OE2
REMARK 470     GLN D 491    CD   OE1  NE2
REMARK 470     ARG D 511    CD   NE   CZ   NH1  NH2
REMARK 470     LYS D 513    CE   NZ
REMARK 470     SER D 525    OG
REMARK 470     LYS D 530    CE   NZ
REMARK 470     GLU D 580    CD   OE1  OE2
REMARK 470     ARG D 581    CG   CD   NE   CZ   NH1  NH2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   N    SER A   199     O    HOH A   840              2.04
REMARK 500   CE   MET C   561     CG2  VAL C   565              2.18
REMARK 500   O    HOH B   770     O    HOH B   771              2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    PHE A   9   CG    PHE A   9   CD2    -0.143
REMARK 500    PHE A   9   CG    PHE A   9   CD1    -0.152
REMARK 500    PHE A   9   CE1   PHE A   9   CZ     -0.215
REMARK 500    PHE A   9   CZ    PHE A   9   CE2    -0.239
REMARK 500    GLU A  17   CB    GLU A  17   CG     -0.222
REMARK 500    ASP A 140   CB    ASP A 140   CG     -0.152
REMARK 500    ARG A 268   CZ    ARG A 268   NH1    -0.132
REMARK 500    ARG A 268   CZ    ARG A 268   NH2    -0.138
REMARK 500    ARG A 287   CZ    ARG A 287   NH1    -0.183
REMARK 500    ARG A 292   CZ    ARG A 292   NH1    -0.139
REMARK 500    ARG A 292   CZ    ARG A 292   NH2    -0.132
REMARK 500    TYR A 444   CG    TYR A 444   CD2    -0.159
REMARK 500    TYR A 444   CG    TYR A 444   CD1    -0.174
REMARK 500    TYR A 444   CE1   TYR A 444   CZ     -0.191
REMARK 500    TYR A 444   CZ    TYR A 444   CE2    -0.202
REMARK 500    LEU A 455   CG    LEU A 455   CD1    -0.230
REMARK 500    ARG B 105   CZ    ARG B 105   NH1    -0.175
REMARK 500    ARG B 160   CZ    ARG B 160   NH1    -0.166
REMARK 500    ARG B 160   CZ    ARG B 160   NH2    -0.101
REMARK 500    ARG B 265   CZ    ARG B 265   NH1    -0.176
REMARK 500    ARG B 292   CZ    ARG B 292   NH1    -0.111
REMARK 500    ARG B 292   CZ    ARG B 292   NH2    -0.123
REMARK 500    GLU B 339   C     GLU B 339   O       0.149
REMARK 500    TYR B 444   CG    TYR B 444   CD2    -0.158
REMARK 500    TYR B 444   CG    TYR B 444   CD1    -0.166
REMARK 500    TYR B 444   CE1   TYR B 444   CZ     -0.188
REMARK 500    TYR B 444   CZ    TYR B 444   CE2    -0.198
REMARK 500    PHE B 485   CG    PHE B 485   CD2    -0.153
REMARK 500    PHE B 485   CG    PHE B 485   CD1    -0.133
REMARK 500    PHE B 485   CE1   PHE B 485   CZ     -0.266
REMARK 500    PHE B 485   CZ    PHE B 485   CE2    -0.232
REMARK 500    PHE C   9   CG    PHE C   9   CD2    -0.162
REMARK 500    PHE C   9   CG    PHE C   9   CD1    -0.161
REMARK 500    PHE C   9   CE1   PHE C   9   CZ     -0.230
REMARK 500    PHE C   9   CZ    PHE C   9   CE2    -0.221
REMARK 500    GLU C  17   CB    GLU C  17   CG     -0.215
REMARK 500    GLU C  55   CD    GLU C  55   OE1    -0.075
REMARK 500    GLU C  55   CD    GLU C  55   OE2    -0.072
REMARK 500    ARG C 268   CZ    ARG C 268   NH1    -0.123
REMARK 500    ARG C 268   CZ    ARG C 268   NH2    -0.131
REMARK 500    ARG C 287   CZ    ARG C 287   NH1    -0.167
REMARK 500    ARG C 292   CZ    ARG C 292   NH1    -0.132
REMARK 500    ARG C 292   CZ    ARG C 292   NH2    -0.120
REMARK 500    TYR C 444   CG    TYR C 444   CD2    -0.154
REMARK 500    TYR C 444   CG    TYR C 444   CD1    -0.152
REMARK 500    TYR C 444   CE1   TYR C 444   CZ     -0.185
REMARK 500    TYR C 444   CZ    TYR C 444   CE2    -0.186
REMARK 500    ARG D 105   CZ    ARG D 105   NH1    -0.165
REMARK 500    ARG D 160   CZ    ARG D 160   NH1    -0.142
REMARK 500    ARG D 160   CZ    ARG D 160   NH2    -0.088
REMARK 500
REMARK 500 THIS ENTRY HAS      66 BOND DEVIATIONS.
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ASP A  34   CB  -  CG  -  OD2 ANGL. DEV. =  -5.8 DEGREES
REMARK 500    ASP A  52   CB  -  CG  -  OD1 ANGL. DEV. =   6.2 DEGREES
REMARK 500    ARG A  61   NE  -  CZ  -  NH1 ANGL. DEV. =   5.1 DEGREES
REMARK 500    GLU A 102   OE1 -  CD  -  OE2 ANGL. DEV. =  -8.0 DEGREES
REMARK 500    ASP A 140   N   -  CA  -  CB  ANGL. DEV. = -15.9 DEGREES
REMARK 500    ASP A 140   CB  -  CG  -  OD1 ANGL. DEV. =   6.4 DEGREES
REMARK 500    ASP A 140   CB  -  CG  -  OD2 ANGL. DEV. = -11.0 DEGREES
REMARK 500    ARG A 268   NH1 -  CZ  -  NH2 ANGL. DEV. = -14.1 DEGREES
REMARK 500    ARG A 268   NE  -  CZ  -  NH1 ANGL. DEV. =   7.8 DEGREES
REMARK 500    ARG A 268   NE  -  CZ  -  NH2 ANGL. DEV. =   6.1 DEGREES
REMARK 500    ARG A 287   CG  -  CD  -  NE  ANGL. DEV. =  14.7 DEGREES
REMARK 500    ARG A 287   NH1 -  CZ  -  NH2 ANGL. DEV. =  -7.0 DEGREES
REMARK 500    ARG A 287   NE  -  CZ  -  NH1 ANGL. DEV. =  -4.7 DEGREES
REMARK 500    ARG A 287   NE  -  CZ  -  NH2 ANGL. DEV. =  11.0 DEGREES
REMARK 500    ARG A 292   NH1 -  CZ  -  NH2 ANGL. DEV. = -16.3 DEGREES
REMARK 500    ARG A 292   NE  -  CZ  -  NH1 ANGL. DEV. =   7.7 DEGREES
REMARK 500    ARG A 292   NE  -  CZ  -  NH2 ANGL. DEV. =   8.4 DEGREES
REMARK 500    TYR A 444   CB  -  CG  -  CD1 ANGL. DEV. =   3.6 DEGREES
REMARK 500    LEU A 492   CB  -  CG  -  CD2 ANGL. DEV. =  10.8 DEGREES
REMARK 500    GLU A 538   CA  -  CB  -  CG  ANGL. DEV. =  14.1 DEGREES
REMARK 500    GLU B  62   OE1 -  CD  -  OE2 ANGL. DEV. =  -8.9 DEGREES
REMARK 500    ARG B 105   NH1 -  CZ  -  NH2 ANGL. DEV. =  -8.6 DEGREES
REMARK 500    ARG B 105   NE  -  CZ  -  NH2 ANGL. DEV. =  10.9 DEGREES
REMARK 500    ARG B 160   NH1 -  CZ  -  NH2 ANGL. DEV. = -10.4 DEGREES
REMARK 500    ARG B 160   NE  -  CZ  -  NH1 ANGL. DEV. =   6.3 DEGREES
REMARK 500    ARG B 265   NE  -  CZ  -  NH1 ANGL. DEV. = -14.2 DEGREES
REMARK 500    ARG B 265   NE  -  CZ  -  NH2 ANGL. DEV. =  14.0 DEGREES
REMARK 500    ARG B 292   NH1 -  CZ  -  NH2 ANGL. DEV. = -11.7 DEGREES
REMARK 500    ARG B 292   NE  -  CZ  -  NH1 ANGL. DEV. =   8.3 DEGREES
REMARK 500    ARG B 292   NE  -  CZ  -  NH2 ANGL. DEV. =   3.3 DEGREES
REMARK 500    GLU B 315   OE1 -  CD  -  OE2 ANGL. DEV. =  -8.0 DEGREES
REMARK 500    ARG B 428   NE  -  CZ  -  NH1 ANGL. DEV. =   3.8 DEGREES
REMARK 500    LEU B 455   CB  -  CG  -  CD2 ANGL. DEV. =  13.1 DEGREES
REMARK 500    ARG B 497   NE  -  CZ  -  NH1 ANGL. DEV. =   3.2 DEGREES
REMARK 500    ASP B 553   CB  -  CG  -  OD1 ANGL. DEV. =   8.5 DEGREES
REMARK 500    LEU B 569   CB  -  CG  -  CD1 ANGL. DEV. =  14.7 DEGREES
REMARK 500    PRO C   6   C   -  N   -  CA  ANGL. DEV. =  11.8 DEGREES
REMARK 500    PRO C   6   C   -  N   -  CD  ANGL. DEV. = -14.0 DEGREES
REMARK 500    GLU C  55   OE1 -  CD  -  OE2 ANGL. DEV. =  -7.8 DEGREES
REMARK 500    GLU C 122   OE1 -  CD  -  OE2 ANGL. DEV. =   7.2 DEGREES
REMARK 500    GLU C 122   CG  -  CD  -  OE2 ANGL. DEV. = -15.5 DEGREES
REMARK 500    ASP C 140   CB  -  CG  -  OD2 ANGL. DEV. =  -8.7 DEGREES
REMARK 500    ARG C 268   NH1 -  CZ  -  NH2 ANGL. DEV. = -15.3 DEGREES
REMARK 500    ARG C 268   NE  -  CZ  -  NH1 ANGL. DEV. =   6.6 DEGREES
REMARK 500    ARG C 268   NE  -  CZ  -  NH2 ANGL. DEV. =   8.7 DEGREES
REMARK 500    ARG C 287   CG  -  CD  -  NE  ANGL. DEV. =  13.2 DEGREES
REMARK 500    ARG C 287   NH1 -  CZ  -  NH2 ANGL. DEV. =  -7.1 DEGREES
REMARK 500    ARG C 287   NE  -  CZ  -  NH1 ANGL. DEV. =  -6.0 DEGREES
REMARK 500    ARG C 287   NE  -  CZ  -  NH2 ANGL. DEV. =  12.5 DEGREES
REMARK 500    ARG C 292   NH1 -  CZ  -  NH2 ANGL. DEV. = -16.3 DEGREES
REMARK 500
REMARK 500 THIS ENTRY HAS      73 ANGLE DEVIATIONS.
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP A  32     -141.21     41.50
REMARK 500    ARG A  61      -82.13   -101.64
REMARK 500    ALA A 148      148.76   -171.77
REMARK 500    PRO A 151       44.93    -82.05
REMARK 500    THR A 214     -169.96   -101.63
REMARK 500    ARG A 216       10.19   -142.72
REMARK 500    SER A 445     -129.92     63.05
REMARK 500    ASP B  32     -139.74     59.21
REMARK 500    ASP B  34       45.78     79.51
REMARK 500    ASP B  52       50.30   -140.43
REMARK 500    ARG B  61      -83.40    -95.71
REMARK 500    ASP B 140     -159.96   -136.79
REMARK 500    ARG B 216     -113.42   -125.07
REMARK 500    ASP B 342      -15.91    -48.84
REMARK 500    ASP B 414       48.42   -140.56
REMARK 500    SER B 445     -130.34     65.46
REMARK 500    ASN B 523       39.67   -144.80
REMARK 500    ARG C  61      -81.73   -102.23
REMARK 500    ALA C 148      149.65   -170.64
REMARK 500    PRO C 151       46.81    -77.73
REMARK 500    THR C 214     -167.80   -103.03
REMARK 500    ASP C 414       45.04   -140.69
REMARK 500    SER C 445     -125.12     65.53
REMARK 500    ASP D  34       53.28   -170.81
REMARK 500    ARG D  61      -84.26    -93.82
REMARK 500    SER D  66     -177.65   -170.72
REMARK 500    GLU D  88       -6.67     66.69
REMARK 500    THR D 130     -169.12   -110.35
REMARK 500    ARG D 216     -112.21   -127.66
REMARK 500    SER D 340     -160.71   -107.95
REMARK 500    ASP D 342        3.55     53.73
REMARK 500    ASP D 414       48.44   -140.22
REMARK 500    SER D 445     -131.46     66.42
REMARK 500    ASN D 523       40.66   -146.22
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 VAL C   31     ASP C   32                 -148.82
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    ASP A 140        46.0      L          L   OUTSIDE RANGE
REMARK 500    ALA D  87        23.2      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA D 583  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH D 667   O
REMARK 620 2 HOH D 691   O    94.6
REMARK 620 3 HOH D 587   O    94.1 171.2
REMARK 620 4 ASP D 374   OD1  94.9  92.8  85.4
REMARK 620 5 HOH D 604   O   108.0  90.1  88.2 156.6
REMARK 620 6 HOH D 608   O   170.3  85.3  86.3  94.7  62.3
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA B 583  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH B 690   O
REMARK 620 2 HOH B 907   O    89.6
REMARK 620 3 HOH B 655   O    97.8 171.4
REMARK 620 4 ASP B 374   OD1  89.2  89.2  86.5
REMARK 620 5 HOH B 687   O   113.0  89.0  92.2 157.7
REMARK 620 6 HOH B 717   O   173.2  88.0  85.2  97.1  60.7
REMARK 620 N                    1     2     3     4     5
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 583
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 584
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA B 583
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL B 584
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA D 583
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL D 584
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL D 585
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3O4G   RELATED DB: PDB
REMARK 900 RELATED ID: 3O4I   RELATED DB: PDB
REMARK 900 RELATED ID: 3O4J   RELATED DB: PDB
DBREF  3O4H A    1   582  UNP    Q9YBQ2   APEH_AERPE       1    582
DBREF  3O4H B    1   582  UNP    Q9YBQ2   APEH_AERPE       1    582
DBREF  3O4H C    1   582  UNP    Q9YBQ2   APEH_AERPE       1    582
DBREF  3O4H D    1   582  UNP    Q9YBQ2   APEH_AERPE       1    582
SEQADV 3O4H ALA A  524  UNP  Q9YBQ2    ASP   524 ENGINEERED
SEQADV 3O4H ALA B  524  UNP  Q9YBQ2    ASP   524 ENGINEERED
SEQADV 3O4H ALA C  524  UNP  Q9YBQ2    ASP   524 ENGINEERED
SEQADV 3O4H ALA D  524  UNP  Q9YBQ2    ASP   524 ENGINEERED
SEQRES   1 A  582  MET ARG ILE ILE MET PRO VAL GLU PHE SER ARG ILE VAL
SEQRES   2 A  582  ARG ASP VAL GLU ARG LEU ILE ALA VAL GLU LYS TYR SER
SEQRES   3 A  582  LEU GLN GLY VAL VAL ASP GLY ASP LYS LEU LEU VAL VAL
SEQRES   4 A  582  GLY PHE SER GLU GLY SER VAL ASN ALA TYR LEU TYR ASP
SEQRES   5 A  582  GLY GLY GLU THR VAL LYS LEU ASN ARG GLU PRO ILE ASN
SEQRES   6 A  582  SER VAL LEU ASP PRO HIS TYR GLY VAL GLY ARG VAL ILE
SEQRES   7 A  582  LEU VAL ARG ASP VAL SER LYS GLY ALA GLU GLN HIS ALA
SEQRES   8 A  582  LEU PHE LYS VAL ASN THR SER ARG PRO GLY GLU GLU GLN
SEQRES   9 A  582  ARG LEU GLU ALA VAL LYS PRO MET ARG ILE LEU SER GLY
SEQRES  10 A  582  VAL ASP THR GLY GLU ALA VAL VAL PHE THR GLY ALA THR
SEQRES  11 A  582  GLU ASP ARG VAL ALA LEU TYR ALA LEU ASP GLY GLY GLY
SEQRES  12 A  582  LEU ARG GLU LEU ALA ARG LEU PRO GLY PHE GLY PHE VAL
SEQRES  13 A  582  SER ASP ILE ARG GLY ASP LEU ILE ALA GLY LEU GLY PHE
SEQRES  14 A  582  PHE GLY GLY GLY ARG VAL SER LEU PHE THR SER ASN LEU
SEQRES  15 A  582  SER SER GLY GLY LEU ARG VAL PHE ASP SER GLY GLU GLY
SEQRES  16 A  582  SER PHE SER SER ALA SER ILE SER PRO GLY MET LYS VAL
SEQRES  17 A  582  THR ALA GLY LEU GLU THR ALA ARG GLU ALA ARG LEU VAL
SEQRES  18 A  582  THR VAL ASP PRO ARG ASP GLY SER VAL GLU ASP LEU GLU
SEQRES  19 A  582  LEU PRO SER LYS ASP PHE SER SER TYR ARG PRO THR ALA
SEQRES  20 A  582  ILE THR TRP LEU GLY TYR LEU PRO ASP GLY ARG LEU ALA
SEQRES  21 A  582  VAL VAL ALA ARG ARG GLU GLY ARG SER ALA VAL PHE ILE
SEQRES  22 A  582  ASP GLY GLU ARG VAL GLU ALA PRO GLN GLY ASN HIS GLY
SEQRES  23 A  582  ARG VAL VAL LEU TRP ARG GLY LYS LEU VAL THR SER HIS
SEQRES  24 A  582  THR SER LEU SER THR PRO PRO ARG ILE VAL SER LEU PRO
SEQRES  25 A  582  SER GLY GLU PRO LEU LEU GLU GLY GLY LEU PRO GLU ASP
SEQRES  26 A  582  LEU ARG ARG SER ILE ALA GLY SER ARG LEU VAL TRP VAL
SEQRES  27 A  582  GLU SER PHE ASP GLY SER ARG VAL PRO THR TYR VAL LEU
SEQRES  28 A  582  GLU SER GLY ARG ALA PRO THR PRO GLY PRO THR VAL VAL
SEQRES  29 A  582  LEU VAL HIS GLY GLY PRO PHE ALA GLU ASP SER ASP SER
SEQRES  30 A  582  TRP ASP THR PHE ALA ALA SER LEU ALA ALA ALA GLY PHE
SEQRES  31 A  582  HIS VAL VAL MET PRO ASN TYR ARG GLY SER THR GLY TYR
SEQRES  32 A  582  GLY GLU GLU TRP ARG LEU LYS ILE ILE GLY ASP PRO CYS
SEQRES  33 A  582  GLY GLY GLU LEU GLU ASP VAL SER ALA ALA ALA ARG TRP
SEQRES  34 A  582  ALA ARG GLU SER GLY LEU ALA SER GLU LEU TYR ILE MET
SEQRES  35 A  582  GLY TYR SER TYR GLY GLY TYR MET THR LEU CYS ALA LEU
SEQRES  36 A  582  THR MET LYS PRO GLY LEU PHE LYS ALA GLY VAL ALA GLY
SEQRES  37 A  582  ALA SER VAL VAL ASP TRP GLU GLU MET TYR GLU LEU SER
SEQRES  38 A  582  ASP ALA ALA PHE ARG ASN PHE ILE GLU GLN LEU THR GLY
SEQRES  39 A  582  GLY SER ARG GLU ILE MET ARG SER ARG SER PRO ILE ASN
SEQRES  40 A  582  HIS VAL ASP ARG ILE LYS GLU PRO LEU ALA LEU ILE HIS
SEQRES  41 A  582  PRO GLN ASN ALA SER ARG THR PRO LEU LYS PRO LEU LEU
SEQRES  42 A  582  ARG LEU MET GLY GLU LEU LEU ALA ARG GLY LYS THR PHE
SEQRES  43 A  582  GLU ALA HIS ILE ILE PRO ASP ALA GLY HIS ALA ILE ASN
SEQRES  44 A  582  THR MET GLU ASP ALA VAL LYS ILE LEU LEU PRO ALA VAL
SEQRES  45 A  582  PHE PHE LEU ALA THR GLN ARG GLU ARG ARG
SEQRES   1 B  582  MET ARG ILE ILE MET PRO VAL GLU PHE SER ARG ILE VAL
SEQRES   2 B  582  ARG ASP VAL GLU ARG LEU ILE ALA VAL GLU LYS TYR SER
SEQRES   3 B  582  LEU GLN GLY VAL VAL ASP GLY ASP LYS LEU LEU VAL VAL
SEQRES   4 B  582  GLY PHE SER GLU GLY SER VAL ASN ALA TYR LEU TYR ASP
SEQRES   5 B  582  GLY GLY GLU THR VAL LYS LEU ASN ARG GLU PRO ILE ASN
SEQRES   6 B  582  SER VAL LEU ASP PRO HIS TYR GLY VAL GLY ARG VAL ILE
SEQRES   7 B  582  LEU VAL ARG ASP VAL SER LYS GLY ALA GLU GLN HIS ALA
SEQRES   8 B  582  LEU PHE LYS VAL ASN THR SER ARG PRO GLY GLU GLU GLN
SEQRES   9 B  582  ARG LEU GLU ALA VAL LYS PRO MET ARG ILE LEU SER GLY
SEQRES  10 B  582  VAL ASP THR GLY GLU ALA VAL VAL PHE THR GLY ALA THR
SEQRES  11 B  582  GLU ASP ARG VAL ALA LEU TYR ALA LEU ASP GLY GLY GLY
SEQRES  12 B  582  LEU ARG GLU LEU ALA ARG LEU PRO GLY PHE GLY PHE VAL
SEQRES  13 B  582  SER ASP ILE ARG GLY ASP LEU ILE ALA GLY LEU GLY PHE
SEQRES  14 B  582  PHE GLY GLY GLY ARG VAL SER LEU PHE THR SER ASN LEU
SEQRES  15 B  582  SER SER GLY GLY LEU ARG VAL PHE ASP SER GLY GLU GLY
SEQRES  16 B  582  SER PHE SER SER ALA SER ILE SER PRO GLY MET LYS VAL
SEQRES  17 B  582  THR ALA GLY LEU GLU THR ALA ARG GLU ALA ARG LEU VAL
SEQRES  18 B  582  THR VAL ASP PRO ARG ASP GLY SER VAL GLU ASP LEU GLU
SEQRES  19 B  582  LEU PRO SER LYS ASP PHE SER SER TYR ARG PRO THR ALA
SEQRES  20 B  582  ILE THR TRP LEU GLY TYR LEU PRO ASP GLY ARG LEU ALA
SEQRES  21 B  582  VAL VAL ALA ARG ARG GLU GLY ARG SER ALA VAL PHE ILE
SEQRES  22 B  582  ASP GLY GLU ARG VAL GLU ALA PRO GLN GLY ASN HIS GLY
SEQRES  23 B  582  ARG VAL VAL LEU TRP ARG GLY LYS LEU VAL THR SER HIS
SEQRES  24 B  582  THR SER LEU SER THR PRO PRO ARG ILE VAL SER LEU PRO
SEQRES  25 B  582  SER GLY GLU PRO LEU LEU GLU GLY GLY LEU PRO GLU ASP
SEQRES  26 B  582  LEU ARG ARG SER ILE ALA GLY SER ARG LEU VAL TRP VAL
SEQRES  27 B  582  GLU SER PHE ASP GLY SER ARG VAL PRO THR TYR VAL LEU
SEQRES  28 B  582  GLU SER GLY ARG ALA PRO THR PRO GLY PRO THR VAL VAL
SEQRES  29 B  582  LEU VAL HIS GLY GLY PRO PHE ALA GLU ASP SER ASP SER
SEQRES  30 B  582  TRP ASP THR PHE ALA ALA SER LEU ALA ALA ALA GLY PHE
SEQRES  31 B  582  HIS VAL VAL MET PRO ASN TYR ARG GLY SER THR GLY TYR
SEQRES  32 B  582  GLY GLU GLU TRP ARG LEU LYS ILE ILE GLY ASP PRO CYS
SEQRES  33 B  582  GLY GLY GLU LEU GLU ASP VAL SER ALA ALA ALA ARG TRP
SEQRES  34 B  582  ALA ARG GLU SER GLY LEU ALA SER GLU LEU TYR ILE MET
SEQRES  35 B  582  GLY TYR SER TYR GLY GLY TYR MET THR LEU CYS ALA LEU
SEQRES  36 B  582  THR MET LYS PRO GLY LEU PHE LYS ALA GLY VAL ALA GLY
SEQRES  37 B  582  ALA SER VAL VAL ASP TRP GLU GLU MET TYR GLU LEU SER
SEQRES  38 B  582  ASP ALA ALA PHE ARG ASN PHE ILE GLU GLN LEU THR GLY
SEQRES  39 B  582  GLY SER ARG GLU ILE MET ARG SER ARG SER PRO ILE ASN
SEQRES  40 B  582  HIS VAL ASP ARG ILE LYS GLU PRO LEU ALA LEU ILE HIS
SEQRES  41 B  582  PRO GLN ASN ALA SER ARG THR PRO LEU LYS PRO LEU LEU
SEQRES  42 B  582  ARG LEU MET GLY GLU LEU LEU ALA ARG GLY LYS THR PHE
SEQRES  43 B  582  GLU ALA HIS ILE ILE PRO ASP ALA GLY HIS ALA ILE ASN
SEQRES  44 B  582  THR MET GLU ASP ALA VAL LYS ILE LEU LEU PRO ALA VAL
SEQRES  45 B  582  PHE PHE LEU ALA THR GLN ARG GLU ARG ARG
SEQRES   1 C  582  MET ARG ILE ILE MET PRO VAL GLU PHE SER ARG ILE VAL
SEQRES   2 C  582  ARG ASP VAL GLU ARG LEU ILE ALA VAL GLU LYS TYR SER
SEQRES   3 C  582  LEU GLN GLY VAL VAL ASP GLY ASP LYS LEU LEU VAL VAL
SEQRES   4 C  582  GLY PHE SER GLU GLY SER VAL ASN ALA TYR LEU TYR ASP
SEQRES   5 C  582  GLY GLY GLU THR VAL LYS LEU ASN ARG GLU PRO ILE ASN
SEQRES   6 C  582  SER VAL LEU ASP PRO HIS TYR GLY VAL GLY ARG VAL ILE
SEQRES   7 C  582  LEU VAL ARG ASP VAL SER LYS GLY ALA GLU GLN HIS ALA
SEQRES   8 C  582  LEU PHE LYS VAL ASN THR SER ARG PRO GLY GLU GLU GLN
SEQRES   9 C  582  ARG LEU GLU ALA VAL LYS PRO MET ARG ILE LEU SER GLY
SEQRES  10 C  582  VAL ASP THR GLY GLU ALA VAL VAL PHE THR GLY ALA THR
SEQRES  11 C  582  GLU ASP ARG VAL ALA LEU TYR ALA LEU ASP GLY GLY GLY
SEQRES  12 C  582  LEU ARG GLU LEU ALA ARG LEU PRO GLY PHE GLY PHE VAL
SEQRES  13 C  582  SER ASP ILE ARG GLY ASP LEU ILE ALA GLY LEU GLY PHE
SEQRES  14 C  582  PHE GLY GLY GLY ARG VAL SER LEU PHE THR SER ASN LEU
SEQRES  15 C  582  SER SER GLY GLY LEU ARG VAL PHE ASP SER GLY GLU GLY
SEQRES  16 C  582  SER PHE SER SER ALA SER ILE SER PRO GLY MET LYS VAL
SEQRES  17 C  582  THR ALA GLY LEU GLU THR ALA ARG GLU ALA ARG LEU VAL
SEQRES  18 C  582  THR VAL ASP PRO ARG ASP GLY SER VAL GLU ASP LEU GLU
SEQRES  19 C  582  LEU PRO SER LYS ASP PHE SER SER TYR ARG PRO THR ALA
SEQRES  20 C  582  ILE THR TRP LEU GLY TYR LEU PRO ASP GLY ARG LEU ALA
SEQRES  21 C  582  VAL VAL ALA ARG ARG GLU GLY ARG SER ALA VAL PHE ILE
SEQRES  22 C  582  ASP GLY GLU ARG VAL GLU ALA PRO GLN GLY ASN HIS GLY
SEQRES  23 C  582  ARG VAL VAL LEU TRP ARG GLY LYS LEU VAL THR SER HIS
SEQRES  24 C  582  THR SER LEU SER THR PRO PRO ARG ILE VAL SER LEU PRO
SEQRES  25 C  582  SER GLY GLU PRO LEU LEU GLU GLY GLY LEU PRO GLU ASP
SEQRES  26 C  582  LEU ARG ARG SER ILE ALA GLY SER ARG LEU VAL TRP VAL
SEQRES  27 C  582  GLU SER PHE ASP GLY SER ARG VAL PRO THR TYR VAL LEU
SEQRES  28 C  582  GLU SER GLY ARG ALA PRO THR PRO GLY PRO THR VAL VAL
SEQRES  29 C  582  LEU VAL HIS GLY GLY PRO PHE ALA GLU ASP SER ASP SER
SEQRES  30 C  582  TRP ASP THR PHE ALA ALA SER LEU ALA ALA ALA GLY PHE
SEQRES  31 C  582  HIS VAL VAL MET PRO ASN TYR ARG GLY SER THR GLY TYR
SEQRES  32 C  582  GLY GLU GLU TRP ARG LEU LYS ILE ILE GLY ASP PRO CYS
SEQRES  33 C  582  GLY GLY GLU LEU GLU ASP VAL SER ALA ALA ALA ARG TRP
SEQRES  34 C  582  ALA ARG GLU SER GLY LEU ALA SER GLU LEU TYR ILE MET
SEQRES  35 C  582  GLY TYR SER TYR GLY GLY TYR MET THR LEU CYS ALA LEU
SEQRES  36 C  582  THR MET LYS PRO GLY LEU PHE LYS ALA GLY VAL ALA GLY
SEQRES  37 C  582  ALA SER VAL VAL ASP TRP GLU GLU MET TYR GLU LEU SER
SEQRES  38 C  582  ASP ALA ALA PHE ARG ASN PHE ILE GLU GLN LEU THR GLY
SEQRES  39 C  582  GLY SER ARG GLU ILE MET ARG SER ARG SER PRO ILE ASN
SEQRES  40 C  582  HIS VAL ASP ARG ILE LYS GLU PRO LEU ALA LEU ILE HIS
SEQRES  41 C  582  PRO GLN ASN ALA SER ARG THR PRO LEU LYS PRO LEU LEU
SEQRES  42 C  582  ARG LEU MET GLY GLU LEU LEU ALA ARG GLY LYS THR PHE
SEQRES  43 C  582  GLU ALA HIS ILE ILE PRO ASP ALA GLY HIS ALA ILE ASN
SEQRES  44 C  582  THR MET GLU ASP ALA VAL LYS ILE LEU LEU PRO ALA VAL
SEQRES  45 C  582  PHE PHE LEU ALA THR GLN ARG GLU ARG ARG
SEQRES   1 D  582  MET ARG ILE ILE MET PRO VAL GLU PHE SER ARG ILE VAL
SEQRES   2 D  582  ARG ASP VAL GLU ARG LEU ILE ALA VAL GLU LYS TYR SER
SEQRES   3 D  582  LEU GLN GLY VAL VAL ASP GLY ASP LYS LEU LEU VAL VAL
SEQRES   4 D  582  GLY PHE SER GLU GLY SER VAL ASN ALA TYR LEU TYR ASP
SEQRES   5 D  582  GLY GLY GLU THR VAL LYS LEU ASN ARG GLU PRO ILE ASN
SEQRES   6 D  582  SER VAL LEU ASP PRO HIS TYR GLY VAL GLY ARG VAL ILE
SEQRES   7 D  582  LEU VAL ARG ASP VAL SER LYS GLY ALA GLU GLN HIS ALA
SEQRES   8 D  582  LEU PHE LYS VAL ASN THR SER ARG PRO GLY GLU GLU GLN
SEQRES   9 D  582  ARG LEU GLU ALA VAL LYS PRO MET ARG ILE LEU SER GLY
SEQRES  10 D  582  VAL ASP THR GLY GLU ALA VAL VAL PHE THR GLY ALA THR
SEQRES  11 D  582  GLU ASP ARG VAL ALA LEU TYR ALA LEU ASP GLY GLY GLY
SEQRES  12 D  582  LEU ARG GLU LEU ALA ARG LEU PRO GLY PHE GLY PHE VAL
SEQRES  13 D  582  SER ASP ILE ARG GLY ASP LEU ILE ALA GLY LEU GLY PHE
SEQRES  14 D  582  PHE GLY GLY GLY ARG VAL SER LEU PHE THR SER ASN LEU
SEQRES  15 D  582  SER SER GLY GLY LEU ARG VAL PHE ASP SER GLY GLU GLY
SEQRES  16 D  582  SER PHE SER SER ALA SER ILE SER PRO GLY MET LYS VAL
SEQRES  17 D  582  THR ALA GLY LEU GLU THR ALA ARG GLU ALA ARG LEU VAL
SEQRES  18 D  582  THR VAL ASP PRO ARG ASP GLY SER VAL GLU ASP LEU GLU
SEQRES  19 D  582  LEU PRO SER LYS ASP PHE SER SER TYR ARG PRO THR ALA
SEQRES  20 D  582  ILE THR TRP LEU GLY TYR LEU PRO ASP GLY ARG LEU ALA
SEQRES  21 D  582  VAL VAL ALA ARG ARG GLU GLY ARG SER ALA VAL PHE ILE
SEQRES  22 D  582  ASP GLY GLU ARG VAL GLU ALA PRO GLN GLY ASN HIS GLY
SEQRES  23 D  582  ARG VAL VAL LEU TRP ARG GLY LYS LEU VAL THR SER HIS
SEQRES  24 D  582  THR SER LEU SER THR PRO PRO ARG ILE VAL SER LEU PRO
SEQRES  25 D  582  SER GLY GLU PRO LEU LEU GLU GLY GLY LEU PRO GLU ASP
SEQRES  26 D  582  LEU ARG ARG SER ILE ALA GLY SER ARG LEU VAL TRP VAL
SEQRES  27 D  582  GLU SER PHE ASP GLY SER ARG VAL PRO THR TYR VAL LEU
SEQRES  28 D  582  GLU SER GLY ARG ALA PRO THR PRO GLY PRO THR VAL VAL
SEQRES  29 D  582  LEU VAL HIS GLY GLY PRO PHE ALA GLU ASP SER ASP SER
SEQRES  30 D  582  TRP ASP THR PHE ALA ALA SER LEU ALA ALA ALA GLY PHE
SEQRES  31 D  582  HIS VAL VAL MET PRO ASN TYR ARG GLY SER THR GLY TYR
SEQRES  32 D  582  GLY GLU GLU TRP ARG LEU LYS ILE ILE GLY ASP PRO CYS
SEQRES  33 D  582  GLY GLY GLU LEU GLU ASP VAL SER ALA ALA ALA ARG TRP
SEQRES  34 D  582  ALA ARG GLU SER GLY LEU ALA SER GLU LEU TYR ILE MET
SEQRES  35 D  582  GLY TYR SER TYR GLY GLY TYR MET THR LEU CYS ALA LEU
SEQRES  36 D  582  THR MET LYS PRO GLY LEU PHE LYS ALA GLY VAL ALA GLY
SEQRES  37 D  582  ALA SER VAL VAL ASP TRP GLU GLU MET TYR GLU LEU SER
SEQRES  38 D  582  ASP ALA ALA PHE ARG ASN PHE ILE GLU GLN LEU THR GLY
SEQRES  39 D  582  GLY SER ARG GLU ILE MET ARG SER ARG SER PRO ILE ASN
SEQRES  40 D  582  HIS VAL ASP ARG ILE LYS GLU PRO LEU ALA LEU ILE HIS
SEQRES  41 D  582  PRO GLN ASN ALA SER ARG THR PRO LEU LYS PRO LEU LEU
SEQRES  42 D  582  ARG LEU MET GLY GLU LEU LEU ALA ARG GLY LYS THR PHE
SEQRES  43 D  582  GLU ALA HIS ILE ILE PRO ASP ALA GLY HIS ALA ILE ASN
SEQRES  44 D  582  THR MET GLU ASP ALA VAL LYS ILE LEU LEU PRO ALA VAL
SEQRES  45 D  582  PHE PHE LEU ALA THR GLN ARG GLU ARG ARG
HET    GOL  A 583      12
HET    GOL  A 584       6
HET     NA  B 583       1
HET    GOL  B 584       6
HET     NA  D 583       1
HET    GOL  D 584       6
HET    GOL  D 585       6
HETNAM     GOL GLYCEROL
HETNAM      NA SODIUM ION
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
FORMUL   5  GOL    5(C3 H8 O3)
FORMUL   7   NA    2(NA 1+)
FORMUL  12  HOH   *928(H2 O)
HELIX    1   1 GLU A    8  VAL A   22  1                                  15
HELIX    2   2 LYS A  238  ARG A  244  1                                   7
HELIX    3   3 PRO A  323  SER A  329  1                                   7
HELIX    4   4 ASP A  379  ALA A  388  1                                  10
HELIX    5   5 GLY A  404  LYS A  410  1                                   7
HELIX    6   6 GLY A  417  SER A  433  1                                  17
HELIX    7   7 SER A  445  LYS A  458  1                                  14
HELIX    8   8 ASP A  473  LEU A  480  1                                   8
HELIX    9   9 ASP A  482  THR A  493  1                                  12
HELIX   10  10 SER A  496  ARG A  503  1                                   8
HELIX   11  11 SER A  504  ILE A  512  5                                   9
HELIX   12  12 PRO A  528  ARG A  542  1                                  15
HELIX   13  13 THR A  560  GLU A  580  1                                  21
HELIX   14  14 GLU B    8  VAL B   22  1                                  15
HELIX   15  15 LYS B  238  ARG B  244  1                                   7
HELIX   16  16 PRO B  323  SER B  329  1                                   7
HELIX   17  17 ASP B  379  ALA B  388  1                                  10
HELIX   18  18 GLY B  404  LYS B  410  1                                   7
HELIX   19  19 GLY B  417  SER B  433  1                                  17
HELIX   20  20 SER B  445  LYS B  458  1                                  14
HELIX   21  21 ASP B  473  SER B  481  1                                   9
HELIX   22  22 ASP B  482  THR B  493  1                                  12
HELIX   23  23 SER B  496  ARG B  503  1                                   8
HELIX   24  24 SER B  504  ILE B  512  5                                   9
HELIX   25  25 LEU B  529  ARG B  542  1                                  14
HELIX   26  26 THR B  560  LEU B  568  1                                   9
HELIX   27  27 LEU B  568  GLU B  580  1                                  13
HELIX   28  28 GLU C    8  VAL C   22  1                                  15
HELIX   29  29 LYS C  238  ARG C  244  1                                   7
HELIX   30  30 PRO C  323  SER C  329  1                                   7
HELIX   31  31 ASP C  379  ALA C  388  1                                  10
HELIX   32  32 GLY C  404  LYS C  410  1                                   7
HELIX   33  33 GLY C  417  SER C  433  1                                  17
HELIX   34  34 SER C  445  LYS C  458  1                                  14
HELIX   35  35 ASP C  473  SER C  481  1                                   9
HELIX   36  36 ASP C  482  GLY C  494  1                                  13
HELIX   37  37 SER C  496  ARG C  503  1                                   8
HELIX   38  38 SER C  504  ILE C  512  5                                   9
HELIX   39  39 LEU C  529  ARG C  542  1                                  14
HELIX   40  40 THR C  560  ARG C  581  1                                  22
HELIX   41  41 GLU D    8  VAL D   22  1                                  15
HELIX   42  42 LYS D  238  ARG D  244  1                                   7
HELIX   43  43 PRO D  323  SER D  329  1                                   7
HELIX   44  44 PHE D  341  GLY D  343  5                                   3
HELIX   45  45 ASP D  379  ALA D  388  1                                  10
HELIX   46  46 GLY D  404  LYS D  410  1                                   7
HELIX   47  47 GLY D  417  SER D  433  1                                  17
HELIX   48  48 SER D  445  LYS D  458  1                                  14
HELIX   49  49 ASP D  473  SER D  481  1                                   9
HELIX   50  50 ASP D  482  THR D  493  1                                  12
HELIX   51  51 SER D  496  ARG D  503  1                                   8
HELIX   52  52 SER D  504  ILE D  512  5                                   9
HELIX   53  53 PRO D  528  ARG D  542  1                                  15
HELIX   54  54 THR D  560  LEU D  568  1                                   9
HELIX   55  55 LEU D  568  ARG D  581  1                                  14
SHEET    1   A 4 LYS A  24  VAL A  31  0
SHEET    2   A 4 LYS A  35  SER A  42 -1  O  PHE A  41   N  LYS A  24
SHEET    3   A 4 SER A  45  ASP A  52 -1  O  TYR A  49   N  VAL A  38
SHEET    4   A 4 GLU A  55  LYS A  58 -1  O  VAL A  57   N  LEU A  50
SHEET    1   B 4 SER A  66  VAL A  67  0
SHEET    2   B 4 ARG A  76  ASP A  82 -1  O  VAL A  80   N  SER A  66
SHEET    3   B 4 HIS A  90  ASN A  96 -1  O  PHE A  93   N  LEU A  79
SHEET    4   B 4 GLN A 104  ARG A 105 -1  O  GLN A 104   N  LYS A  94
SHEET    1   C 5 ASP A  69  PRO A  70  0
SHEET    2   C 5 ARG A 113  ASP A 119  1  O  ASP A 119   N  ASP A  69
SHEET    3   C 5 VAL A 124  ALA A 129 -1  O  VAL A 125   N  VAL A 118
SHEET    4   C 5 VAL A 134  ASP A 140 -1  O  TYR A 137   N  PHE A 126
SHEET    5   C 5 GLY A 143  LEU A 150 -1  O  ARG A 145   N  ALA A 138
SHEET    1   D 4 GLY A 154  ARG A 160  0
SHEET    2   D 4 LEU A 163  GLY A 171 -1  O  ALA A 165   N  ASP A 158
SHEET    3   D 4 ARG A 174  ASN A 181 -1  O  SER A 176   N  GLY A 168
SHEET    4   D 4 ARG A 188  PHE A 190 -1  O  PHE A 190   N  LEU A 177
SHEET    1   E 4 SER A 196  ILE A 202  0
SHEET    2   E 4 VAL A 208  GLU A 213 -1  O  GLU A 213   N  SER A 196
SHEET    3   E 4 ALA A 218  VAL A 223 -1  O  VAL A 223   N  VAL A 208
SHEET    4   E 4 VAL A 230  ASP A 232 -1  O  GLU A 231   N  THR A 222
SHEET    1   F 4 ALA A 247  TYR A 253  0
SHEET    2   F 4 LEU A 259  ARG A 265 -1  O  VAL A 262   N  TRP A 250
SHEET    3   F 4 ARG A 268  ILE A 273 -1  O  PHE A 272   N  VAL A 261
SHEET    4   F 4 GLU A 276  VAL A 278 -1  O  VAL A 278   N  VAL A 271
SHEET    1   G 4 ASN A 284  TRP A 291  0
SHEET    2   G 4 LYS A 294  SER A 301 -1  O  LYS A 294   N  TRP A 291
SHEET    3   G 4 THR A 304  LEU A 311 -1  O  VAL A 309   N  THR A 297
SHEET    4   G 4 PRO A 316  LEU A 318 -1  O  LEU A 318   N  ILE A 308
SHEET    1   H16 ILE A 330  GLU A 339  0
SHEET    2   H16 ARG A 345  SER A 353 -1  O  VAL A 346   N  VAL A 338
SHEET    3   H16 HIS A 391  PRO A 395 -1  O  MET A 394   N  TYR A 349
SHEET    4   H16 GLY A 360  VAL A 366  1  N  VAL A 363   O  HIS A 391
SHEET    5   H16 ALA A 436  TYR A 444  1  O  TYR A 440   N  VAL A 364
SHEET    6   H16 GLY A 465  GLY A 468  1  O  GLY A 468   N  GLY A 443
SHEET    7   H16 LEU A 516  PRO A 521  1  O  ALA A 517   N  ALA A 467
SHEET    8   H16 PHE A 546  ILE A 551  1  O  GLU A 547   N  LEU A 518
SHEET    9   H16 PHE B 546  ILE B 551 -1  O  ILE B 550   N  ALA A 548
SHEET   10   H16 LEU B 516  PRO B 521  1  N  HIS B 520   O  HIS B 549
SHEET   11   H16 ALA B 464  GLY B 468  1  N  ALA B 467   O  ALA B 517
SHEET   12   H16 ALA B 436  TYR B 444  1  N  GLY B 443   O  GLY B 468
SHEET   13   H16 GLY B 360  VAL B 366  1  N  VAL B 364   O  TYR B 440
SHEET   14   H16 HIS B 391  PRO B 395  1  O  HIS B 391   N  VAL B 363
SHEET   15   H16 ARG B 345  SER B 353 -1  N  TYR B 349   O  MET B 394
SHEET   16   H16 ILE B 330  GLU B 339 -1  N  VAL B 338   O  VAL B 346
SHEET    1   I 4 LYS B  24  VAL B  31  0
SHEET    2   I 4 LYS B  35  PHE B  41 -1  O  LEU B  37   N  GLN B  28
SHEET    3   I 4 VAL B  46  TYR B  51 -1  O  TYR B  49   N  VAL B  38
SHEET    4   I 4 THR B  56  LYS B  58 -1  O  VAL B  57   N  LEU B  50
SHEET    1   J 4 SER B  66  VAL B  67  0
SHEET    2   J 4 ARG B  76  ASP B  82 -1  O  VAL B  80   N  SER B  66
SHEET    3   J 4 HIS B  90  ASN B  96 -1  O  PHE B  93   N  LEU B  79
SHEET    4   J 4 GLU B 103  ARG B 105 -1  O  GLN B 104   N  LYS B  94
SHEET    1   K 5 ASP B  69  PRO B  70  0
SHEET    2   K 5 ARG B 113  ASP B 119  1  O  ASP B 119   N  ASP B  69
SHEET    3   K 5 VAL B 124  ALA B 129 -1  O  VAL B 125   N  VAL B 118
SHEET    4   K 5 VAL B 134  ASP B 140 -1  O  TYR B 137   N  PHE B 126
SHEET    5   K 5 GLY B 143  LEU B 150 -1  O  LEU B 150   N  VAL B 134
SHEET    1   L 4 GLY B 154  ARG B 160  0
SHEET    2   L 4 LEU B 163  GLY B 168 -1  O  ALA B 165   N  ASP B 158
SHEET    3   L 4 SER B 176  ASN B 181 -1  O  SER B 176   N  GLY B 168
SHEET    4   L 4 ARG B 188  PHE B 190 -1  O  PHE B 190   N  LEU B 177
SHEET    1   M 4 SER B 196  ILE B 202  0
SHEET    2   M 4 VAL B 208  GLU B 213 -1  O  GLU B 213   N  SER B 196
SHEET    3   M 4 ARG B 219  VAL B 223 -1  O  ARG B 219   N  LEU B 212
SHEET    4   M 4 VAL B 230  ASP B 232 -1  O  GLU B 231   N  THR B 222
SHEET    1   N 4 ALA B 247  TYR B 253  0
SHEET    2   N 4 LEU B 259  ARG B 265 -1  O  ARG B 264   N  ALA B 247
SHEET    3   N 4 ARG B 268  ILE B 273 -1  O  PHE B 272   N  VAL B 261
SHEET    4   N 4 GLU B 276  VAL B 278 -1  O  VAL B 278   N  VAL B 271
SHEET    1   O 4 ASN B 284  TRP B 291  0
SHEET    2   O 4 LYS B 294  SER B 301 -1  O  LYS B 294   N  TRP B 291
SHEET    3   O 4 THR B 304  LEU B 311 -1  O  LEU B 311   N  LEU B 295
SHEET    4   O 4 PRO B 316  LEU B 318 -1  O  LEU B 317   N  ILE B 308
SHEET    1   P 4 LYS C  24  VAL C  31  0
SHEET    2   P 4 LYS C  35  SER C  42 -1  O  VAL C  39   N  SER C  26
SHEET    3   P 4 SER C  45  ASP C  52 -1  O  TYR C  49   N  VAL C  38
SHEET    4   P 4 GLU C  55  LYS C  58 -1  O  GLU C  55   N  ASP C  52
SHEET    1   Q 4 SER C  66  VAL C  67  0
SHEET    2   Q 4 ARG C  76  ASP C  82 -1  O  VAL C  80   N  SER C  66
SHEET    3   Q 4 HIS C  90  ASN C  96 -1  O  PHE C  93   N  LEU C  79
SHEET    4   Q 4 GLN C 104  ARG C 105 -1  O  GLN C 104   N  LYS C  94
SHEET    1   R 5 ASP C  69  PRO C  70  0
SHEET    2   R 5 ARG C 113  ASP C 119  1  O  ASP C 119   N  ASP C  69
SHEET    3   R 5 VAL C 124  ALA C 129 -1  O  VAL C 125   N  VAL C 118
SHEET    4   R 5 VAL C 134  ASP C 140 -1  O  TYR C 137   N  PHE C 126
SHEET    5   R 5 GLY C 143  LEU C 150 -1  O  ARG C 145   N  ALA C 138
SHEET    1   S 4 GLY C 154  ARG C 160  0
SHEET    2   S 4 LEU C 163  GLY C 171 -1  O  ALA C 165   N  ASP C 158
SHEET    3   S 4 ARG C 174  ASN C 181 -1  O  SER C 176   N  GLY C 168
SHEET    4   S 4 ARG C 188  PHE C 190 -1  O  PHE C 190   N  LEU C 177
SHEET    1   T 4 GLY C 195  ILE C 202  0
SHEET    2   T 4 VAL C 208  THR C 214 -1  O  GLU C 213   N  SER C 196
SHEET    3   T 4 ALA C 218  VAL C 223 -1  O  VAL C 223   N  VAL C 208
SHEET    4   T 4 VAL C 230  ASP C 232 -1  O  GLU C 231   N  THR C 222
SHEET    1   U 4 ALA C 247  TYR C 253  0
SHEET    2   U 4 LEU C 259  ARG C 265 -1  O  VAL C 262   N  TRP C 250
SHEET    3   U 4 ARG C 268  ILE C 273 -1  O  PHE C 272   N  VAL C 261
SHEET    4   U 4 GLU C 276  VAL C 278 -1  O  VAL C 278   N  VAL C 271
SHEET    1   V 4 ASN C 284  TRP C 291  0
SHEET    2   V 4 LYS C 294  SER C 301 -1  O  LYS C 294   N  TRP C 291
SHEET    3   V 4 THR C 304  LEU C 311 -1  O  LEU C 311   N  LEU C 295
SHEET    4   V 4 PRO C 316  LEU C 318 -1  O  LEU C 318   N  ILE C 308
SHEET    1   W16 ILE C 330  GLU C 339  0
SHEET    2   W16 ARG C 345  SER C 353 -1  O  THR C 348   N  VAL C 336
SHEET    3   W16 HIS C 391  PRO C 395 -1  O  VAL C 392   N  LEU C 351
SHEET    4   W16 GLY C 360  VAL C 366  1  N  VAL C 363   O  HIS C 391
SHEET    5   W16 ALA C 436  TYR C 444  1  O  TYR C 440   N  VAL C 364
SHEET    6   W16 GLY C 465  GLY C 468  1  O  GLY C 468   N  GLY C 443
SHEET    7   W16 LEU C 516  PRO C 521  1  O  ALA C 517   N  ALA C 467
SHEET    8   W16 PHE C 546  ILE C 551  1  O  HIS C 549   N  HIS C 520
SHEET    9   W16 PHE D 546  ILE D 551 -1  O  ILE D 550   N  ALA C 548
SHEET   10   W16 LEU D 516  PRO D 521  1  N  HIS D 520   O  HIS D 549
SHEET   11   W16 ALA D 464  GLY D 468  1  N  ALA D 467   O  ALA D 517
SHEET   12   W16 ALA D 436  TYR D 444  1  N  GLY D 443   O  GLY D 468
SHEET   13   W16 GLY D 360  VAL D 366  1  N  VAL D 364   O  TYR D 440
SHEET   14   W16 HIS D 391  PRO D 395  1  O  HIS D 391   N  VAL D 363
SHEET   15   W16 ARG D 345  SER D 353 -1  N  TYR D 349   O  MET D 394
SHEET   16   W16 ILE D 330  GLU D 339 -1  N  VAL D 336   O  THR D 348
SHEET    1   X 4 LYS D  24  VAL D  31  0
SHEET    2   X 4 LYS D  35  PHE D  41 -1  O  LEU D  37   N  GLN D  28
SHEET    3   X 4 VAL D  46  ASP D  52 -1  O  TYR D  49   N  VAL D  38
SHEET    4   X 4 GLU D  55  LYS D  58 -1  O  VAL D  57   N  LEU D  50
SHEET    1   Y 4 SER D  66  VAL D  67  0
SHEET    2   Y 4 ARG D  76  ASP D  82 -1  O  VAL D  80   N  SER D  66
SHEET    3   Y 4 HIS D  90  ASN D  96 -1  O  PHE D  93   N  LEU D  79
SHEET    4   Y 4 GLU D 103  ARG D 105 -1  O  GLN D 104   N  LYS D  94
SHEET    1   Z 5 ASP D  69  PRO D  70  0
SHEET    2   Z 5 ARG D 113  ASP D 119  1  O  ASP D 119   N  ASP D  69
SHEET    3   Z 5 VAL D 124  ALA D 129 -1  O  VAL D 125   N  VAL D 118
SHEET    4   Z 5 VAL D 134  ASP D 140 -1  O  TYR D 137   N  PHE D 126
SHEET    5   Z 5 GLY D 143  LEU D 150 -1  O  ALA D 148   N  LEU D 136
SHEET    1  AA 4 GLY D 154  ARG D 160  0
SHEET    2  AA 4 LEU D 163  GLY D 168 -1  O  ALA D 165   N  ASP D 158
SHEET    3  AA 4 SER D 176  ASN D 181 -1  O  SER D 176   N  GLY D 168
SHEET    4  AA 4 ARG D 188  PHE D 190 -1  O  PHE D 190   N  LEU D 177
SHEET    1  AB 4 SER D 196  ILE D 202  0
SHEET    2  AB 4 VAL D 208  GLU D 213 -1  O  THR D 209   N  SER D 201
SHEET    3  AB 4 ARG D 219  VAL D 223 -1  O  ARG D 219   N  LEU D 212
SHEET    4  AB 4 VAL D 230  ASP D 232 -1  O  GLU D 231   N  THR D 222
SHEET    1  AC 4 ALA D 247  TYR D 253  0
SHEET    2  AC 4 LEU D 259  ARG D 265 -1  O  ALA D 260   N  GLY D 252
SHEET    3  AC 4 ARG D 268  ILE D 273 -1  O  PHE D 272   N  VAL D 261
SHEET    4  AC 4 GLU D 276  VAL D 278 -1  O  VAL D 278   N  VAL D 271
SHEET    1  AD 4 ASN D 284  TRP D 291  0
SHEET    2  AD 4 LYS D 294  SER D 301 -1  O  LYS D 294   N  TRP D 291
SHEET    3  AD 4 THR D 304  LEU D 311 -1  O  LEU D 311   N  LEU D 295
SHEET    4  AD 4 PRO D 316  LEU D 318 -1  O  LEU D 317   N  ILE D 308
LINK        NA    NA D 583                 O   HOH D 667     1555   1555  2.21
LINK        NA    NA B 583                 O   HOH B 690     1555   1555  2.28
LINK        NA    NA D 583                 O   HOH D 691     1555   1555  2.33
LINK        NA    NA B 583                 O   HOH B 907     1555   1555  2.36
LINK        NA    NA B 583                 O   HOH B 655     1555   1555  2.41
LINK        NA    NA D 583                 O   HOH D 587     1555   1555  2.53
LINK         OD1 ASP B 374                NA    NA B 583     1555   1555  2.53
LINK        NA    NA B 583                 O   HOH B 687     1555   1555  2.58
LINK         OD1 ASP D 374                NA    NA D 583     1555   1555  2.60
LINK        NA    NA D 583                 O   HOH D 604     1555   1555  2.70
LINK        NA    NA D 583                 O   HOH D 608     1555   1555  2.74
LINK        NA    NA B 583                 O   HOH B 717     1555   1555  2.76
CISPEP   1 LEU A  311    PRO A  312          0         4.45
CISPEP   2 THR A  358    PRO A  359          0        -0.42
CISPEP   3 GLY A  369    PRO A  370          0        12.22
CISPEP   4 LEU B  311    PRO B  312          0         5.11
CISPEP   5 THR B  358    PRO B  359          0         4.47
CISPEP   6 GLY B  369    PRO B  370          0         9.30
CISPEP   7 LEU C  311    PRO C  312          0         6.03
CISPEP   8 THR C  358    PRO C  359          0        -2.12
CISPEP   9 GLY C  369    PRO C  370          0        13.65
CISPEP  10 LEU D  311    PRO D  312          0         6.29
CISPEP  11 THR D  358    PRO D  359          0         2.79
CISPEP  12 GLY D  369    PRO D  370          0         9.83
SITE     1 AC1  6 GLY A 369  SER A 445  MET A 477  PHE A 488
SITE     2 AC1  6 ARG A 526  THR A 527
SITE     1 AC2  2 ARG A 345  TYR A 403
SITE     1 AC3  6 ASP B 374  HOH B 655  HOH B 687  HOH B 690
SITE     2 AC3  6 HOH B 717  HOH B 907
SITE     1 AC4  3 SER B 445  VAL B 471  PHE B 488
SITE     1 AC5  6 ASP D 374  HOH D 587  HOH D 604  HOH D 608
SITE     2 AC5  6 HOH D 667  HOH D 691
SITE     1 AC6  4 TYR D 444  ALA D 469  GLY D 555  HOH D 711
SITE     1 AC7  5 SER D 445  VAL D 471  PHE D 488  HOH D 713
SITE     2 AC7  5 HOH D 714
CRYST1   71.405   97.979   98.797 105.69 103.52 100.36 P 1           4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.014005  0.002560  0.004447        0.00000
SCALE2      0.000000  0.010375  0.003586        0.00000
SCALE3      0.000000  0.000000  0.011014        0.00000
TER    4351      ARG A 581
TER    8607      GLU B 580
TER   12917      ARG C 581
TER   17205      ARG D 581
MASTER      914    0    7   55  148    0   11    618078    4   52  180
END