longtext: 3PIC-pdb

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HEADER    HYDROLASE                               05-NOV-10   3PIC
TITLE     GLUCURONOYL ESTERASE CATALYTIC DOMAIN (CIP2_GE) FROM HYPOCREA JECORINA
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: CIP2;
COMPND   3 CHAIN: A, B, C;
COMPND   4 FRAGMENT: CATALYTIC DOMAIN (UNP RESIDUES 90-460)
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HYPOCREA JECORINA;
SOURCE   3 ORGANISM_COMMON: TRICHODERMA REESEI;
SOURCE   4 ORGANISM_TAXID: 51453
KEYWDS    ALPHA/BETA HYDROLASE FOLD, GLUCURONOYL ESTERASE, CARBOHYDRATE
KEYWDS   2 ESTERASE FAMILY 15 (CE-15), N-LINKED GLYCOSYLATION, SECRETED,
KEYWDS   3 HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    P.R.POKKULURI
REVDAT   3   20-JUL-11 3PIC    1       JRNL
REVDAT   2   22-JUN-11 3PIC    1       JRNL
REVDAT   1   01-JUN-11 3PIC    0
JRNL        AUTH   P.R.POKKULURI,N.E.DUKE,S.J.WOOD,M.A.COTTA,X.L.LI,P.BIELY,
JRNL        AUTH 2 M.SCHIFFER
JRNL        TITL   STRUCTURE OF THE CATALYTIC DOMAIN OF GLUCURONOYL ESTERASE
JRNL        TITL 2 CIP2 FROM HYPOCREA JECORINA.
JRNL        REF    PROTEINS                      V.  79  2588 2011
JRNL        REFN                   ISSN 0887-3585
JRNL        PMID   21661060
JRNL        DOI    10.1002/PROT.23088
REMARK   2
REMARK   2 RESOLUTION.    1.90 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.2
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.90
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 31.20
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 58390.550
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 90.1
REMARK   3   NUMBER OF REFLECTIONS             : 87710
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.195
REMARK   3   FREE R VALUE                     : 0.234
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 10.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 8748
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.003
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.90
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.02
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 85.30
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 12321
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2470
REMARK   3   BIN FREE R VALUE                    : 0.2920
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 9.80
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 1340
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.008
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 8066
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 42
REMARK   3   SOLVENT ATOMS            : 1113
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 9.40
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 15.00
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 1.67000
REMARK   3    B22 (A**2) : -3.51000
REMARK   3    B33 (A**2) : 1.85000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.21
REMARK   3   ESD FROM SIGMAA              (A) : 0.17
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.26
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.22
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.005
REMARK   3   BOND ANGLES            (DEGREES) : 1.20
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 23.70
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.76
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.040 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 1.470 ; 2.000
REMARK   3   SIDE-CHAIN BOND              (A**2) : 1.690 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 2.180 ; 2.500
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : FLAT MODEL
REMARK   3   KSOL        : 0.35
REMARK   3   BSOL        : 36.62
REMARK   3
REMARK   3  NCS MODEL : NONE
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM
REMARK   3  PARAMETER FILE  2  : WATER_REP.PARAM
REMARK   3  PARAMETER FILE  3  : CARBOHYDRATE.PARAM
REMARK   3  PARAMETER FILE  4  : NULL
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP
REMARK   3  TOPOLOGY FILE  2   : WATER.TOP
REMARK   3  TOPOLOGY FILE  3   : CARB.TOP
REMARK   3  TOPOLOGY FILE  4   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: BULK SOLVENT MODEL USED
REMARK   4
REMARK   4 3PIC COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 11-NOV-10.
REMARK 100 THE RCSB ID CODE IS RCSB062406.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 25-JUN-07
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.9
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 19-BM
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.91946
REMARK 200  MONOCHROMATOR                  : MONOCHROMATIC
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : CUSTOM-MADE
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 91361
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.900
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 92.0
REMARK 200  DATA REDUNDANCY                : 5.000
REMARK 200  R MERGE                    (I) : 0.08500
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 17.0000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.90
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.93
REMARK 200  COMPLETENESS FOR SHELL     (%) : 84.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 4.00
REMARK 200  R MERGE FOR SHELL          (I) : 0.30500
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 4.000
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: MAD
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MAD
REMARK 200 SOFTWARE USED: HKL-3000
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 52.09
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.57
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: QUIK SCREEN C4 (1.4 M SODIUM/POTASSIUM
REMARK 280  PHOSPHATE, PH 6.9), VAPOR DIFFUSION, HANGING DROP, TEMPERATURE
REMARK 280  298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       40.13000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       92.94500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       41.00500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       92.94500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       40.13000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       41.00500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 300 REMARK: MONOMER
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLN A    86
REMARK 465     GLN A    87
REMARK 465     THR A    88
REMARK 465     SER A    89
REMARK 465     GLY A    90
REMARK 465     ALA A    91
REMARK 465     GLY A    92
REMARK 465     GLY A    93
REMARK 465     ALA A    94
REMARK 465     THR A    95
REMARK 465     CYS A    96
REMARK 465     SER A    97
REMARK 465     ALA A    98
REMARK 465     GLN B    86
REMARK 465     GLN B    87
REMARK 465     THR B    88
REMARK 465     SER B    89
REMARK 465     GLY B    90
REMARK 465     ALA B    91
REMARK 465     GLY B    92
REMARK 465     GLY B    93
REMARK 465     ALA B    94
REMARK 465     GLN C    86
REMARK 465     GLN C    87
REMARK 465     THR C    88
REMARK 465     SER C    89
REMARK 465     GLY C    90
REMARK 465     ALA C    91
REMARK 465     GLY C    92
REMARK 465     GLY C    93
REMARK 465     ALA C    94
REMARK 465     THR C    95
REMARK 465     CYS C    96
REMARK 465     SER C    97
REMARK 465     ALA C    98
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     SER A 102    OG
REMARK 470     LYS A 122    CG   CD   CE   NZ
REMARK 470     LYS A 173    CG   CD   CE   NZ
REMARK 470     LYS A 292    CG   CD   CE   NZ
REMARK 470     LYS A 317    CG   CD   CE   NZ
REMARK 470     GLN A 393    CG   CD   OE1  NE2
REMARK 470     SER A 460    OG
REMARK 470     THR B  95    OG1  CG2
REMARK 470     LYS B 173    CG   CD   CE   NZ
REMARK 470     THR B 188    OG1  CG2
REMARK 470     ARG C 106    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS C 126    CG   CD   CE   NZ
REMARK 470     ASP C 127    CG   OD1  OD2
REMARK 470     LYS C 128    CG   CD   CE   NZ
REMARK 470     THR C 156    OG1  CG2
REMARK 470     LYS C 173    CG   CD   CE   NZ
REMARK 470     SER C 174    OG
REMARK 470     ILE C 181    CG1  CG2  CD1
REMARK 470     VAL C 208    CG1  CG2
REMARK 470     ASN C 212    CG   OD1
REMARK 470     LEU C 261    CG   CD1  CD2
REMARK 470     ARG C 266    CZ   NH1  NH2
REMARK 470     LYS C 271    CG   CD   CE   NZ
REMARK 470     LYS C 292    CG   CD   CE   NZ
REMARK 470     ASP C 399    CG   OD1  OD2
REMARK 470     HIS C 400    CG   ND1  CD2  CE1  NE2
REMARK 470     LEU C 431    CG   CD1  CD2
REMARK 470     GLN C 433    CG   CD   OE1  NE2
REMARK 470     SER C 434    OG
REMARK 470     GLN C 441    CG   CD   OE1  NE2
REMARK 470     GLN C 448    CG   CD   OE1  NE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    GLU A 144      -51.75   -121.93
REMARK 500    GLN A 221       46.03   -143.76
REMARK 500    SER A 278     -126.53     57.24
REMARK 500    PRO A 334       42.68    -83.96
REMARK 500    GLN A 346       45.46   -151.91
REMARK 500    PRO A 351       30.09    -77.68
REMARK 500    SER B 278     -127.57     58.21
REMARK 500    PRO B 334       41.67    -81.69
REMARK 500    GLN B 346       41.11   -142.33
REMARK 500    PRO B 351       27.01    -75.09
REMARK 500    PHE B 440      100.41   -162.92
REMARK 500    GLU C 144      -52.82   -120.36
REMARK 500    GLN C 221       53.28   -144.75
REMARK 500    SER C 278     -121.27     57.06
REMARK 500    PRO C 334       41.86    -81.07
REMARK 500    GLN C 346       49.77   -143.64
REMARK 500    PRO C 351       26.79    -78.07
REMARK 500    SER C 434       75.02   -110.35
REMARK 500    PHE C 440       81.11   -166.77
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 724        DISTANCE =  5.10 ANGSTROMS
REMARK 525    HOH A 728        DISTANCE =  6.25 ANGSTROMS
REMARK 525    HOH A 923        DISTANCE =  5.04 ANGSTROMS
REMARK 525    HOH B 693        DISTANCE =  5.05 ANGSTROMS
REMARK 525    HOH B 973        DISTANCE =  5.36 ANGSTROMS
REMARK 525    HOH B1017        DISTANCE =  6.98 ANGSTROMS
REMARK 525    HOH B1023        DISTANCE =  5.02 ANGSTROMS
REMARK 525    HOH B1100        DISTANCE =  5.26 ANGSTROMS
REMARK 525    HOH B1108        DISTANCE =  6.59 ANGSTROMS
REMARK 525    HOH B1112        DISTANCE =  5.33 ANGSTROMS
REMARK 525    HOH C 717        DISTANCE =  5.06 ANGSTROMS
REMARK 525    HOH C 725        DISTANCE =  7.39 ANGSTROMS
REMARK 525    HOH C 726        DISTANCE =  6.58 ANGSTROMS
REMARK 525    HOH C 729        DISTANCE =  5.30 ANGSTROMS
REMARK 525    HOH C 865        DISTANCE =  6.31 ANGSTROMS
REMARK 525    HOH C 968        DISTANCE =  5.22 ANGSTROMS
REMARK 525    HOH C1101        DISTANCE =  5.66 ANGSTROMS
REMARK 610
REMARK 610 MISSING HETEROATOM
REMARK 610 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 610 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 610 I=INSERTION CODE):
REMARK 610   M RES C SSEQI
REMARK 610     NAG A  500
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 500
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 500
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG C 500
DBREF  3PIC A   90   460  UNP    Q7Z9N1   Q7Z9N1_TRIRE    90    460
DBREF  3PIC B   90   460  UNP    Q7Z9N1   Q7Z9N1_TRIRE    90    460
DBREF  3PIC C   90   460  UNP    Q7Z9N1   Q7Z9N1_TRIRE    90    460
SEQADV 3PIC GLN A   86  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC GLN A   87  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC THR A   88  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC SER A   89  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC GLN B   86  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC GLN B   87  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC THR B   88  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC SER B   89  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC GLN C   86  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC GLN C   87  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC THR C   88  UNP  Q7Z9N1              CLONING ARTIFACT
SEQADV 3PIC SER C   89  UNP  Q7Z9N1              CLONING ARTIFACT
SEQRES   1 A  375  GLN GLN THR SER GLY ALA GLY GLY ALA THR CYS SER ALA
SEQRES   2 A  375  LEU PRO GLY SER ILE THR LEU ARG SER ASN ALA LYS LEU
SEQRES   3 A  375  ASN ASP LEU PHE THR MET PHE ASN GLY ASP LYS VAL THR
SEQRES   4 A  375  THR LYS ASP LYS PHE SER CYS ARG GLN ALA GLU MET SER
SEQRES   5 A  375  GLU LEU ILE GLN ARG TYR GLU LEU GLY THR LEU PRO GLY
SEQRES   6 A  375  ARG PRO SER THR LEU THR ALA SER PHE SER GLY ASN THR
SEQRES   7 A  375  LEU THR ILE ASN CYS GLY GLU ALA GLY LYS SER ILE SER
SEQRES   8 A  375  PHE THR VAL THR ILE THR TYR PRO SER SER GLY THR ALA
SEQRES   9 A  375  PRO TYR PRO ALA ILE ILE GLY TYR GLY GLY GLY SER LEU
SEQRES  10 A  375  PRO ALA PRO ALA GLY VAL ALA MET ILE ASN PHE ASN ASN
SEQRES  11 A  375  ASP ASN ILE ALA ALA GLN VAL ASN THR GLY SER ARG GLY
SEQRES  12 A  375  GLN GLY LYS PHE TYR ASP LEU TYR GLY SER SER HIS SER
SEQRES  13 A  375  ALA GLY ALA MET THR ALA TRP ALA TRP GLY VAL SER ARG
SEQRES  14 A  375  VAL ILE ASP ALA LEU GLU LEU VAL PRO GLY ALA ARG ILE
SEQRES  15 A  375  ASP THR THR LYS ILE GLY VAL THR GLY CYS SER ARG ASN
SEQRES  16 A  375  GLY LYS GLY ALA MET VAL ALA GLY ALA PHE GLU LYS ARG
SEQRES  17 A  375  ILE VAL LEU THR LEU PRO GLN GLU SER GLY ALA GLY GLY
SEQRES  18 A  375  SER ALA CYS TRP ARG ILE SER ASP TYR LEU LYS SER GLN
SEQRES  19 A  375  GLY ALA ASN ILE GLN THR ALA SER GLU ILE ILE GLY GLU
SEQRES  20 A  375  ASP PRO TRP PHE SER THR THR PHE ASN SER TYR VAL ASN
SEQRES  21 A  375  GLN VAL PRO VAL LEU PRO PHE ASP HIS HIS SER LEU ALA
SEQRES  22 A  375  ALA LEU ILE ALA PRO ARG GLY LEU PHE VAL ILE ASP ASN
SEQRES  23 A  375  ASN ILE ASP TRP LEU GLY PRO GLN SER CYS PHE GLY CYS
SEQRES  24 A  375  MET THR ALA ALA HIS MET ALA TRP GLN ALA LEU GLY VAL
SEQRES  25 A  375  SER ASP HIS MET GLY TYR SER GLN ILE GLY ALA HIS ALA
SEQRES  26 A  375  HIS CYS ALA PHE PRO SER ASN GLN GLN SER GLN LEU THR
SEQRES  27 A  375  ALA PHE VAL GLN LYS PHE LEU LEU GLY GLN SER THR ASN
SEQRES  28 A  375  THR ALA ILE PHE GLN SER ASP PHE SER ALA ASN GLN SER
SEQRES  29 A  375  GLN TRP ILE ASP TRP THR THR PRO THR LEU SER
SEQRES   1 B  375  GLN GLN THR SER GLY ALA GLY GLY ALA THR CYS SER ALA
SEQRES   2 B  375  LEU PRO GLY SER ILE THR LEU ARG SER ASN ALA LYS LEU
SEQRES   3 B  375  ASN ASP LEU PHE THR MET PHE ASN GLY ASP LYS VAL THR
SEQRES   4 B  375  THR LYS ASP LYS PHE SER CYS ARG GLN ALA GLU MET SER
SEQRES   5 B  375  GLU LEU ILE GLN ARG TYR GLU LEU GLY THR LEU PRO GLY
SEQRES   6 B  375  ARG PRO SER THR LEU THR ALA SER PHE SER GLY ASN THR
SEQRES   7 B  375  LEU THR ILE ASN CYS GLY GLU ALA GLY LYS SER ILE SER
SEQRES   8 B  375  PHE THR VAL THR ILE THR TYR PRO SER SER GLY THR ALA
SEQRES   9 B  375  PRO TYR PRO ALA ILE ILE GLY TYR GLY GLY GLY SER LEU
SEQRES  10 B  375  PRO ALA PRO ALA GLY VAL ALA MET ILE ASN PHE ASN ASN
SEQRES  11 B  375  ASP ASN ILE ALA ALA GLN VAL ASN THR GLY SER ARG GLY
SEQRES  12 B  375  GLN GLY LYS PHE TYR ASP LEU TYR GLY SER SER HIS SER
SEQRES  13 B  375  ALA GLY ALA MET THR ALA TRP ALA TRP GLY VAL SER ARG
SEQRES  14 B  375  VAL ILE ASP ALA LEU GLU LEU VAL PRO GLY ALA ARG ILE
SEQRES  15 B  375  ASP THR THR LYS ILE GLY VAL THR GLY CYS SER ARG ASN
SEQRES  16 B  375  GLY LYS GLY ALA MET VAL ALA GLY ALA PHE GLU LYS ARG
SEQRES  17 B  375  ILE VAL LEU THR LEU PRO GLN GLU SER GLY ALA GLY GLY
SEQRES  18 B  375  SER ALA CYS TRP ARG ILE SER ASP TYR LEU LYS SER GLN
SEQRES  19 B  375  GLY ALA ASN ILE GLN THR ALA SER GLU ILE ILE GLY GLU
SEQRES  20 B  375  ASP PRO TRP PHE SER THR THR PHE ASN SER TYR VAL ASN
SEQRES  21 B  375  GLN VAL PRO VAL LEU PRO PHE ASP HIS HIS SER LEU ALA
SEQRES  22 B  375  ALA LEU ILE ALA PRO ARG GLY LEU PHE VAL ILE ASP ASN
SEQRES  23 B  375  ASN ILE ASP TRP LEU GLY PRO GLN SER CYS PHE GLY CYS
SEQRES  24 B  375  MET THR ALA ALA HIS MET ALA TRP GLN ALA LEU GLY VAL
SEQRES  25 B  375  SER ASP HIS MET GLY TYR SER GLN ILE GLY ALA HIS ALA
SEQRES  26 B  375  HIS CYS ALA PHE PRO SER ASN GLN GLN SER GLN LEU THR
SEQRES  27 B  375  ALA PHE VAL GLN LYS PHE LEU LEU GLY GLN SER THR ASN
SEQRES  28 B  375  THR ALA ILE PHE GLN SER ASP PHE SER ALA ASN GLN SER
SEQRES  29 B  375  GLN TRP ILE ASP TRP THR THR PRO THR LEU SER
SEQRES   1 C  375  GLN GLN THR SER GLY ALA GLY GLY ALA THR CYS SER ALA
SEQRES   2 C  375  LEU PRO GLY SER ILE THR LEU ARG SER ASN ALA LYS LEU
SEQRES   3 C  375  ASN ASP LEU PHE THR MET PHE ASN GLY ASP LYS VAL THR
SEQRES   4 C  375  THR LYS ASP LYS PHE SER CYS ARG GLN ALA GLU MET SER
SEQRES   5 C  375  GLU LEU ILE GLN ARG TYR GLU LEU GLY THR LEU PRO GLY
SEQRES   6 C  375  ARG PRO SER THR LEU THR ALA SER PHE SER GLY ASN THR
SEQRES   7 C  375  LEU THR ILE ASN CYS GLY GLU ALA GLY LYS SER ILE SER
SEQRES   8 C  375  PHE THR VAL THR ILE THR TYR PRO SER SER GLY THR ALA
SEQRES   9 C  375  PRO TYR PRO ALA ILE ILE GLY TYR GLY GLY GLY SER LEU
SEQRES  10 C  375  PRO ALA PRO ALA GLY VAL ALA MET ILE ASN PHE ASN ASN
SEQRES  11 C  375  ASP ASN ILE ALA ALA GLN VAL ASN THR GLY SER ARG GLY
SEQRES  12 C  375  GLN GLY LYS PHE TYR ASP LEU TYR GLY SER SER HIS SER
SEQRES  13 C  375  ALA GLY ALA MET THR ALA TRP ALA TRP GLY VAL SER ARG
SEQRES  14 C  375  VAL ILE ASP ALA LEU GLU LEU VAL PRO GLY ALA ARG ILE
SEQRES  15 C  375  ASP THR THR LYS ILE GLY VAL THR GLY CYS SER ARG ASN
SEQRES  16 C  375  GLY LYS GLY ALA MET VAL ALA GLY ALA PHE GLU LYS ARG
SEQRES  17 C  375  ILE VAL LEU THR LEU PRO GLN GLU SER GLY ALA GLY GLY
SEQRES  18 C  375  SER ALA CYS TRP ARG ILE SER ASP TYR LEU LYS SER GLN
SEQRES  19 C  375  GLY ALA ASN ILE GLN THR ALA SER GLU ILE ILE GLY GLU
SEQRES  20 C  375  ASP PRO TRP PHE SER THR THR PHE ASN SER TYR VAL ASN
SEQRES  21 C  375  GLN VAL PRO VAL LEU PRO PHE ASP HIS HIS SER LEU ALA
SEQRES  22 C  375  ALA LEU ILE ALA PRO ARG GLY LEU PHE VAL ILE ASP ASN
SEQRES  23 C  375  ASN ILE ASP TRP LEU GLY PRO GLN SER CYS PHE GLY CYS
SEQRES  24 C  375  MET THR ALA ALA HIS MET ALA TRP GLN ALA LEU GLY VAL
SEQRES  25 C  375  SER ASP HIS MET GLY TYR SER GLN ILE GLY ALA HIS ALA
SEQRES  26 C  375  HIS CYS ALA PHE PRO SER ASN GLN GLN SER GLN LEU THR
SEQRES  27 C  375  ALA PHE VAL GLN LYS PHE LEU LEU GLY GLN SER THR ASN
SEQRES  28 C  375  THR ALA ILE PHE GLN SER ASP PHE SER ALA ASN GLN SER
SEQRES  29 C  375  GLN TRP ILE ASP TRP THR THR PRO THR LEU SER
MODRES 3PIC ASN C  447  ASN  GLYCOSYLATION SITE
MODRES 3PIC ASN B  447  ASN  GLYCOSYLATION SITE
HET    NAG  A 500      14
HET    NAG  B 500      14
HET    NAG  C 500      14
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   4  NAG    3(C8 H15 N O6)
FORMUL   7  HOH   *1113(H2 O)
HELIX    1   1 THR A  125  GLU A  144  1                                  20
HELIX    2   2 ASN A  214  ALA A  219  1                                   6
HELIX    3   3 ASN A  223  ARG A  227  5                                   5
HELIX    4   4 GLY A  230  GLY A  237  1                                   8
HELIX    5   5 GLY A  243  VAL A  262  1                                  20
HELIX    6   6 PRO A  263  ALA A  265  5                                   3
HELIX    7   7 SER A  278  GLU A  291  1                                  14
HELIX    8   8 CYS A  309  GLN A  319  1                                  11
HELIX    9   9 THR A  325  ILE A  330  1                                   6
HELIX   10  10 SER A  337  TYR A  343  5                                   7
HELIX   11  11 GLN A  346  LEU A  350  5                                   5
HELIX   12  12 ASP A  353  LEU A  360  1                                   8
HELIX   13  13 ILE A  373  LEU A  376  5                                   4
HELIX   14  14 GLY A  377  LEU A  395  1                                  19
HELIX   15  15 VAL A  397  ASP A  399  5                                   3
HELIX   16  16 PRO A  415  ASN A  417  5                                   3
HELIX   17  17 GLN A  418  LEU A  430  1                                  13
HELIX   18  18 ASN A  447  ILE A  452  1                                   6
HELIX   19  19 THR B  125  GLU B  144  1                                  20
HELIX   20  20 ASN B  214  ALA B  219  1                                   6
HELIX   21  21 ASN B  223  ARG B  227  5                                   5
HELIX   22  22 GLY B  230  GLY B  237  1                                   8
HELIX   23  23 GLY B  243  VAL B  262  1                                  20
HELIX   24  24 PRO B  263  ALA B  265  5                                   3
HELIX   25  25 ASN B  280  GLU B  291  1                                  12
HELIX   26  26 CYS B  309  GLN B  319  1                                  11
HELIX   27  27 THR B  325  ILE B  330  1                                   6
HELIX   28  28 SER B  337  TYR B  343  5                                   7
HELIX   29  29 GLN B  346  LEU B  350  5                                   5
HELIX   30  30 ASP B  353  LEU B  360  1                                   8
HELIX   31  31 ILE B  373  LEU B  376  5                                   4
HELIX   32  32 GLY B  377  LEU B  395  1                                  19
HELIX   33  33 VAL B  397  ASP B  399  5                                   3
HELIX   34  34 PRO B  415  ASN B  417  5                                   3
HELIX   35  35 GLN B  418  LEU B  430  1                                  13
HELIX   36  36 ASN B  447  ILE B  452  1                                   6
HELIX   37  37 THR C  125  GLU C  144  1                                  20
HELIX   38  38 ASN C  214  ALA C  219  1                                   6
HELIX   39  39 ASN C  223  ARG C  227  5                                   5
HELIX   40  40 GLY C  230  GLY C  237  1                                   8
HELIX   41  41 GLY C  243  VAL C  262  1                                  20
HELIX   42  42 PRO C  263  ALA C  265  5                                   3
HELIX   43  43 SER C  278  GLU C  291  1                                  14
HELIX   44  44 CYS C  309  GLY C  320  1                                  12
HELIX   45  45 THR C  325  ILE C  330  1                                   6
HELIX   46  46 SER C  337  TYR C  343  5                                   7
HELIX   47  47 GLN C  346  LEU C  350  5                                   5
HELIX   48  48 ASP C  353  LEU C  360  1                                   8
HELIX   49  49 ILE C  373  LEU C  376  5                                   4
HELIX   50  50 GLY C  377  LEU C  395  1                                  19
HELIX   51  51 VAL C  397  ASP C  399  5                                   3
HELIX   52  52 PRO C  415  ASN C  417  5                                   3
HELIX   53  53 GLN C  418  LEU C  430  1                                  13
HELIX   54  54 ASN C  447  ILE C  452  1                                   6
SHEET    1   A10 THR A 154  SER A 160  0
SHEET    2   A10 THR A 163  GLU A 170 -1  O  ASN A 167   N  THR A 156
SHEET    3   A10 LYS A 173  THR A 182 -1  O  PHE A 177   N  ILE A 166
SHEET    4   A10 ALA A 209  PHE A 213 -1  O  ASN A 212   N  THR A 180
SHEET    5   A10 TYR A 191  TYR A 197  1  N  GLY A 196   O  ILE A 211
SHEET    6   A10 ILE A 267  CYS A 277  1  O  ASP A 268   N  TYR A 191
SHEET    7   A10 ILE A 294  GLN A 300  1  O  LEU A 298   N  VAL A 274
SHEET    8   A10 GLY A 365  ILE A 369  1  O  PHE A 367   N  THR A 297
SHEET    9   A10 MET A 401  SER A 404  1  O  GLY A 402   N  LEU A 366
SHEET   10   A10 PHE A 440  GLN A 441  1  O  GLN A 441   N  TYR A 403
SHEET    1   B10 THR B 154  SER B 160  0
SHEET    2   B10 THR B 163  GLU B 170 -1  O  ASN B 167   N  THR B 156
SHEET    3   B10 LYS B 173  THR B 182 -1  O  VAL B 179   N  LEU B 164
SHEET    4   B10 ALA B 209  PHE B 213 -1  O  MET B 210   N  THR B 182
SHEET    5   B10 TYR B 191  TYR B 197  1  N  PRO B 192   O  ALA B 209
SHEET    6   B10 ILE B 267  CYS B 277  1  O  GLY B 273   N  ALA B 193
SHEET    7   B10 ILE B 294  GLN B 300  1  O  LEU B 298   N  VAL B 274
SHEET    8   B10 GLY B 365  ILE B 369  1  O  PHE B 367   N  THR B 297
SHEET    9   B10 MET B 401  SER B 404  1  O  GLY B 402   N  LEU B 366
SHEET   10   B10 PHE B 440  GLN B 441  1  O  GLN B 441   N  TYR B 403
SHEET    1   C10 THR C 154  SER C 160  0
SHEET    2   C10 THR C 163  GLU C 170 -1  O  GLY C 169   N  THR C 154
SHEET    3   C10 LYS C 173  THR C 182 -1  O  LYS C 173   N  GLU C 170
SHEET    4   C10 ALA C 209  PHE C 213 -1  O  ASN C 212   N  THR C 180
SHEET    5   C10 TYR C 191  TYR C 197  1  N  GLY C 196   O  ILE C 211
SHEET    6   C10 ILE C 267  CYS C 277  1  O  ASP C 268   N  TYR C 191
SHEET    7   C10 ILE C 294  GLN C 300  1  O  LEU C 298   N  VAL C 274
SHEET    8   C10 GLY C 365  ASP C 370  1  O  GLY C 365   N  THR C 297
SHEET    9   C10 MET C 401  GLN C 405  1  O  GLY C 402   N  LEU C 366
SHEET   10   C10 PHE C 440  GLN C 441  1  O  GLN C 441   N  TYR C 403
SSBOND   1 CYS A  277    CYS A  412                          1555   1555  2.04
SSBOND   2 CYS A  309    CYS A  384                          1555   1555  2.03
SSBOND   3 CYS B   96    CYS B  131                          1555   1555  2.03
SSBOND   4 CYS B  277    CYS B  412                          1555   1555  2.04
SSBOND   5 CYS B  309    CYS B  384                          1555   1555  2.03
SSBOND   6 CYS C  277    CYS C  412                          1555   1555  2.04
SSBOND   7 CYS C  309    CYS C  384                          1555   1555  2.03
LINK         ND2 ASN C 447                 C1  NAG C 500     1555   1555  1.45
LINK         ND2 ASN B 447                 C1  NAG B 500     1555   1555  1.45
CISPEP   1 ALA A  189    PRO A  190          0         0.32
CISPEP   2 ALA A  362    PRO A  363          0         0.65
CISPEP   3 ALA B  189    PRO B  190          0        -2.17
CISPEP   4 ALA B  362    PRO B  363          0         1.09
CISPEP   5 ALA C  189    PRO C  190          0         0.26
CISPEP   6 ALA C  362    PRO C  363          0         0.08
SITE     1 AC1  7 TYR A 315  GLN A 379  ASN A 447  GLN A 450
SITE     2 AC1  7 TRP A 451  HOH A 751  HOH A 815
SITE     1 AC2  7 TYR B 315  GLN B 379  ASN B 447  GLN B 450
SITE     2 AC2  7 TRP B 451  HOH B 748  HOH B 845
SITE     1 AC3  6 TYR C 315  GLN C 379  ASN C 447  GLN C 450
SITE     2 AC3  6 TRP C 451  HOH C 850
CRYST1   80.260   82.010  185.890  90.00  90.00  90.00 P 21 21 21   12
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.012460  0.000000  0.000000        0.00000
SCALE2      0.000000  0.012194  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005380        0.00000
TER    2693      SER A 460
TER    5424      SER B 460
TER    8069      SER C 460
MASTER      400    0    3   54   30    0    6    6 9221    3   58   87
END