longtext: 3PUI-pdb

content
HEADER    HYDROLASE                               04-DEC-10   3PUI
TITLE     COCAINE ESTERASE WITH MUTATIONS G4C, S10C
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: COCAINE ESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.1.1.-;
COMPND   5 ENGINEERED: YES;
COMPND   6 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: RHODOCOCCUS SP.;
SOURCE   3 ORGANISM_TAXID: 104109;
SOURCE   4 STRAIN: MB1 BRESLER;
SOURCE   5 GENE: COCE;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: BL21 GOLD DE3;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PET-22B (+)
KEYWDS    ALPHA/BETA HYDROLASE, JELLY-ROLL BETA-BARREL, CLEAVAGE OF COCAINE,
KEYWDS   2 HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.R.NANCE,J.J.G.TESMER
REVDAT   1   21-SEP-11 3PUI    0
JRNL        AUTH   D.NARASIMHAN,G.T.COLLINS,M.R.NANCE,J.NICHOLS,E.EDWALD,
JRNL        AUTH 2 J.CHAN,M.C.KO,J.H.WOODS,J.J.TESMER,R.K.SUNAHARA
JRNL        TITL   SUBUNIT STABILIZATION AND PEGYLATION OF COCAINE ESTERASE
JRNL        TITL 2 IMPROVES IN VIVO RESIDENCE TIME.
JRNL        REF    MOL.PHARMACOL.                             2011
JRNL        REFN                   ESSN 1521-0111
JRNL        PMID   21890748
JRNL        DOI    10.1124/MOL.111.074997
REMARK   2
REMARK   2 RESOLUTION.    1.53 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.5.0102
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.53
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 25.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 94.7
REMARK   3   NUMBER OF REFLECTIONS             : 99518
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.156
REMARK   3   R VALUE            (WORKING SET) : 0.155
REMARK   3   FREE R VALUE                     : 0.177
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 5240
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.53
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.57
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 7070
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 92.88
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2560
REMARK   3   BIN FREE R VALUE SET COUNT          : 371
REMARK   3   BIN FREE R VALUE                    : 0.2900
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4386
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 6
REMARK   3   SOLVENT ATOMS            : 691
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 13.28
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.00000
REMARK   3    B22 (A**2) : 0.00000
REMARK   3    B33 (A**2) : -0.01000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.080
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.065
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.040
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 2.467
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.970
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.963
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4868 ; 0.007 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A):  3190 ; 0.001 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  6706 ; 1.158 ; 1.954
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  7798 ; 0.858 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   654 ; 6.201 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   215 ;32.282 ;23.814
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   743 ;12.259 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    35 ;15.290 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   742 ; 0.072 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  5743 ; 0.005 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  1019 ; 0.002 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  3103 ; 0.693 ; 1.500
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  1246 ; 0.177 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  5063 ; 1.185 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1765 ; 1.791 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  1643 ; 2.754 ; 4.500
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  8058 ; 0.696 ; 3.000
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 3PUI COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 07-DEC-10.
REMARK 100 THE RCSB ID CODE IS RCSB062827.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 04-MAR-09
REMARK 200  TEMPERATURE           (KELVIN) : NULL
REMARK 200  PH                             : 7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 21-ID-F
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97872
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : NULL
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL
REMARK 200  DATA SCALING SOFTWARE          : HKL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 110836
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.530
REMARK 200  RESOLUTION RANGE LOW       (A) : 25.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 94.6
REMARK 200  DATA REDUNDANCY                : 3.200
REMARK 200  R MERGE                    (I) : 0.06600
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 17.0000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.53
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.56
REMARK 200  COMPLETENESS FOR SHELL     (%) : 90.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00
REMARK 200  R MERGE FOR SHELL          (I) : 0.59900
REMARK 200  R SYM FOR SHELL            (I) : 0.63400
REMARK 200   FOR SHELL         : 1.400
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: PDB ENTRY 1JU3
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: 1JU3
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 56.23
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.81
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 10MM TRIS PH 7.5, 25 MM NACL, 1.6 M
REMARK 280  AMMONIUM SULFATE, 10MM DTT, VAPOR DIFFUSION, HANGING DROP,
REMARK 280  TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 65 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+2/3
REMARK 290       3555   -X+Y,-X,Z+1/3
REMARK 290       4555   -X,-Y,Z+1/2
REMARK 290       5555   Y,-X+Y,Z+1/6
REMARK 290       6555   X-Y,X,Z+5/6
REMARK 290       7555   Y,X,-Z+2/3
REMARK 290       8555   X-Y,-Y,-Z
REMARK 290       9555   -X,-X+Y,-Z+1/3
REMARK 290      10555   -Y,-X,-Z+1/6
REMARK 290      11555   -X+Y,Y,-Z+1/2
REMARK 290      12555   X,X-Y,-Z+5/6
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      148.27067
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       74.13533
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      111.20300
REMARK 290   SMTRY1   5  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   5 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000       37.06767
REMARK 290   SMTRY1   6  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000      185.33833
REMARK 290   SMTRY1   7 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   7  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000      148.27067
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   9 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   9 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   9  0.000000  0.000000 -1.000000       74.13533
REMARK 290   SMTRY1  10  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  10 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000       37.06767
REMARK 290   SMTRY1  11 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  11  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000      111.20300
REMARK 290   SMTRY1  12  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  12  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000      185.33833
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 4340 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 39140 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -139.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2  0.500000  0.866025  0.000000      -52.83500
REMARK 350   BIOMT2   2  0.866025 -0.500000  0.000000       91.51290
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      -37.06767
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     VAL A     2
REMARK 465     ALA A   577
REMARK 465     ALA A   578
REMARK 465     ALA A   579
REMARK 465     LEU A   580
REMARK 465     GLU A   581
REMARK 465     HIS A   582
REMARK 465     HIS A   583
REMARK 465     HIS A   584
REMARK 465     HIS A   585
REMARK 465     HIS A   586
REMARK 465     HIS A   587
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OG   SER A   159     O    HOH A  1112              2.09
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   OD2  ASP A   198     O    HOH A   941     8675     2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    CYS A   4      -45.84     69.58
REMARK 500    ASP A  45      106.74     78.72
REMARK 500    SER A  56      -83.78   -133.79
REMARK 500    PHE A  78     -120.28     48.15
REMARK 500    HIS A  87      -38.50     72.84
REMARK 500    SER A 117     -117.31     62.09
REMARK 500    ALA A 136      148.74   -170.08
REMARK 500    TYR A 152     -119.56   -119.08
REMARK 500    ASP A 198       71.94   -154.87
REMARK 500    GLN A 224      -60.39    -96.76
REMARK 500    PRO A 284       37.48    -88.67
REMARK 500    THR A 371      163.34     71.34
REMARK 500    ASN A 413       52.98   -146.56
REMARK 500    LEU A 508     -121.22     61.20
REMARK 500    ASN A 528       80.62   -164.71
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    CYS A   4        24.2      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA A 592  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 259   OD1
REMARK 620 2 SER A 265   OG  116.1
REMARK 620 N                    1
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL A 588
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL A 589
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL A 590
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL A 591
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA A 592
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL A 593
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3I2F   RELATED DB: PDB
REMARK 900 RELATED ID: 3I2G   RELATED DB: PDB
REMARK 900 RELATED ID: 3I2H   RELATED DB: PDB
REMARK 900 RELATED ID: 3I2I   RELATED DB: PDB
REMARK 900 RELATED ID: 3I2J   RELATED DB: PDB
REMARK 900 RELATED ID: 1JU3   RELATED DB: PDB
REMARK 900 RELATED ID: 1JU4   RELATED DB: PDB
REMARK 900 RELATED ID: 3IDA   RELATED DB: PDB
REMARK 900 RELATED ID: 3PUH   RELATED DB: PDB
DBREF  3PUI A    1   574  UNP    Q9L9D7   COCE_RHOSM       1    574
SEQADV 3PUI CYS A    4  UNP  Q9L9D7    GLY     4 ENGINEERED MUTATION
SEQADV 3PUI CYS A   10  UNP  Q9L9D7    SER    10 ENGINEERED MUTATION
SEQADV 3PUI LYS A  575  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI LEU A  576  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI ALA A  577  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI ALA A  578  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI ALA A  579  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI LEU A  580  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI GLU A  581  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI HIS A  582  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI HIS A  583  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI HIS A  584  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI HIS A  585  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI HIS A  586  UNP  Q9L9D7              EXPRESSION TAG
SEQADV 3PUI HIS A  587  UNP  Q9L9D7              EXPRESSION TAG
SEQRES   1 A  587  MET VAL ASP CYS ASN TYR SER VAL ALA CYS ASN VAL MET
SEQRES   2 A  587  VAL PRO MET ARG ASP GLY VAL ARG LEU ALA VAL ASP LEU
SEQRES   3 A  587  TYR ARG PRO ASP ALA ASP GLY PRO VAL PRO VAL LEU LEU
SEQRES   4 A  587  VAL ARG ASN PRO TYR ASP LYS PHE ASP VAL PHE ALA TRP
SEQRES   5 A  587  SER THR GLN SER THR ASN TRP LEU GLU PHE VAL ARG ASP
SEQRES   6 A  587  GLY TYR ALA VAL VAL ILE GLN ASP THR ARG GLY LEU PHE
SEQRES   7 A  587  ALA SER GLU GLY GLU PHE VAL PRO HIS VAL ASP ASP GLU
SEQRES   8 A  587  ALA ASP ALA GLU ASP THR LEU SER TRP ILE LEU GLU GLN
SEQRES   9 A  587  ALA TRP CYS ASP GLY ASN VAL GLY MET PHE GLY VAL SER
SEQRES  10 A  587  TYR LEU GLY VAL THR GLN TRP GLN ALA ALA VAL SER GLY
SEQRES  11 A  587  VAL GLY GLY LEU LYS ALA ILE ALA PRO SER MET ALA SER
SEQRES  12 A  587  ALA ASP LEU TYR ARG ALA PRO TRP TYR GLY PRO GLY GLY
SEQRES  13 A  587  ALA LEU SER VAL GLU ALA LEU LEU GLY TRP SER ALA LEU
SEQRES  14 A  587  ILE GLY THR GLY LEU ILE THR SER ARG SER ASP ALA ARG
SEQRES  15 A  587  PRO GLU ASP ALA ALA ASP PHE VAL GLN LEU ALA ALA ILE
SEQRES  16 A  587  LEU ASN ASP VAL ALA GLY ALA ALA SER VAL THR PRO LEU
SEQRES  17 A  587  ALA GLU GLN PRO LEU LEU GLY ARG LEU ILE PRO TRP VAL
SEQRES  18 A  587  ILE ASP GLN VAL VAL ASP HIS PRO ASP ASN ASP GLU SER
SEQRES  19 A  587  TRP GLN SER ILE SER LEU PHE GLU ARG LEU GLY GLY LEU
SEQRES  20 A  587  ALA THR PRO ALA LEU ILE THR ALA GLY TRP TYR ASP GLY
SEQRES  21 A  587  PHE VAL GLY GLU SER LEU ARG THR PHE VAL ALA VAL LYS
SEQRES  22 A  587  ASP ASN ALA ASP ALA ARG LEU VAL VAL GLY PRO TRP SER
SEQRES  23 A  587  HIS SER ASN LEU THR GLY ARG ASN ALA ASP ARG LYS PHE
SEQRES  24 A  587  GLY ILE ALA ALA THR TYR PRO ILE GLN GLU ALA THR THR
SEQRES  25 A  587  MET HIS LYS ALA PHE PHE ASP ARG HIS LEU ARG GLY GLU
SEQRES  26 A  587  THR ASP ALA LEU ALA GLY VAL PRO LYS VAL ARG LEU PHE
SEQRES  27 A  587  VAL MET GLY ILE ASP GLU TRP ARG ASP GLU THR ASP TRP
SEQRES  28 A  587  PRO LEU PRO ASP THR ALA TYR THR PRO PHE TYR LEU GLY
SEQRES  29 A  587  GLY SER GLY ALA ALA ASN THR SER THR GLY GLY GLY THR
SEQRES  30 A  587  LEU SER THR SER ILE SER GLY THR GLU SER ALA ASP THR
SEQRES  31 A  587  TYR LEU TYR ASP PRO ALA ASP PRO VAL PRO SER LEU GLY
SEQRES  32 A  587  GLY THR LEU LEU PHE HIS ASN GLY ASP ASN GLY PRO ALA
SEQRES  33 A  587  ASP GLN ARG PRO ILE HIS ASP ARG ASP ASP VAL LEU CYS
SEQRES  34 A  587  TYR SER THR GLU VAL LEU THR ASP PRO VAL GLU VAL THR
SEQRES  35 A  587  GLY THR VAL SER ALA ARG LEU PHE VAL SER SER SER ALA
SEQRES  36 A  587  VAL ASP THR ASP PHE THR ALA LYS LEU VAL ASP VAL PHE
SEQRES  37 A  587  PRO ASP GLY ARG ALA ILE ALA LEU CYS ASP GLY ILE VAL
SEQRES  38 A  587  ARG MET ARG TYR ARG GLU THR LEU VAL ASN PRO THR LEU
SEQRES  39 A  587  ILE GLU ALA GLY GLU ILE TYR GLU VAL ALA ILE ASP MET
SEQRES  40 A  587  LEU ALA THR SER ASN VAL PHE LEU PRO GLY HIS ARG ILE
SEQRES  41 A  587  MET VAL GLN VAL SER SER SER ASN PHE PRO LYS TYR ASP
SEQRES  42 A  587  ARG ASN SER ASN THR GLY GLY VAL ILE ALA ARG GLU GLN
SEQRES  43 A  587  LEU GLU GLU MET CYS THR ALA VAL ASN ARG ILE HIS ARG
SEQRES  44 A  587  GLY PRO GLU HIS PRO SER HIS ILE VAL LEU PRO ILE ILE
SEQRES  45 A  587  LYS ARG LYS LEU ALA ALA ALA LEU GLU HIS HIS HIS HIS
SEQRES  46 A  587  HIS HIS
HET     CL  A 588       1
HET     CL  A 589       1
HET     CL  A 590       1
HET     CL  A 591       1
HET     NA  A 592       1
HET     CL  A 593       1
HETNAM      CL CHLORIDE ION
HETNAM      NA SODIUM ION
FORMUL   2   CL    5(CL 1-)
FORMUL   6   NA    NA 1+
FORMUL   8  HOH   *691(H2 O)
HELIX    1   1 ASP A   48  THR A   54  1                                   7
HELIX    2   2 TRP A   59  ASP A   65  1                                   7
HELIX    3   3 ASP A   89  GLN A  104  1                                  16
HELIX    4   4 SER A  117  VAL A  128  1                                  12
HELIX    5   5 SER A  159  SER A  177  1                                  19
HELIX    6   6 GLU A  184  ASP A  198  1                                  15
HELIX    7   7 ASP A  198  VAL A  205  1                                   8
HELIX    8   8 GLN A  211  ILE A  218  1                                   8
HELIX    9   9 PRO A  219  GLN A  224  1                                   6
HELIX   10  10 ASP A  232  SER A  237  1                                   6
HELIX   11  11 LEU A  240  GLY A  245  1                                   6
HELIX   12  12 PHE A  261  LYS A  273  1                                  13
HELIX   13  13 GLY A  300  THR A  304  5                                   5
HELIX   14  14 PRO A  306  ARG A  323  1                                  18
HELIX   15  15 GLN A  418  HIS A  422  5                                   5
HELIX   16  16 ARG A  484  ARG A  486  5                                   3
HELIX   17  17 VAL A  541  GLU A  545  5                                   5
HELIX   18  18 GLN A  546  MET A  550  5                                   5
SHEET    1   A 6 TYR A   6  PRO A  15  0
SHEET    2   A 6 ARG A  21  PRO A  29 -1  O  LEU A  22   N  VAL A  14
SHEET    3   A 6 ALA A  68  ASP A  73 -1  O  VAL A  69   N  TYR A  27
SHEET    4   A 6 VAL A  35  ASN A  42  1  N  LEU A  38   O  VAL A  70
SHEET    5   A 6 CYS A 107  MET A 113  1  O  GLY A 112   N  LEU A  39
SHEET    6   A 6 LEU A 134  ALA A 136  1  O  LYS A 135   N  VAL A 111
SHEET    1   B 2 GLY A 115  VAL A 116  0
SHEET    2   B 2 PRO A 139  SER A 140  1  O  SER A 140   N  GLY A 115
SHEET    1   C 4 ALA A 251  TYR A 258  0
SHEET    2   C 4 ALA A 278  SER A 286  1  O  ARG A 279   N  ILE A 253
SHEET    3   C 4 VAL A 335  VAL A 339  1  O  PHE A 338   N  VAL A 282
SHEET    4   C 4 GLU A 344  GLU A 348 -1  O  GLU A 344   N  VAL A 339
SHEET    1   D 2 ARG A 293  ASN A 294  0
SHEET    2   D 2 ARG A 297  LYS A 298 -1  O  ARG A 297   N  ASN A 294
SHEET    1   E 6 THR A 377  SER A 379  0
SHEET    2   E 6 ALA A 357  GLY A 364 -1  N  TYR A 362   O  SER A 379
SHEET    3   E 6 HIS A 566  ILE A 572 -1  O  LEU A 569   N  THR A 359
SHEET    4   E 6 VAL A 439  SER A 453 -1  N  SER A 446   O  VAL A 568
SHEET    5   E 6 ALA A 553  ARG A 559 -1  O  HIS A 558   N  PHE A 450
SHEET    6   E 6 SER A 387  TYR A 393 -1  N  TYR A 393   O  ALA A 553
SHEET    1   F 5 THR A 377  SER A 379  0
SHEET    2   F 5 ALA A 357  GLY A 364 -1  N  TYR A 362   O  SER A 379
SHEET    3   F 5 HIS A 566  ILE A 572 -1  O  LEU A 569   N  THR A 359
SHEET    4   F 5 VAL A 439  SER A 453 -1  N  SER A 446   O  VAL A 568
SHEET    5   F 5 TYR A 501  PHE A 514 -1  O  VAL A 503   N  LEU A 449
SHEET    1   G 4 CYS A 429  SER A 431  0
SHEET    2   G 4 ARG A 519  SER A 525 -1  O  VAL A 522   N  TYR A 430
SHEET    3   G 4 ASP A 459  VAL A 467 -1  N  THR A 461   O  SER A 525
SHEET    4   G 4 ALA A 473  ARG A 482 -1  O  CYS A 477   N  LEU A 464
SSBOND   1 CYS A    4    CYS A   10                          1555  12564  2.02
LINK         OD1 ASP A 259                NA    NA A 592     1555   1555  2.90
LINK         OG  SER A 265                NA    NA A 592     1555   1555  2.98
CISPEP   1 ALA A  149    PRO A  150          0        11.45
CISPEP   2 THR A  206    PRO A  207          0        -2.92
CISPEP   3 TRP A  351    PRO A  352          0        -6.62
CISPEP   4 PHE A  529    PRO A  530          0         9.63
SITE     1 AC1  2 LYS A 298  ARG A 472
SITE     1 AC2  2 GLY A 365  HOH A1268
SITE     1 AC3  6 ASN A 197  GLY A 300  ILE A 301  ALA A 302
SITE     2 AC3  6 HOH A 829  HOH A 909
SITE     1 AC4  4 TRP A 345  HOH A 904  HOH A1013  HOH A1060
SITE     1 AC5  7 MET A 141  ALA A 255  GLY A 256  ASP A 259
SITE     2 AC5  7 PHE A 261  VAL A 262  SER A 265
SITE     1 AC6  3 ASP A 470  ARG A 472  HOH A 697
CRYST1  105.670  105.670  222.406  90.00  90.00 120.00 P 65 2 2     12
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.009463  0.005464  0.000000        0.00000
SCALE2      0.000000  0.010927  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004496        0.00000
TER    4718      LEU A 576
MASTER      415    0    6   18   29    0    8    6 5083    1    3   46
END