longtext: 3QBJ-pdb

content
HEADER    HYDROLASE/HYDROLASE INHIBITOR           13-JAN-11   3QBJ
TITLE     CRYSTAL STRUCTURE OF DIPEPTIDYL PEPTIDASE IV IN COMPLEX WITH INHIBITOR
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: DIPEPTIDYL PEPTIDASE 4;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: ADABP, ADENOSINE DEAMINASE COMPLEXING PROTEIN 2, ADCP-2,
COMPND   5 DIPEPTIDYL PEPTIDASE IV, DPP IV, T-CELL ACTIVATION ANTIGEN CD26,
COMPND   6 TP103, DIPEPTIDYL PEPTIDASE IV SOLUBLE FORM;
COMPND   7 EC: 3.4.14.5;
COMPND   8 ENGINEERED: YES;
COMPND   9 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: DPP4, ADCP2, CD26;
SOURCE   6 EXPRESSION_SYSTEM: SPODOPTERA FRUGIPERDA;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 7108
KEYWDS    DIABETES MELLITUS, DIPEPTIDYL-PEPTIDASE IV INHIBITOR, HYPOGLYCEMIC
KEYWDS   2 AGENT, PYRROLIDINES, SPRAGUE-DAWLEY, STRUCTURE-ACTIVITY
KEYWDS   3 RELATIONSHIP, HYDROLASE, GLYCOSYLATION, HYDROLASE-HYDROLASE
KEYWDS   4 INHIBITOR COMPLEX
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.P.LIU
REVDAT   1   25-JAN-12 3QBJ    0
JRNL        AUTH   S.P.LIU
JRNL        TITL   CRYSTAL STRUCTURE OF DIPEPTIDYL PEPTIDASE IV IN COMPLEX WITH
JRNL        TITL 2 INHIBITOR
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.21 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : BUSTER 2.9.6
REMARK   3   AUTHORS     : BRICOGNE,BLANC,BRANDL,FLENSBURG,KELLER,
REMARK   3               : PACIOREK,ROVERSI,SHARFF,SMART,VONRHEIN,
REMARK   3               : WOMACK,MATTHEWS,TEN EYCK,TRONRUD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.21
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 18.11
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 80.3
REMARK   3   NUMBER OF REFLECTIONS             : 78758
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD           : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION   : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET)  : 0.203
REMARK   3   R VALUE            (WORKING SET)  : 0.202
REMARK   3   FREE R VALUE                      : 0.235
REMARK   3   FREE R VALUE TEST SET SIZE   (%)  : 4.970
REMARK   3   FREE R VALUE TEST SET COUNT       : 3918
REMARK   3   ESTIMATED ERROR OF FREE R VALUE   : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED               : 20
REMARK   3   BIN RESOLUTION RANGE HIGH   (ANGSTROMS) : 2.21
REMARK   3   BIN RESOLUTION RANGE LOW    (ANGSTROMS) : 2.27
REMARK   3   BIN COMPLETENESS (WORKING+TEST)     (%) : 80.26
REMARK   3   REFLECTIONS IN BIN (WORKING + TEST SET) : 3919
REMARK   3   BIN R VALUE        (WORKING + TEST SET) : 0.2440
REMARK   3   REFLECTIONS IN BIN        (WORKING SET) : 3724
REMARK   3   BIN R VALUE               (WORKING SET) : 0.2432
REMARK   3   BIN FREE R VALUE                        : 0.2588
REMARK   3   BIN FREE R VALUE TEST SET SIZE      (%) : 4.98
REMARK   3   BIN FREE R VALUE TEST SET COUNT         : 195
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE     : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 11914
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 198
REMARK   3   SOLVENT ATOMS            : 71
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 42.75
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 72.23
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 18.50950
REMARK   3    B22 (A**2) : -35.32870
REMARK   3    B33 (A**2) : 16.81920
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT                    (A) : 0.46
REMARK   3   DPI (BLOW EQ-10) BASED ON R VALUE        (A) : NULL
REMARK   3   DPI (BLOW EQ-9) BASED ON FREE R VALUE    (A) : NULL
REMARK   3   DPI (CRUICKSHANK) BASED ON R VALUE       (A) : NULL
REMARK   3   DPI (CRUICKSHANK) BASED ON FREE R VALUE  (A) : NULL
REMARK   3
REMARK   3   REFERENCES: BLOW, D. (2002) ACTA CRYST D58, 792-797
REMARK   3               CRUICKSHANK, D.W.J. (1999) ACTA CRYST D55, 583-601
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.896
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.880
REMARK   3
REMARK   3   NUMBER OF GEOMETRIC FUNCTION TERMS DEFINED : 15
REMARK   3   TERM                          COUNT    WEIGHT   FUNCTION.
REMARK   3    BOND LENGTHS              : 12483  ; 2.000  ; HARMONIC
REMARK   3    BOND ANGLES               : 17004  ; 2.000  ; HARMONIC
REMARK   3    TORSION ANGLES            : 4116   ; 2.000  ; SINUSOIDAL
REMARK   3    TRIGONAL CARBON PLANES    : 321    ; 2.000  ; HARMONIC
REMARK   3    GENERAL PLANES            : 1816   ; 5.000  ; HARMONIC
REMARK   3    ISOTROPIC THERMAL FACTORS : 12483  ; 20.000 ; HARMONIC
REMARK   3    BAD NON-BONDED CONTACTS   : NULL   ; NULL   ; NULL
REMARK   3    IMPROPER TORSIONS         : NULL   ; NULL   ; NULL
REMARK   3    PSEUDOROTATION ANGLES     : NULL   ; NULL   ; NULL
REMARK   3    CHIRAL IMPROPER TORSION   : 1632   ; 5.000  ; NULL
REMARK   3    SUM OF OCCUPANCIES        : NULL   ; NULL   ; NULL
REMARK   3    UTILITY DISTANCES         : NULL   ; NULL   ; NULL
REMARK   3    UTILITY ANGLES            : NULL   ; NULL   ; NULL
REMARK   3    UTILITY TORSION           : NULL   ; NULL   ; NULL
REMARK   3    IDEAL-DIST CONTACT TERM   : 13264  ; 4.000  ; NULL
REMARK   3
REMARK   3   RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3    BOND LENGTHS                       (A) : 0.010
REMARK   3    BOND ANGLES                  (DEGREES) : 1.11
REMARK   3    PEPTIDE OMEGA TORSION ANGLES (DEGREES) : 2.99
REMARK   3    OTHER TORSION ANGLES         (DEGREES) : 19.05
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION:   A   39    A  796
REMARK   3    ORIGIN FOR THE GROUP (A):   16.7180   16.3799   27.6709
REMARK   3    T TENSOR
REMARK   3     T11:   -0.2752 T22:    0.3103
REMARK   3     T33:   -0.1729 T12:    0.0095
REMARK   3     T13:    0.0039 T23:    0.1827
REMARK   3    L TENSOR
REMARK   3     L11:    0.6264 L22:    0.4177
REMARK   3     L33:    1.8418 L12:   -0.0404
REMARK   3     L13:   -0.1631 L23:   -0.4122
REMARK   3    S TENSOR
REMARK   3     S11:    0.0411 S12:    0.2268 S13:    0.1080
REMARK   3     S21:   -0.0336 S22:    0.0268 S23:    0.0086
REMARK   3     S31:    0.0228 S32:   -0.0822 S33:   -0.0679
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION:   B   39    B  796
REMARK   3    ORIGIN FOR THE GROUP (A):    0.0608   -2.5173   78.4587
REMARK   3    T TENSOR
REMARK   3     T11:   -0.2700 T22:   -0.1414
REMARK   3     T33:   -0.2778 T12:    0.0125
REMARK   3     T13:    0.0217 T23:    0.0448
REMARK   3    L TENSOR
REMARK   3     L11:    0.9531 L22:    0.5814
REMARK   3     L33:    2.0308 L12:    0.1197
REMARK   3     L13:   -0.4564 L23:   -0.4377
REMARK   3    S TENSOR
REMARK   3     S11:    0.0020 S12:    0.0852 S13:    0.0250
REMARK   3     S21:    0.0850 S22:    0.1245 S23:    0.0564
REMARK   3     S31:    0.0884 S32:   -0.2422 S33:   -0.1265
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3QBJ COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 18-JAN-11.
REMARK 100 THE RCSB ID CODE IS RCSB063436.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 29-JUN-05
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : NULL
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : APS
REMARK 200  BEAMLINE                       : 17-ID
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : SI 111 CHANNEL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XSCALE
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 79656
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.200
REMARK 200  RESOLUTION RANGE LOW       (A) : 42.409
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 80.3
REMARK 200  DATA REDUNDANCY                : 5.600
REMARK 200  R MERGE                    (I) : 0.10100
REMARK 200  R SYM                      (I) : 0.10100
REMARK 200   FOR THE DATA SET  : 14.7000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.32
REMARK 200  COMPLETENESS FOR SHELL     (%) : 55.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 4.50
REMARK 200  R MERGE FOR SHELL          (I) : 0.39200
REMARK 200  R SYM FOR SHELL            (I) : 0.39200
REMARK 200   FOR SHELL         : 1.800
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 55.79
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.78
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       32.87550
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      212.08000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       34.54100
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      212.08000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       32.87550
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       34.54100
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 6970 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 60700 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: 20.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A    31
REMARK 465     THR A    32
REMARK 465     ASP A    33
REMARK 465     ASP A    34
REMARK 465     ALA A    35
REMARK 465     THR A    36
REMARK 465     ALA A    37
REMARK 465     ASP A    38
REMARK 465     LEU A   767
REMARK 465     VAL A   768
REMARK 465     PRO A   769
REMARK 465     ARG A   770
REMARK 465     GLY A   771
REMARK 465     SER A   772
REMARK 465     HIS A   773
REMARK 465     HIS A   774
REMARK 465     HIS A   775
REMARK 465     HIS A   776
REMARK 465     HIS A   777
REMARK 465     HIS A   778
REMARK 465     GLY B    31
REMARK 465     THR B    32
REMARK 465     ASP B    33
REMARK 465     ASP B    34
REMARK 465     ALA B    35
REMARK 465     THR B    36
REMARK 465     ALA B    37
REMARK 465     ASP B    38
REMARK 465     LEU B   767
REMARK 465     VAL B   768
REMARK 465     PRO B   769
REMARK 465     ARG B   770
REMARK 465     GLY B   771
REMARK 465     SER B   772
REMARK 465     HIS B   773
REMARK 465     HIS B   774
REMARK 465     HIS B   775
REMARK 465     HIS B   776
REMARK 465     HIS B   777
REMARK 465     HIS B   778
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O4   NAG A   796     O5   NAG A   798              2.17
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A  40     -145.02   -115.40
REMARK 500    SER A  64     -168.04   -165.25
REMARK 500    HIS A  66       -2.41   -145.08
REMARK 500    GLN A  72       68.08   -100.68
REMARK 500    GLU A  73      107.91     48.91
REMARK 500    GLN A 123     -100.58   -113.16
REMARK 500    TRP A 124     -151.78    -95.39
REMARK 500    ASN A 151       36.69     70.53
REMARK 500    HIS A 162       30.42   -144.13
REMARK 500    ILE A 193      -62.77   -121.84
REMARK 500    VAL A 207      -60.82   -103.20
REMARK 500    SER A 242     -154.83     61.83
REMARK 500    GLN A 320       28.85    -70.06
REMARK 500    SER A 333       30.40    -95.82
REMARK 500    SER A 334      -29.89   -160.34
REMARK 500    LYS A 392      -73.60    -46.05
REMARK 500    ASP A 393     -143.71   -115.75
REMARK 500    ASN A 450       82.50   -161.33
REMARK 500    TYR A 547      -72.47   -131.77
REMARK 500    ARG A 596       14.00     59.63
REMARK 500    ARG A 597       44.86   -141.38
REMARK 500    THR A 600      -89.55   -118.14
REMARK 500    SER A 630     -113.58     61.96
REMARK 500    ALA A 654       54.05     39.92
REMARK 500    ASP A 678      -95.52    -91.61
REMARK 500    ASN A 710      -68.97    -98.54
REMARK 500    GLU B  73      102.56     41.84
REMARK 500    GLN B 123     -100.18   -113.27
REMARK 500    TRP B 124     -151.19    -95.47
REMARK 500    ASN B 151       36.68     70.12
REMARK 500    HIS B 162       30.08   -144.77
REMARK 500    ILE B 193      -62.64   -121.69
REMARK 500    VAL B 207      -60.76   -103.19
REMARK 500    SER B 242     -156.53     61.23
REMARK 500    GLN B 320       34.83    -69.39
REMARK 500    SER B 334      -31.56   -157.76
REMARK 500    LYS B 392      -71.92    -47.22
REMARK 500    ASP B 393     -143.50   -115.20
REMARK 500    ASN B 450       82.78   -160.09
REMARK 500    TYR B 547      -71.38   -133.10
REMARK 500    ARG B 597       54.68   -142.56
REMARK 500    THR B 600      -89.53   -118.25
REMARK 500    SER B 630     -114.83     65.04
REMARK 500    ASP B 678      -97.28   -115.67
REMARK 500    ASN B 710      -68.07    -97.38
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    ARG A  40        23.1      L          L   OUTSIDE RANGE
REMARK 500    LYS A  41        23.6      L          L   OUTSIDE RANGE
REMARK 500    ILE A 389        22.0      L          L   OUTSIDE RANGE
REMARK 500    LYS A 392        22.1      L          L   OUTSIDE RANGE
REMARK 500    HIS A 682        24.9      L          L   OUTSIDE RANGE
REMARK 500    LYS B  41        23.7      L          L   OUTSIDE RANGE
REMARK 500    ASN B  92        23.7      L          L   OUTSIDE RANGE
REMARK 500    VAL B 202        25.0      L          L   OUTSIDE RANGE
REMARK 500    GLN B 320        24.8      L          L   OUTSIDE RANGE
REMARK 500    ILE B 389        21.5      L          L   OUTSIDE RANGE
REMARK 500    LYS B 392        23.6      L          L   OUTSIDE RANGE
REMARK 500    VAL B 486        24.0      L          L   OUTSIDE RANGE
REMARK 500    VAL B 558        22.3      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 794
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 797
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 796
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 798
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE BMA A 802
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MAN A 803
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 799
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 800
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NXZ A 900
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 794
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 797
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 796
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NXZ B 900
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3F8S   RELATED DB: PDB
REMARK 900 CRYSTAL STRUCTURE OF DIPEPTIDYL PEPTIDASE IV IN COMPLEX
REMARK 900 WITH INHIBITOR
DBREF  3QBJ A   31   766  UNP    P27487   DPP4_HUMAN      31    766
DBREF  3QBJ B   31   766  UNP    P27487   DPP4_HUMAN      31    766
SEQADV 3QBJ ALA A  150  UNP  P27487    ASN   150 ENGINEERED MUTATION
SEQADV 3QBJ ALA A  520  UNP  P27487    ASN   520 ENGINEERED MUTATION
SEQADV 3QBJ LEU A  767  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ VAL A  768  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ PRO A  769  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ ARG A  770  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ GLY A  771  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ SER A  772  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS A  773  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS A  774  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS A  775  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS A  776  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS A  777  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS A  778  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ ALA B  150  UNP  P27487    ASN   150 ENGINEERED MUTATION
SEQADV 3QBJ ALA B  520  UNP  P27487    ASN   520 ENGINEERED MUTATION
SEQADV 3QBJ LEU B  767  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ VAL B  768  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ PRO B  769  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ ARG B  770  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ GLY B  771  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ SER B  772  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS B  773  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS B  774  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS B  775  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS B  776  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS B  777  UNP  P27487              EXPRESSION TAG
SEQADV 3QBJ HIS B  778  UNP  P27487              EXPRESSION TAG
SEQRES   1 A  748  GLY THR ASP ASP ALA THR ALA ASP SER ARG LYS THR TYR
SEQRES   2 A  748  THR LEU THR ASP TYR LEU LYS ASN THR TYR ARG LEU LYS
SEQRES   3 A  748  LEU TYR SER LEU ARG TRP ILE SER ASP HIS GLU TYR LEU
SEQRES   4 A  748  TYR LYS GLN GLU ASN ASN ILE LEU VAL PHE ASN ALA GLU
SEQRES   5 A  748  TYR GLY ASN SER SER VAL PHE LEU GLU ASN SER THR PHE
SEQRES   6 A  748  ASP GLU PHE GLY HIS SER ILE ASN ASP TYR SER ILE SER
SEQRES   7 A  748  PRO ASP GLY GLN PHE ILE LEU LEU GLU TYR ASN TYR VAL
SEQRES   8 A  748  LYS GLN TRP ARG HIS SER TYR THR ALA SER TYR ASP ILE
SEQRES   9 A  748  TYR ASP LEU ASN LYS ARG GLN LEU ILE THR GLU GLU ARG
SEQRES  10 A  748  ILE PRO ALA ASN THR GLN TRP VAL THR TRP SER PRO VAL
SEQRES  11 A  748  GLY HIS LYS LEU ALA TYR VAL TRP ASN ASN ASP ILE TYR
SEQRES  12 A  748  VAL LYS ILE GLU PRO ASN LEU PRO SER TYR ARG ILE THR
SEQRES  13 A  748  TRP THR GLY LYS GLU ASP ILE ILE TYR ASN GLY ILE THR
SEQRES  14 A  748  ASP TRP VAL TYR GLU GLU GLU VAL PHE SER ALA TYR SER
SEQRES  15 A  748  ALA LEU TRP TRP SER PRO ASN GLY THR PHE LEU ALA TYR
SEQRES  16 A  748  ALA GLN PHE ASN ASP THR GLU VAL PRO LEU ILE GLU TYR
SEQRES  17 A  748  SER PHE TYR SER ASP GLU SER LEU GLN TYR PRO LYS THR
SEQRES  18 A  748  VAL ARG VAL PRO TYR PRO LYS ALA GLY ALA VAL ASN PRO
SEQRES  19 A  748  THR VAL LYS PHE PHE VAL VAL ASN THR ASP SER LEU SER
SEQRES  20 A  748  SER VAL THR ASN ALA THR SER ILE GLN ILE THR ALA PRO
SEQRES  21 A  748  ALA SER MET LEU ILE GLY ASP HIS TYR LEU CYS ASP VAL
SEQRES  22 A  748  THR TRP ALA THR GLN GLU ARG ILE SER LEU GLN TRP LEU
SEQRES  23 A  748  ARG ARG ILE GLN ASN TYR SER VAL MET ASP ILE CYS ASP
SEQRES  24 A  748  TYR ASP GLU SER SER GLY ARG TRP ASN CYS LEU VAL ALA
SEQRES  25 A  748  ARG GLN HIS ILE GLU MET SER THR THR GLY TRP VAL GLY
SEQRES  26 A  748  ARG PHE ARG PRO SER GLU PRO HIS PHE THR LEU ASP GLY
SEQRES  27 A  748  ASN SER PHE TYR LYS ILE ILE SER ASN GLU GLU GLY TYR
SEQRES  28 A  748  ARG HIS ILE CYS TYR PHE GLN ILE ASP LYS LYS ASP CYS
SEQRES  29 A  748  THR PHE ILE THR LYS GLY THR TRP GLU VAL ILE GLY ILE
SEQRES  30 A  748  GLU ALA LEU THR SER ASP TYR LEU TYR TYR ILE SER ASN
SEQRES  31 A  748  GLU TYR LYS GLY MET PRO GLY GLY ARG ASN LEU TYR LYS
SEQRES  32 A  748  ILE GLN LEU SER ASP TYR THR LYS VAL THR CYS LEU SER
SEQRES  33 A  748  CYS GLU LEU ASN PRO GLU ARG CYS GLN TYR TYR SER VAL
SEQRES  34 A  748  SER PHE SER LYS GLU ALA LYS TYR TYR GLN LEU ARG CYS
SEQRES  35 A  748  SER GLY PRO GLY LEU PRO LEU TYR THR LEU HIS SER SER
SEQRES  36 A  748  VAL ASN ASP LYS GLY LEU ARG VAL LEU GLU ASP ASN SER
SEQRES  37 A  748  ALA LEU ASP LYS MET LEU GLN ASN VAL GLN MET PRO SER
SEQRES  38 A  748  LYS LYS LEU ASP PHE ILE ILE LEU ALA GLU THR LYS PHE
SEQRES  39 A  748  TRP TYR GLN MET ILE LEU PRO PRO HIS PHE ASP LYS SER
SEQRES  40 A  748  LYS LYS TYR PRO LEU LEU LEU ASP VAL TYR ALA GLY PRO
SEQRES  41 A  748  CYS SER GLN LYS ALA ASP THR VAL PHE ARG LEU ASN TRP
SEQRES  42 A  748  ALA THR TYR LEU ALA SER THR GLU ASN ILE ILE VAL ALA
SEQRES  43 A  748  SER PHE ASP GLY ARG GLY SER GLY TYR GLN GLY ASP LYS
SEQRES  44 A  748  ILE MET HIS ALA ILE ASN ARG ARG LEU GLY THR PHE GLU
SEQRES  45 A  748  VAL GLU ASP GLN ILE GLU ALA ALA ARG GLN PHE SER LYS
SEQRES  46 A  748  MET GLY PHE VAL ASP ASN LYS ARG ILE ALA ILE TRP GLY
SEQRES  47 A  748  TRP SER TYR GLY GLY TYR VAL THR SER MET VAL LEU GLY
SEQRES  48 A  748  SER GLY SER GLY VAL PHE LYS CYS GLY ILE ALA VAL ALA
SEQRES  49 A  748  PRO VAL SER ARG TRP GLU TYR TYR ASP SER VAL TYR THR
SEQRES  50 A  748  GLU ARG TYR MET GLY LEU PRO THR PRO GLU ASP ASN LEU
SEQRES  51 A  748  ASP HIS TYR ARG ASN SER THR VAL MET SER ARG ALA GLU
SEQRES  52 A  748  ASN PHE LYS GLN VAL GLU TYR LEU LEU ILE HIS GLY THR
SEQRES  53 A  748  ALA ASP ASP ASN VAL HIS PHE GLN GLN SER ALA GLN ILE
SEQRES  54 A  748  SER LYS ALA LEU VAL ASP VAL GLY VAL ASP PHE GLN ALA
SEQRES  55 A  748  MET TRP TYR THR ASP GLU ASP HIS GLY ILE ALA SER SER
SEQRES  56 A  748  THR ALA HIS GLN HIS ILE TYR THR HIS MET SER HIS PHE
SEQRES  57 A  748  ILE LYS GLN CYS PHE SER LEU PRO LEU VAL PRO ARG GLY
SEQRES  58 A  748  SER HIS HIS HIS HIS HIS HIS
SEQRES   1 B  748  GLY THR ASP ASP ALA THR ALA ASP SER ARG LYS THR TYR
SEQRES   2 B  748  THR LEU THR ASP TYR LEU LYS ASN THR TYR ARG LEU LYS
SEQRES   3 B  748  LEU TYR SER LEU ARG TRP ILE SER ASP HIS GLU TYR LEU
SEQRES   4 B  748  TYR LYS GLN GLU ASN ASN ILE LEU VAL PHE ASN ALA GLU
SEQRES   5 B  748  TYR GLY ASN SER SER VAL PHE LEU GLU ASN SER THR PHE
SEQRES   6 B  748  ASP GLU PHE GLY HIS SER ILE ASN ASP TYR SER ILE SER
SEQRES   7 B  748  PRO ASP GLY GLN PHE ILE LEU LEU GLU TYR ASN TYR VAL
SEQRES   8 B  748  LYS GLN TRP ARG HIS SER TYR THR ALA SER TYR ASP ILE
SEQRES   9 B  748  TYR ASP LEU ASN LYS ARG GLN LEU ILE THR GLU GLU ARG
SEQRES  10 B  748  ILE PRO ALA ASN THR GLN TRP VAL THR TRP SER PRO VAL
SEQRES  11 B  748  GLY HIS LYS LEU ALA TYR VAL TRP ASN ASN ASP ILE TYR
SEQRES  12 B  748  VAL LYS ILE GLU PRO ASN LEU PRO SER TYR ARG ILE THR
SEQRES  13 B  748  TRP THR GLY LYS GLU ASP ILE ILE TYR ASN GLY ILE THR
SEQRES  14 B  748  ASP TRP VAL TYR GLU GLU GLU VAL PHE SER ALA TYR SER
SEQRES  15 B  748  ALA LEU TRP TRP SER PRO ASN GLY THR PHE LEU ALA TYR
SEQRES  16 B  748  ALA GLN PHE ASN ASP THR GLU VAL PRO LEU ILE GLU TYR
SEQRES  17 B  748  SER PHE TYR SER ASP GLU SER LEU GLN TYR PRO LYS THR
SEQRES  18 B  748  VAL ARG VAL PRO TYR PRO LYS ALA GLY ALA VAL ASN PRO
SEQRES  19 B  748  THR VAL LYS PHE PHE VAL VAL ASN THR ASP SER LEU SER
SEQRES  20 B  748  SER VAL THR ASN ALA THR SER ILE GLN ILE THR ALA PRO
SEQRES  21 B  748  ALA SER MET LEU ILE GLY ASP HIS TYR LEU CYS ASP VAL
SEQRES  22 B  748  THR TRP ALA THR GLN GLU ARG ILE SER LEU GLN TRP LEU
SEQRES  23 B  748  ARG ARG ILE GLN ASN TYR SER VAL MET ASP ILE CYS ASP
SEQRES  24 B  748  TYR ASP GLU SER SER GLY ARG TRP ASN CYS LEU VAL ALA
SEQRES  25 B  748  ARG GLN HIS ILE GLU MET SER THR THR GLY TRP VAL GLY
SEQRES  26 B  748  ARG PHE ARG PRO SER GLU PRO HIS PHE THR LEU ASP GLY
SEQRES  27 B  748  ASN SER PHE TYR LYS ILE ILE SER ASN GLU GLU GLY TYR
SEQRES  28 B  748  ARG HIS ILE CYS TYR PHE GLN ILE ASP LYS LYS ASP CYS
SEQRES  29 B  748  THR PHE ILE THR LYS GLY THR TRP GLU VAL ILE GLY ILE
SEQRES  30 B  748  GLU ALA LEU THR SER ASP TYR LEU TYR TYR ILE SER ASN
SEQRES  31 B  748  GLU TYR LYS GLY MET PRO GLY GLY ARG ASN LEU TYR LYS
SEQRES  32 B  748  ILE GLN LEU SER ASP TYR THR LYS VAL THR CYS LEU SER
SEQRES  33 B  748  CYS GLU LEU ASN PRO GLU ARG CYS GLN TYR TYR SER VAL
SEQRES  34 B  748  SER PHE SER LYS GLU ALA LYS TYR TYR GLN LEU ARG CYS
SEQRES  35 B  748  SER GLY PRO GLY LEU PRO LEU TYR THR LEU HIS SER SER
SEQRES  36 B  748  VAL ASN ASP LYS GLY LEU ARG VAL LEU GLU ASP ASN SER
SEQRES  37 B  748  ALA LEU ASP LYS MET LEU GLN ASN VAL GLN MET PRO SER
SEQRES  38 B  748  LYS LYS LEU ASP PHE ILE ILE LEU ALA GLU THR LYS PHE
SEQRES  39 B  748  TRP TYR GLN MET ILE LEU PRO PRO HIS PHE ASP LYS SER
SEQRES  40 B  748  LYS LYS TYR PRO LEU LEU LEU ASP VAL TYR ALA GLY PRO
SEQRES  41 B  748  CYS SER GLN LYS ALA ASP THR VAL PHE ARG LEU ASN TRP
SEQRES  42 B  748  ALA THR TYR LEU ALA SER THR GLU ASN ILE ILE VAL ALA
SEQRES  43 B  748  SER PHE ASP GLY ARG GLY SER GLY TYR GLN GLY ASP LYS
SEQRES  44 B  748  ILE MET HIS ALA ILE ASN ARG ARG LEU GLY THR PHE GLU
SEQRES  45 B  748  VAL GLU ASP GLN ILE GLU ALA ALA ARG GLN PHE SER LYS
SEQRES  46 B  748  MET GLY PHE VAL ASP ASN LYS ARG ILE ALA ILE TRP GLY
SEQRES  47 B  748  TRP SER TYR GLY GLY TYR VAL THR SER MET VAL LEU GLY
SEQRES  48 B  748  SER GLY SER GLY VAL PHE LYS CYS GLY ILE ALA VAL ALA
SEQRES  49 B  748  PRO VAL SER ARG TRP GLU TYR TYR ASP SER VAL TYR THR
SEQRES  50 B  748  GLU ARG TYR MET GLY LEU PRO THR PRO GLU ASP ASN LEU
SEQRES  51 B  748  ASP HIS TYR ARG ASN SER THR VAL MET SER ARG ALA GLU
SEQRES  52 B  748  ASN PHE LYS GLN VAL GLU TYR LEU LEU ILE HIS GLY THR
SEQRES  53 B  748  ALA ASP ASP ASN VAL HIS PHE GLN GLN SER ALA GLN ILE
SEQRES  54 B  748  SER LYS ALA LEU VAL ASP VAL GLY VAL ASP PHE GLN ALA
SEQRES  55 B  748  MET TRP TYR THR ASP GLU ASP HIS GLY ILE ALA SER SER
SEQRES  56 B  748  THR ALA HIS GLN HIS ILE TYR THR HIS MET SER HIS PHE
SEQRES  57 B  748  ILE LYS GLN CYS PHE SER LEU PRO LEU VAL PRO ARG GLY
SEQRES  58 B  748  SER HIS HIS HIS HIS HIS HIS
MODRES 3QBJ ASN A  219  ASN  GLYCOSYLATION SITE
MODRES 3QBJ ASN A   85  ASN  GLYCOSYLATION SITE
MODRES 3QBJ ASN A  229  ASN  GLYCOSYLATION SITE
MODRES 3QBJ ASN B  229  ASN  GLYCOSYLATION SITE
MODRES 3QBJ ASN B   85  ASN  GLYCOSYLATION SITE
MODRES 3QBJ ASN A  321  ASN  GLYCOSYLATION SITE
HET    NAG  A 794      14
HET    NAG  A 797      14
HET    NAG  A 796      14
HET    NAG  A 798      14
HET    BMA  A 802      11
HET    MAN  A 803      11
HET    NAG  A 799      14
HET    NAG  A 800      14
HET    NXZ  A 900      25
HET    NAG  B 794      14
HET    NAG  B 797      14
HET    NAG  B 796      14
HET    NXZ  B 900      25
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     BMA BETA-D-MANNOSE
HETNAM     MAN ALPHA-D-MANNOSE
HETNAM     NXZ 1-[(3S,4S)-4-AMINO-1-(6-PHENYLPYRIMIDIN-4-YL)
HETNAM   2 NXZ  PYRROLIDIN-3-YL]PIPERIDIN-2-ONE
FORMUL   3  NAG    9(C8 H15 N O6)
FORMUL   4  BMA    C6 H12 O6
FORMUL   4  MAN    C6 H12 O6
FORMUL   7  NXZ    2(C19 H23 N5 O)
FORMUL  11  HOH   *71(H2 O)
HELIX    1   1 THR A   44  LYS A   50  1                                   7
HELIX    2   2 SER A   93  ASP A   96  5                                   4
HELIX    3   3 ASP A  200  VAL A  207  1                                   8
HELIX    4   4 PRO A  290  ILE A  295  1                                   6
HELIX    5   5 LEU A  340  GLN A  344  5                                   5
HELIX    6   6 GLU A  421  MET A  425  5                                   5
HELIX    7   7 LYS A  463  ALA A  465  5                                   3
HELIX    8   8 ASN A  497  ASN A  506  1                                  10
HELIX    9   9 ASN A  562  THR A  570  1                                   9
HELIX   10  10 GLY A  587  HIS A  592  1                                   6
HELIX   11  11 ALA A  593  ASN A  595  5                                   3
HELIX   12  12 THR A  600  LYS A  615  1                                  16
HELIX   13  13 SER A  630  GLY A  641  1                                  12
HELIX   14  14 ARG A  658  TYR A  662  5                                   5
HELIX   15  15 ASP A  663  GLY A  672  1                                  10
HELIX   16  16 ASN A  679  SER A  686  1                                   8
HELIX   17  17 VAL A  688  VAL A  698  5                                  11
HELIX   18  18 PHE A  713  ASP A  725  1                                  13
HELIX   19  19 SER A  744  PHE A  763  1                                  20
HELIX   20  20 THR B   44  LYS B   50  1                                   7
HELIX   21  21 SER B   93  ASP B   96  5                                   4
HELIX   22  22 ASP B  200  VAL B  207  1                                   8
HELIX   23  23 PRO B  290  ILE B  295  1                                   6
HELIX   24  24 LEU B  340  GLN B  344  5                                   5
HELIX   25  25 GLU B  421  MET B  425  5                                   5
HELIX   26  26 LYS B  463  ALA B  465  5                                   3
HELIX   27  27 ASN B  497  ASN B  506  1                                  10
HELIX   28  28 ASN B  562  THR B  570  1                                   9
HELIX   29  29 GLY B  587  HIS B  592  1                                   6
HELIX   30  30 ALA B  593  ASN B  595  5                                   3
HELIX   31  31 THR B  600  MET B  616  1                                  17
HELIX   32  32 SER B  630  GLY B  641  1                                  12
HELIX   33  33 ARG B  658  TYR B  662  5                                   5
HELIX   34  34 ASP B  663  GLY B  672  1                                  10
HELIX   35  35 ASN B  679  SER B  686  1                                   8
HELIX   36  36 VAL B  688  VAL B  698  5                                  11
HELIX   37  37 PHE B  713  ASP B  725  1                                  13
HELIX   38  38 SER B  744  PHE B  763  1                                  20
SHEET    1   A 4 ARG A  61  TRP A  62  0
SHEET    2   A 4 GLU A  67  LYS A  71 -1  O  LEU A  69   N  ARG A  61
SHEET    3   A 4 ASN A  75  ASN A  80 -1  O  PHE A  79   N  TYR A  68
SHEET    4   A 4 SER A  86  GLU A  91 -1  O  LEU A  90   N  ILE A  76
SHEET    1   B 4 ASP A 104  ILE A 107  0
SHEET    2   B 4 PHE A 113  LYS A 122 -1  O  GLU A 117   N  ASP A 104
SHEET    3   B 4 TYR A 128  ASP A 136 -1  O  ASP A 133   N  LEU A 116
SHEET    4   B 4 GLN A 141  LEU A 142 -1  O  GLN A 141   N  ASP A 136
SHEET    1   C 4 TRP A 154  TRP A 157  0
SHEET    2   C 4 LEU A 164  TRP A 168 -1  O  ALA A 165   N  THR A 156
SHEET    3   C 4 ASP A 171  LYS A 175 -1  O  LYS A 175   N  LEU A 164
SHEET    4   C 4 TYR A 183  ARG A 184 -1  O  TYR A 183   N  VAL A 174
SHEET    1   D 3 ILE A 194  ASN A 196  0
SHEET    2   D 3 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3   D 3 LEU A 214  TRP A 216 -1  N  TRP A 215   O  ALA A 224
SHEET    1   E 4 ILE A 194  ASN A 196  0
SHEET    2   E 4 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3   E 4 THR A 265  ASN A 272 -1  O  PHE A 269   N  TYR A 225
SHEET    4   E 4 SER A 284  ILE A 287 -1  O  ILE A 285   N  VAL A 270
SHEET    1   F 2 LEU A 235  PHE A 240  0
SHEET    2   F 2 LYS A 250  PRO A 255 -1  O  VAL A 252   N  TYR A 238
SHEET    1   G 4 HIS A 298  TRP A 305  0
SHEET    2   G 4 ARG A 310  ARG A 317 -1  O  SER A 312   N  THR A 304
SHEET    3   G 4 TYR A 322  TYR A 330 -1  O  ASP A 326   N  LEU A 313
SHEET    4   G 4 TRP A 337  ASN A 338 -1  O  ASN A 338   N  ASP A 329
SHEET    1   H 4 HIS A 298  TRP A 305  0
SHEET    2   H 4 ARG A 310  ARG A 317 -1  O  SER A 312   N  THR A 304
SHEET    3   H 4 TYR A 322  TYR A 330 -1  O  ASP A 326   N  LEU A 313
SHEET    4   H 4 HIS A 345  MET A 348 -1  O  GLU A 347   N  SER A 323
SHEET    1   I 4 HIS A 363  PHE A 364  0
SHEET    2   I 4 SER A 370  SER A 376 -1  O  TYR A 372   N  HIS A 363
SHEET    3   I 4 ARG A 382  GLN A 388 -1  O  PHE A 387   N  PHE A 371
SHEET    4   I 4 CYS A 394  PHE A 396 -1  O  THR A 395   N  TYR A 386
SHEET    1   J 4 VAL A 404  LEU A 410  0
SHEET    2   J 4 TYR A 414  SER A 419 -1  O  TYR A 416   N  ALA A 409
SHEET    3   J 4 ASN A 430  GLN A 435 -1  O  TYR A 432   N  TYR A 417
SHEET    4   J 4 ASP A 438  CYS A 444 -1  O  THR A 443   N  LYS A 433
SHEET    1   K 4 TYR A 457  PHE A 461  0
SHEET    2   K 4 TYR A 467  CYS A 472 -1  O  ARG A 471   N  SER A 458
SHEET    3   K 4 LEU A 479  SER A 484 -1  O  HIS A 483   N  TYR A 468
SHEET    4   K 4 LYS A 489  GLU A 495 -1  O  LEU A 494   N  TYR A 480
SHEET    1   L 8 SER A 511  LEU A 519  0
SHEET    2   L 8 THR A 522  LEU A 530 -1  O  TYR A 526   N  ASP A 515
SHEET    3   L 8 ILE A 574  PHE A 578 -1  O  VAL A 575   N  ILE A 529
SHEET    4   L 8 TYR A 540  VAL A 546  1  N  ASP A 545   O  ALA A 576
SHEET    5   L 8 VAL A 619  TRP A 629  1  O  ALA A 625   N  LEU A 544
SHEET    6   L 8 CYS A 649  VAL A 653  1  O  VAL A 653   N  GLY A 628
SHEET    7   L 8 GLU A 699  GLY A 705  1  O  ILE A 703   N  ALA A 652
SHEET    8   L 8 GLN A 731  TYR A 735  1  O  GLN A 731   N  TYR A 700
SHEET    1   M 4 LEU B  60  TRP B  62  0
SHEET    2   M 4 GLU B  67  LYS B  71 -1  O  LEU B  69   N  ARG B  61
SHEET    3   M 4 ASN B  75  ASN B  80 -1  O  PHE B  79   N  TYR B  68
SHEET    4   M 4 SER B  86  GLU B  91 -1  O  PHE B  89   N  ILE B  76
SHEET    1   N 4 ASP B 104  ILE B 107  0
SHEET    2   N 4 PHE B 113  LYS B 122 -1  O  GLU B 117   N  ASP B 104
SHEET    3   N 4 TYR B 128  ASP B 136 -1  O  ASP B 133   N  LEU B 116
SHEET    4   N 4 GLN B 141  LEU B 142 -1  O  GLN B 141   N  ASP B 136
SHEET    1   O 4 TRP B 154  TRP B 157  0
SHEET    2   O 4 LEU B 164  TRP B 168 -1  O  ALA B 165   N  THR B 156
SHEET    3   O 4 ASP B 171  LYS B 175 -1  O  LYS B 175   N  LEU B 164
SHEET    4   O 4 TYR B 183  ARG B 184 -1  O  TYR B 183   N  VAL B 174
SHEET    1   P 3 ILE B 194  ASN B 196  0
SHEET    2   P 3 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3   P 3 LEU B 214  TRP B 216 -1  N  TRP B 215   O  ALA B 224
SHEET    1   Q 4 ILE B 194  ASN B 196  0
SHEET    2   Q 4 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3   Q 4 THR B 265  ASN B 272 -1  O  PHE B 269   N  TYR B 225
SHEET    4   Q 4 SER B 284  ILE B 287 -1  O  ILE B 285   N  VAL B 270
SHEET    1   R 2 LEU B 235  PHE B 240  0
SHEET    2   R 2 LYS B 250  PRO B 255 -1  O  VAL B 252   N  TYR B 238
SHEET    1   S 4 HIS B 298  TRP B 305  0
SHEET    2   S 4 ARG B 310  ARG B 317 -1  O  SER B 312   N  THR B 304
SHEET    3   S 4 TYR B 322  TYR B 330 -1  O  ASP B 326   N  LEU B 313
SHEET    4   S 4 TRP B 337  ASN B 338 -1  O  ASN B 338   N  ASP B 329
SHEET    1   T 4 HIS B 298  TRP B 305  0
SHEET    2   T 4 ARG B 310  ARG B 317 -1  O  SER B 312   N  THR B 304
SHEET    3   T 4 TYR B 322  TYR B 330 -1  O  ASP B 326   N  LEU B 313
SHEET    4   T 4 HIS B 345  MET B 348 -1  O  GLU B 347   N  SER B 323
SHEET    1   U 4 HIS B 363  PHE B 364  0
SHEET    2   U 4 SER B 370  SER B 376 -1  O  TYR B 372   N  HIS B 363
SHEET    3   U 4 ARG B 382  GLN B 388 -1  O  PHE B 387   N  PHE B 371
SHEET    4   U 4 CYS B 394  PHE B 396 -1  O  THR B 395   N  TYR B 386
SHEET    1   V 4 VAL B 404  LEU B 410  0
SHEET    2   V 4 TYR B 414  SER B 419 -1  O  TYR B 416   N  ALA B 409
SHEET    3   V 4 ASN B 430  GLN B 435 -1  O  TYR B 432   N  TYR B 417
SHEET    4   V 4 ASP B 438  CYS B 444 -1  O  THR B 443   N  LYS B 433
SHEET    1   W 4 TYR B 457  PHE B 461  0
SHEET    2   W 4 TYR B 467  CYS B 472 -1  O  GLN B 469   N  SER B 460
SHEET    3   W 4 LEU B 479  SER B 484 -1  O  LEU B 479   N  CYS B 472
SHEET    4   W 4 LYS B 489  GLU B 495 -1  O  LEU B 494   N  TYR B 480
SHEET    1   X 8 SER B 511  LEU B 519  0
SHEET    2   X 8 THR B 522  LEU B 530 -1  O  TYR B 526   N  ASP B 515
SHEET    3   X 8 ILE B 574  PHE B 578 -1  O  VAL B 575   N  ILE B 529
SHEET    4   X 8 TYR B 540  VAL B 546  1  N  ASP B 545   O  ALA B 576
SHEET    5   X 8 VAL B 619  TRP B 629  1  O  ALA B 625   N  LEU B 544
SHEET    6   X 8 CYS B 649  VAL B 653  1  O  VAL B 653   N  GLY B 628
SHEET    7   X 8 GLU B 699  GLY B 705  1  O  ILE B 703   N  ALA B 652
SHEET    8   X 8 GLN B 731  TYR B 735  1  O  GLN B 731   N  TYR B 700
SSBOND   1 CYS A  328    CYS A  339                          1555   1555  2.06
SSBOND   2 CYS A  385    CYS A  394                          1555   1555  2.06
SSBOND   3 CYS A  444    CYS A  447                          1555   1555  2.03
SSBOND   4 CYS A  454    CYS A  472                          1555   1555  2.08
SSBOND   5 CYS A  649    CYS A  762                          1555   1555  2.05
SSBOND   6 CYS B  328    CYS B  339                          1555   1555  2.05
SSBOND   7 CYS B  385    CYS B  394                          1555   1555  2.06
SSBOND   8 CYS B  444    CYS B  447                          1555   1555  2.04
SSBOND   9 CYS B  454    CYS B  472                          1555   1555  2.10
SSBOND  10 CYS B  649    CYS B  762                          1555   1555  2.05
LINK         O4  NAG A 796                 C1  NAG A 798     1555   1555  1.38
LINK         ND2 ASN A 219                 C1  NAG A 799     1555   1555  1.43
LINK         ND2 ASN A  85                 C1  NAG A 794     1555   1555  1.45
LINK         O4  NAG A 798                 C1  BMA A 802     1555   1555  1.45
LINK         ND2 ASN A 229                 C1  NAG A 796     1555   1555  1.46
LINK         O4  NAG B 794                 C1  NAG B 797     1555   1555  1.46
LINK         ND2 ASN B 229                 C1  NAG B 796     1555   1555  1.46
LINK         ND2 ASN B  85                 C1  NAG B 794     1555   1555  1.46
LINK         O4  NAG A 794                 C1  NAG A 797     1555   1555  1.49
LINK         ND2 ASN A 321                 C1  NAG A 800     1555   1555  1.52
LINK         O3  BMA A 802                 C1  MAN A 803     1555   1555  1.54
CISPEP   1 GLY A  474    PRO A  475          0         2.36
CISPEP   2 GLY B  474    PRO B  475          0         1.74
SITE     1 AC1  8 VAL A  78  ASN A  85  SER A  86  SER A  87
SITE     2 AC1  8 TYR A 386  GLN A 388  THR A 395  NAG A 797
SITE     1 AC2  1 NAG A 794
SITE     1 AC3  6 ILE A 194  ASN A 229  THR A 231  GLU A 232
SITE     2 AC3  6 LYS A 267  NAG A 798
SITE     1 AC4  3 GLU A 232  NAG A 796  BMA A 802
SITE     1 AC5  2 NAG A 798  MAN A 803
SITE     1 AC6  2 BMA A 802  SER B 690
SITE     1 AC7  4 ASN A 219  THR A 221  GLN A 308  GLU A 309
SITE     1 AC8  2 ASN A 321  SER A 349
SITE     1 AC9 11 ARG A 125  GLU A 205  GLU A 206  SER A 209
SITE     2 AC9 11 PHE A 357  ARG A 358  TYR A 547  TYR A 662
SITE     3 AC9 11 TYR A 666  ASN A 710  VAL A 711
SITE     1 BC1  9 GLU B  67  VAL B  78  ASN B  85  SER B  86
SITE     2 BC1  9 SER B  87  TYR B 386  GLN B 388  THR B 395
SITE     3 BC1  9 NAG B 797
SITE     1 BC2  2 LYS B 391  NAG B 794
SITE     1 BC3  5 ILE B 194  ASN B 229  THR B 231  GLU B 232
SITE     2 BC3  5 LYS B 267
SITE     1 BC4 11 ARG B 125  GLU B 205  GLU B 206  SER B 209
SITE     2 BC4 11 PHE B 357  ARG B 358  TYR B 547  TYR B 662
SITE     3 BC4 11 TYR B 666  ASN B 710  VAL B 711
CRYST1   65.751   69.082  424.160  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.015209  0.000000  0.000000        0.00000
SCALE2      0.000000  0.014476  0.000000        0.00000
SCALE3      0.000000  0.000000  0.002358        0.00000
TER    5958      PRO A 766
TER   11916      PRO B 766
MASTER      454    0   13   38   98    0   22    612183    2  224  116
END