longtext: 3RAR-pdb

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HEADER    HYDROLASE                               28-MAR-11   3RAR
TITLE     X-RAY STRUCTURE OF A BOUND PHOSPHONATE TRANSITION STATE ANALOG AND
TITLE    2 ENANTIOSELECTIVITY OF CANDIDA RUGOSA LIPASE TOWARD CHIRAL CARBOXYLIC
TITLE    3 ACIDS
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE 1;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: UNP RESIDUES 16-549;
COMPND   5 EC: 3.1.1.3;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: CANDIDA RUGOSA;
SOURCE   3 ORGANISM_COMMON: YEAST;
SOURCE   4 ORGANISM_TAXID: 5481;
SOURCE   5 GENE: LIP1;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    ALPHA/BETA HYDROLASE FOLD, HYDROLASE, CALCIUM BINDING, (RC,RP)-
KEYWDS   2 METHYL[1-(METHOXY)PHENYLMETHYL]PHOSPHONYL ADDUCT AT O OF SERINE 209
EXPDTA    X-RAY DIFFRACTION
AUTHOR    P.GROCHULSKI,R.KAZLAUSKAS
REVDAT   1   27-APR-11 3RAR    0
JRNL        AUTH   P.GROCHULSKI,R.KAZLAUSKAS
JRNL        TITL   X-RAY STRUCTURE OF A BOUND PHOSPHONATE TRANSITION STATE
JRNL        TITL 2 ANALOG AND ENANTIOSELECTIVITY OF CANDIDA RUGOSA LIPASE
JRNL        TITL 3 TOWARD CHIRAL CARBOXYLIC ACIDS
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.19 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.3
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.19
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 54.10
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 1.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 564032.690
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 80.9
REMARK   3   NUMBER OF REFLECTIONS             : 23670
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.171
REMARK   3   FREE R VALUE                     : 0.211
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 9.900
REMARK   3   FREE R VALUE TEST SET COUNT      : 2334
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.004
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.19
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.33
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 62.50
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 2716
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2330
REMARK   3   BIN FREE R VALUE                    : 0.2730
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 9.90
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 297
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.016
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4022
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 57
REMARK   3   SOLVENT ATOMS            : 174
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 10.90
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 22.50
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -0.62000
REMARK   3    B22 (A**2) : -0.80000
REMARK   3    B33 (A**2) : 1.42000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : -0.00000
REMARK   3    B23 (A**2) : -0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.21
REMARK   3   ESD FROM SIGMAA              (A) : 0.27
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.26
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.29
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.006
REMARK   3   BOND ANGLES            (DEGREES) : 1.30
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 23.40
REMARK   3   IMPROPER ANGLES        (DEGREES) : 1.76
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.160 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 1.810 ; 2.000
REMARK   3   SIDE-CHAIN BOND              (A**2) : 1.880 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 2.630 ; 2.500
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : FLAT MODEL
REMARK   3   KSOL        : 0.37
REMARK   3   BSOL        : 76.86
REMARK   3
REMARK   3  NCS MODEL : NONE
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM
REMARK   3  PARAMETER FILE  2  : DNA-RNA_REP.PARAM
REMARK   3  PARAMETER FILE  3  : WATER_REP.PARAM
REMARK   3  PARAMETER FILE  4  : ION.PARAM
REMARK   3  PARAMETER FILE  5  : CARBOHYDRATE.PARAM
REMARK   3  PARAMETER FILE  6  : IAN.PAR
REMARK   3  PARAMETER FILE  7  : NULL
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP
REMARK   3  TOPOLOGY FILE  2   : DNA-RNA.TOP
REMARK   3  TOPOLOGY FILE  3   : WATER.TOP
REMARK   3  TOPOLOGY FILE  4   : ION.TOP
REMARK   3  TOPOLOGY FILE  5   : CARBOHYDRATE.TOP
REMARK   3  TOPOLOGY FILE  6   : IAN.TOP
REMARK   3  TOPOLOGY FILE  7   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3RAR COMPLIES WITH FORMAT V. 3.20, 01-DEC-08
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 29-MAR-11.
REMARK 100 THE RCSB ID CODE IS RCSB064700.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 17-JUL-95; NULL
REMARK 200  TEMPERATURE           (KELVIN) : 298; NULL
REMARK 200  PH                             : 5.3
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N; NULL
REMARK 200  RADIATION SOURCE               : ROTATING ANODE; NULL
REMARK 200  BEAMLINE                       : NULL; NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU; OTHER
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M; NULL
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418; NULL
REMARK 200  MONOCHROMATOR                  : GRAPHITE; NULL
REMARK 200  OPTICS                         : NULL; NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE; NULL
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU RAXIS IIC; NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : RAXIS
REMARK 200  DATA SCALING SOFTWARE          : RAXIS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 23670
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.190
REMARK 200  RESOLUTION RANGE LOW       (A) : 54.100
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 81.0
REMARK 200  DATA REDUNDANCY                : NULL
REMARK 200  R MERGE                    (I) : 0.09200
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.19
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.25
REMARK 200  COMPLETENESS FOR SHELL     (%) : 58.1
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.23500
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH; NULL
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: FOURIER SYNTHESIS
REMARK 200 SOFTWARE USED: CNS
REMARK 200 STARTING MODEL: PDB ENTRY 1LPP
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 49.62
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.44
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 40% (V/V) 2-METHYL-2,4-PENTANEDIOL, 30
REMARK 280  MM SODIUM ACETATE BUFFER AND 30 MM CACL2., PH 5.3, VAPOR
REMARK 280  DIFFUSION, HANGING DROP, TEMPERATURE 296K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       88.05000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       88.05000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       32.50000
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       48.75000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       32.50000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       48.75000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       88.05000
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       32.50000
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       48.75000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000       88.05000
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       32.50000
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       48.75000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 2830 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 36250 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -15.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000      130.00000
REMARK 350   BIOMT2   2  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      -88.05000
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ILE A  18      -61.94     74.28
REMARK 500    PRO A  32       50.46    -69.69
REMARK 500    PHE A  38       -4.65     62.44
REMARK 500    THR A  68      156.90    179.34
REMARK 500    PHE A 125       19.66     59.45
REMARK 500    SER A 159     -154.00     66.06
REMARK 500    SER A 209     -116.90     55.70
REMARK 500    SER A 241       75.18     66.04
REMARK 500    ALA A 243     -121.76   -116.37
REMARK 500    SER A 301     -121.97     43.24
REMARK 500    VAL A 313      -59.36   -125.47
REMARK 500    ASP A 317     -178.53   -171.41
REMARK 500    ASP A 340       69.50   -112.57
REMARK 500    ASP A 384      106.47    -29.74
REMARK 500    PHE A 415      -71.91   -138.71
REMARK 500    VAL A 444      -46.00     67.96
REMARK 500    ASN A 468      -80.60   -117.36
REMARK 500
REMARK 500 REMARK: NULL
REMARK 610
REMARK 610 MISSING HETEROATOM
REMARK 610 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 610 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 610 I=INSERTION CODE):
REMARK 610   M RES C SSEQI
REMARK 610     IAN A  560
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CA A 590  CA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLY A 326   O
REMARK 620 2 HOH A 538   O   128.2
REMARK 620 N                    1
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CA A 589  CA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 ASP A 260   OD1
REMARK 620 2 HOH A 682   O    82.2
REMARK 620 3 ASP A 260   OD2  45.4 101.4
REMARK 620 N                    1     2
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 991
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 992
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 994
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CA A 589
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CA A 590
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE IAN A 560
DBREF  3RAR A    1   534  UNP    P20261   LIP1_CANRU      16    549
SEQRES   1 A  534  ALA PRO THR ALA THR LEU ALA ASN GLY ASP THR ILE THR
SEQRES   2 A  534  GLY LEU ASN ALA ILE ILE ASN GLU ALA PHE LEU GLY ILE
SEQRES   3 A  534  PRO PHE ALA GLU PRO PRO VAL GLY ASN LEU ARG PHE LYS
SEQRES   4 A  534  ASP PRO VAL PRO TYR SER GLY SER LEU ASP GLY GLN LYS
SEQRES   5 A  534  PHE THR SER TYR GLY PRO SER CYS MET GLN GLN ASN PRO
SEQRES   6 A  534  GLU GLY THR TYR GLU GLU ASN LEU PRO LYS ALA ALA LEU
SEQRES   7 A  534  ASP LEU VAL MET GLN SER LYS VAL PHE GLU ALA VAL SER
SEQRES   8 A  534  PRO SER SER GLU ASP CYS LEU THR ILE ASN VAL VAL ARG
SEQRES   9 A  534  PRO PRO GLY THR LYS ALA GLY ALA ASN LEU PRO VAL MET
SEQRES  10 A  534  LEU TRP ILE PHE GLY GLY GLY PHE GLU VAL GLY GLY THR
SEQRES  11 A  534  SER THR PHE PRO PRO ALA GLN MET ILE THR LYS SER ILE
SEQRES  12 A  534  ALA MET GLY LYS PRO ILE ILE HIS VAL SER VAL ASN TYR
SEQRES  13 A  534  ARG VAL SER SER TRP GLY PHE LEU ALA GLY ASP GLU ILE
SEQRES  14 A  534  LYS ALA GLU GLY SER ALA ASN ALA GLY LEU LYS ASP GLN
SEQRES  15 A  534  ARG LEU GLY MET GLN TRP VAL ALA ASP ASN ILE ALA ALA
SEQRES  16 A  534  PHE GLY GLY ASP PRO THR LYS VAL THR ILE PHE GLY GLU
SEQRES  17 A  534  SER ALA GLY SER MET SER VAL MET CYS HIS ILE LEU TRP
SEQRES  18 A  534  ASN ASP GLY ASP ASN THR TYR LYS GLY LYS PRO LEU PHE
SEQRES  19 A  534  ARG ALA GLY ILE MET GLN SER GLY ALA MET VAL PRO SER
SEQRES  20 A  534  ASP ALA VAL ASP GLY ILE TYR GLY ASN GLU ILE PHE ASP
SEQRES  21 A  534  LEU LEU ALA SER ASN ALA GLY CYS GLY SER ALA SER ASP
SEQRES  22 A  534  LYS LEU ALA CYS LEU ARG GLY VAL SER SER ASP THR LEU
SEQRES  23 A  534  GLU ASP ALA THR ASN ASN THR PRO GLY PHE LEU ALA TYR
SEQRES  24 A  534  SER SER LEU ARG LEU SER TYR LEU PRO ARG PRO ASP GLY
SEQRES  25 A  534  VAL ASN ILE THR ASP ASP MET TYR ALA LEU VAL ARG GLU
SEQRES  26 A  534  GLY LYS TYR ALA ASN ILE PRO VAL ILE ILE GLY ASP GLN
SEQRES  27 A  534  ASN ASP GLU GLY THR PHE PHE GLY THR SER SER LEU ASN
SEQRES  28 A  534  VAL THR THR ASP ALA GLN ALA ARG GLU TYR PHE LYS GLN
SEQRES  29 A  534  SER PHE VAL HIS ALA SER ASP ALA GLU ILE ASP THR LEU
SEQRES  30 A  534  MET THR ALA TYR PRO GLY ASP ILE THR GLN GLY SER PRO
SEQRES  31 A  534  PHE ASP THR GLY ILE LEU ASN ALA LEU THR PRO GLN PHE
SEQRES  32 A  534  LYS ARG ILE SER ALA VAL LEU GLY ASP LEU GLY PHE THR
SEQRES  33 A  534  LEU ALA ARG ARG TYR PHE LEU ASN HIS TYR THR GLY GLY
SEQRES  34 A  534  THR LYS TYR SER PHE LEU SER LYS GLN LEU SER GLY LEU
SEQRES  35 A  534  PRO VAL LEU GLY THR PHE HIS SER ASN ASP ILE VAL PHE
SEQRES  36 A  534  GLN ASP TYR LEU LEU GLY SER GLY SER LEU ILE TYR ASN
SEQRES  37 A  534  ASN ALA PHE ILE ALA PHE ALA THR ASP LEU ASP PRO ASN
SEQRES  38 A  534  THR ALA GLY LEU LEU VAL LYS TRP PRO GLU TYR THR SER
SEQRES  39 A  534  SER SER GLN SER GLY ASN ASN LEU MET MET ILE ASN ALA
SEQRES  40 A  534  LEU GLY LEU TYR THR GLY LYS ASP ASN PHE ARG THR ALA
SEQRES  41 A  534  GLY TYR ASP ALA LEU PHE SER ASN PRO PRO SER PHE PHE
SEQRES  42 A  534  VAL
MODRES 3RAR ASN A  351  ASN  GLYCOSYLATION SITE
MODRES 3RAR ASN A  314  ASN  GLYCOSYLATION SITE
HET    NAG  A 991      14
HET    NAG  A 992      14
HET    NAG  A 994      14
HET     CA  A 589       1
HET     CA  A 590       1
HET    IAN  A 560      13
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM      CA CALCIUM ION
HETNAM     IAN METHYL HYDROGEN (R)-[(R)-METHOXY(PHENYL)
HETNAM   2 IAN  METHYL]PHOSPHONATE
FORMUL   2  NAG    3(C8 H15 N O6)
FORMUL   4   CA    2(CA 2+)
FORMUL   6  IAN    C9 H13 O4 P
FORMUL   7  HOH   *174(H2 O)
HELIX    1   1 VAL A   33  ARG A   37  5                                   5
HELIX    2   2 ASN A   72  GLN A   83  1                                  12
HELIX    3   3 SER A   84  SER A   91  1                                   8
HELIX    4   4 GLY A  129  PHE A  133  5                                   5
HELIX    5   5 PRO A  135  MET A  145  1                                  11
HELIX    6   6 SER A  159  LEU A  164  1                                   6
HELIX    7   7 GLY A  166  GLY A  173  1                                   8
HELIX    8   8 ASN A  176  ILE A  193  1                                  18
HELIX    9   9 ALA A  194  PHE A  196  5                                   3
HELIX   10  10 SER A  209  TRP A  221  1                                  13
HELIX   11  11 ASN A  222  ASP A  225  5                                   4
HELIX   12  12 GLY A  252  ALA A  266  1                                  15
HELIX   13  13 ASP A  273  ARG A  279  1                                   7
HELIX   14  14 SER A  282  ASN A  291  1                                  10
HELIX   15  15 ASP A  318  GLU A  325  1                                   8
HELIX   16  16 GLY A  342  THR A  347  1                                   6
HELIX   17  17 SER A  348  LEU A  350  5                                   3
HELIX   18  18 THR A  354  PHE A  366  1                                  13
HELIX   19  19 SER A  370  TYR A  381  1                                  12
HELIX   20  20 ASP A  384  GLY A  388  5                                   5
HELIX   21  21 GLN A  402  PHE A  415  1                                  14
HELIX   22  22 PHE A  415  ASN A  424  1                                  10
HELIX   23  23 SER A  450  TYR A  458  1                                   9
HELIX   24  24 GLY A  463  ASN A  468  1                                   6
HELIX   25  25 ASN A  468  LEU A  478  1                                  11
HELIX   26  26 ASP A  479  GLY A  484  5                                   6
HELIX   27  27 ARG A  518  SER A  527  1                                  10
HELIX   28  28 ASN A  528  PHE A  533  5                                   6
SHEET    1   A 2 THR A   3  THR A   5  0
SHEET    2   A 2 THR A  11  THR A  13 -1  O  ILE A  12   N  ALA A   4
SHEET    1   B11 LEU A  15  ASN A  16  0
SHEET    2   B11 GLU A  21  PRO A  27 -1  O  ALA A  22   N  LEU A  15
SHEET    3   B11 THR A  99  ARG A 104 -1  O  ILE A 100   N  ILE A  26
SHEET    4   B11 ILE A 150  VAL A 154 -1  O  HIS A 151   N  VAL A 103
SHEET    5   B11 LEU A 114  ILE A 120  1  N  TRP A 119   O  VAL A 152
SHEET    6   B11 GLY A 198  GLU A 208  1  O  LYS A 202   N  VAL A 116
SHEET    7   B11 ALA A 236  GLN A 240  1  O  ALA A 236   N  ILE A 205
SHEET    8   B11 VAL A 333  GLN A 338  1  O  ILE A 334   N  MET A 239
SHEET    9   B11 LYS A 431  SER A 436  1  O  SER A 436   N  ASP A 337
SHEET   10   B11 LEU A 502  ILE A 505  1  O  MET A 503   N  SER A 433
SHEET   11   B11 LEU A 510  GLY A 513 -1  O  TYR A 511   N  MET A 504
SHEET    1   C 2 THR A 227  TYR A 228  0
SHEET    2   C 2 LYS A 231  PRO A 232 -1  O  LYS A 231   N  TYR A 228
SSBOND   1 CYS A   60    CYS A   97                          1555   1555  2.03
SSBOND   2 CYS A  268    CYS A  277                          1555   1555  2.03
LINK         O4  NAG A 992                 C1  NAG A 994     1555   1555  1.40
LINK         ND2 ASN A 351                 C1  NAG A 992     1555   1555  1.45
LINK         ND2 ASN A 314                 C1  NAG A 991     1555   1555  1.46
LINK         O   GLY A 326                CA    CA A 590     1555   1555  2.72
LINK         OD1 ASP A 260                CA    CA A 589     1555   1555  2.77
LINK        CA    CA A 590                 O   HOH A 538     1555   1555  2.81
LINK        CA    CA A 589                 O   HOH A 682     1555   1555  2.86
LINK         OD2 ASP A 260                CA    CA A 589     1555   1555  2.91
LINK         OG  SER A 209                 P1  IAN A 560     1555   1555  1.65
CISPEP   1 SER A  389    PRO A  390          0         0.02
SITE     1 AC1  5 TRP A 221  ASN A 222  VAL A 313  ASN A 314
SITE     2 AC1  5 HOH A 709
SITE     1 AC2  5 GLU A  70  TYR A 299  ASN A 351  GLN A 357
SITE     2 AC2  5 NAG A 994
SITE     1 AC3  3 TYR A  69  GLU A  70  NAG A 992
SITE     1 AC4  2 ASP A 260  HOH A 682
SITE     1 AC5  2 GLY A 326  HOH A 538
SITE     1 AC6 11 GLY A 122  GLY A 123  GLY A 124  SER A 209
SITE     2 AC6 11 ALA A 210  MET A 213  PHE A 296  LEU A 302
SITE     3 AC6 11 LEU A 304  PHE A 345  HIS A 449
CRYST1   65.000   97.500  176.100  90.00  90.00  90.00 C 2 2 21      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.015385  0.000000  0.000000        0.00000
SCALE2      0.000000  0.010256  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005679        0.00000
TER    4023      VAL A 534
MASTER      336    0    6   28   15    0   10    6 4253    1   69   42
END