longtext: 3VJK-pdb

content
HEADER    HYDROLASE/HYDROLASE INHIBITOR           24-OCT-11   3VJK
TITLE     CRYSTAL STRUCTURE OF HUMAN DEPIPTIDYL PEPTIDASE IV (DPP-4) IN COMPLEX
TITLE    2 WITH MP-513
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: DIPEPTIDYL PEPTIDASE 4;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: UNP RESIDUES 33-766;
COMPND   5 SYNONYM: ADABP, ADENOSINE DEAMINASE COMPLEXING PROTEIN 2, ADCP-2,
COMPND   6 DIPEPTIDYL PEPTIDASE IV, DPP IV, T-CELL ACTIVATION ANTIGEN CD26,
COMPND   7 TP103, DIPEPTIDYL PEPTIDASE 4 MEMBRANE FORM, DIPEPTIDYL PEPTIDASE IV
COMPND   8 MEMBRANE FORM, DIPEPTIDYL PEPTIDASE 4 SOLUBLE FORM, DIPEPTIDYL
COMPND   9 PEPTIDASE IV SOLUBLE FORM;
COMPND  10 EC: 3.4.14.5;
COMPND  11 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: DPP4, ADCP2, CD26;
SOURCE   6 EXPRESSION_SYSTEM: SPODOPTERA FRUGIPERDA;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 7108;
SOURCE   8 EXPRESSION_SYSTEM_CELL_LINE: EXPRESSF+;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: BACULOVIRUS;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PPSC8
KEYWDS    ALPHA/BETA, BETA-PROPELLER, AMINOPEPTIDASE, SERINE PROTEASE, SIGNAL-
KEYWDS   2 ANCHOR, TRANSMEMBRANE, DIABETES, GLYCOPROTEIN, CELL MEMBRANE,
KEYWDS   3 HYDROLASE-HYDROLASE INHIBITOR COMPLEX
EXPDTA    X-RAY DIFFRACTION
AUTHOR    F.AKAHOSHI,H.KISHIDA,I.MIYAGUCHI,T.YOSHIDA,S.ISHII
REVDAT   1   24-OCT-12 3VJK    0
JRNL        AUTH   T.YOSHIDA,F.AKAHOSHI,H.SAKASHITA,H.KITAJIMA,M.NAKAMURA,
JRNL        AUTH 2 S.SONDA,M.TAKEUCHI,Y.TANAKA,N.UEDA,S.SEKIGUCHI,T.ISHIGE,
JRNL        AUTH 3 K.SHIMA,M.NABENO,Y.ABE,J.ANABUKI,A.SOEJIMA,K.YOSHIDA,
JRNL        AUTH 4 Y.TAKASHINA,S.ISHII,S.KIUCHI,S.FUKUDA,R.TSUTSUMIUCHI,
JRNL        AUTH 5 K.KOSAKA,T.MUROZONO,Y.NAKAMARU,H.UTSUMI,N.MASUTOMI,
JRNL        AUTH 6 H.KISHIDA,I.MIYAGUCHI,Y.HAYASHI
JRNL        TITL   DISCOVERY AND PRECLINICAL PROFILE OF TENELIGLIPTIN
JRNL        TITL 2 (3-[(2S,4S)-4-[4-(3-METHYL-1-PHENYL-1H-PYRAZOL-5-YL)
JRNL        TITL 3 PIPERAZIN-1-YL]PYRROLIDIN-2-YLCARBONYL]THIAZOLIDINE): A
JRNL        TITL 4 HIGHLY POTENT, SELECTIVE, LONG-LASTING AND ORALLY ACTIVE
JRNL        TITL 5 DIPEPTIDYL PEPTIDASE IV INHIBITOR FOR THE TREATMENT OF TYPE
JRNL        TITL 6 2 DIABETES
JRNL        REF    BIOORG.MED.CHEM.              V.  20  5705 2012
JRNL        REFN                   ISSN 0968-0896
JRNL        PMID   22959556
JRNL        DOI    10.1016/J.BMC.2012.08.012
REMARK   2
REMARK   2 RESOLUTION.    2.49 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.5.0110
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.49
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 30.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 88.7
REMARK   3   NUMBER OF REFLECTIONS             : 61379
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.228
REMARK   3   R VALUE            (WORKING SET) : 0.225
REMARK   3   FREE R VALUE                     : 0.279
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 3211
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.49
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.55
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 4707
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 93.63
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2810
REMARK   3   BIN FREE R VALUE SET COUNT          : 243
REMARK   3   BIN FREE R VALUE                    : 0.3330
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 11942
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 298
REMARK   3   SOLVENT ATOMS            : 456
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 51.40
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 37.50
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -2.41000
REMARK   3    B22 (A**2) : 3.66000
REMARK   3    B33 (A**2) : -1.25000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): NULL
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.334
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.213
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 9.424
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.929
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.897
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A): 12619 ; 0.011 ; 0.022
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 17188 ; 1.435 ; 1.958
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  1456 ; 6.439 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   618 ;34.688 ;23.981
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1996 ;16.119 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    60 ;20.168 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  1837 ; 0.092 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  9721 ; 0.005 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  7262 ; 0.551 ; 1.500
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2): 11782 ; 1.047 ; 2.000
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  5357 ; 1.508 ; 3.000
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  5406 ; 2.488 ; 4.500
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.40
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3VJK COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 09-NOV-11.
REMARK 100 THE RCSB ID CODE IS RCSB095124.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 28-NOV-06
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.8
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PHOTON FACTORY
REMARK 200  BEAMLINE                       : BL-5A
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0000
REMARK 200  MONOCHROMATOR                  : SI 111
REMARK 200  OPTICS                         : MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : AREA DETECTOR
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 65671
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.490
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 90.2
REMARK 200  DATA REDUNDANCY                : 3.800
REMARK 200  R MERGE                    (I) : 0.09200
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 15.6000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.49
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.58
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.7
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.10
REMARK 200  R MERGE FOR SHELL          (I) : 0.49100
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 1J2E
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 58.94
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.00
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 20% PEG 4000, 0.18M SODIUM ACETATE,
REMARK 280  0.18M GLYCINE-SODIUM HYDROXIDE , PH 8.8, VAPOR DIFFUSION, SITTING
REMARK 280  DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       58.97750
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       69.00550
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       63.20650
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       69.00550
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       58.97750
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       63.20650
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 9250 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 60130 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: 46.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ASP A    33
REMARK 465     ASP A    34
REMARK 465     ALA A    35
REMARK 465     THR A    36
REMARK 465     ALA A    37
REMARK 465     HIS A   767
REMARK 465     HIS A   768
REMARK 465     HIS A   769
REMARK 465     HIS A   770
REMARK 465     HIS A   771
REMARK 465     HIS A   772
REMARK 465     ASP B    33
REMARK 465     ASP B    34
REMARK 465     ALA B    35
REMARK 465     THR B    36
REMARK 465     ALA B    37
REMARK 465     HIS B   767
REMARK 465     HIS B   768
REMARK 465     HIS B   769
REMARK 465     HIS B   770
REMARK 465     HIS B   771
REMARK 465     HIS B   772
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    GLU A  97       32.86    -91.59
REMARK 500    GLN A 123      -95.40    -97.40
REMARK 500    TRP A 124     -145.46   -100.56
REMARK 500    HIS A 162       30.14   -150.01
REMARK 500    ILE A 193      -54.87   -126.86
REMARK 500    SER A 209       46.52     38.40
REMARK 500    SER A 242     -171.95     70.64
REMARK 500    GLN A 320       42.03    -80.90
REMARK 500    PHE A 357      -35.60   -132.98
REMARK 500    ASN A 377     -168.81    -72.89
REMARK 500    ALA A 409      154.02    179.32
REMARK 500    ASN A 450       72.82   -166.13
REMARK 500    ARG A 492      158.70    179.14
REMARK 500    GLU A 521       37.37     77.85
REMARK 500    TYR A 547      -79.32   -120.94
REMARK 500    ARG A 597       41.09   -140.23
REMARK 500    THR A 600      -97.41   -118.31
REMARK 500    LYS A 615      -19.62    -48.70
REMARK 500    SER A 630     -123.54     56.20
REMARK 500    ASP A 678      -94.06   -107.11
REMARK 500    ASN A 710      -68.49    -94.67
REMARK 500    ILE A 742       48.68     26.84
REMARK 500    TYR B  58       76.82   -152.74
REMARK 500    ASN B  74      -15.05     68.62
REMARK 500    GLU B  97       62.78   -101.89
REMARK 500    GLN B 123     -105.04   -102.47
REMARK 500    TRP B 124     -143.02    -90.66
REMARK 500    TRP B 154      141.71   -173.18
REMARK 500    HIS B 162       34.17   -159.42
REMARK 500    LYS B 190      114.97   -163.54
REMARK 500    GLU B 191      126.73    -33.36
REMARK 500    ASP B 192        9.48     58.06
REMARK 500    ILE B 193      -57.84   -126.55
REMARK 500    SER B 242     -162.23     61.37
REMARK 500    GLN B 320       43.48    -82.93
REMARK 500    ALA B 342      -14.76    -49.64
REMARK 500    ASN B 450       78.69   -163.38
REMARK 500    GLU B 521       11.06     58.24
REMARK 500    TYR B 547      -72.71   -126.94
REMARK 500    TYR B 585       19.54     80.40
REMARK 500    THR B 600      -90.00   -115.51
REMARK 500    SER B 630     -124.38     63.92
REMARK 500    ASP B 678      -94.90   -123.64
REMARK 500    ASN B 710      -77.15    -92.13
REMARK 500    ASP B 739     -152.48    -99.78
REMARK 500    ILE B 742       60.12     31.60
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE M51 A 1000
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1085
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1092
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1219
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1220
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1229
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1230
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1281
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1282
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 1321
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE M51 B 1000
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1085
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1150
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1219
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1220
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1229
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1230
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1281
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 1321
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3VJL   RELATED DB: PDB
REMARK 900 RELATED ID: 3VJM   RELATED DB: PDB
DBREF  3VJK A   33   766  UNP    P27487   DPP4_HUMAN      33    766
DBREF  3VJK B   33   766  UNP    P27487   DPP4_HUMAN      33    766
SEQADV 3VJK HIS A  767  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS A  768  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS A  769  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS A  770  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS A  771  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS A  772  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS B  767  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS B  768  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS B  769  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS B  770  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS B  771  UNP  P27487              EXPRESSION TAG
SEQADV 3VJK HIS B  772  UNP  P27487              EXPRESSION TAG
SEQRES   1 A  740  ASP ASP ALA THR ALA ASP SER ARG LYS THR TYR THR LEU
SEQRES   2 A  740  THR ASP TYR LEU LYS ASN THR TYR ARG LEU LYS LEU TYR
SEQRES   3 A  740  SER LEU ARG TRP ILE SER ASP HIS GLU TYR LEU TYR LYS
SEQRES   4 A  740  GLN GLU ASN ASN ILE LEU VAL PHE ASN ALA GLU TYR GLY
SEQRES   5 A  740  ASN SER SER VAL PHE LEU GLU ASN SER THR PHE ASP GLU
SEQRES   6 A  740  PHE GLY HIS SER ILE ASN ASP TYR SER ILE SER PRO ASP
SEQRES   7 A  740  GLY GLN PHE ILE LEU LEU GLU TYR ASN TYR VAL LYS GLN
SEQRES   8 A  740  TRP ARG HIS SER TYR THR ALA SER TYR ASP ILE TYR ASP
SEQRES   9 A  740  LEU ASN LYS ARG GLN LEU ILE THR GLU GLU ARG ILE PRO
SEQRES  10 A  740  ASN ASN THR GLN TRP VAL THR TRP SER PRO VAL GLY HIS
SEQRES  11 A  740  LYS LEU ALA TYR VAL TRP ASN ASN ASP ILE TYR VAL LYS
SEQRES  12 A  740  ILE GLU PRO ASN LEU PRO SER TYR ARG ILE THR TRP THR
SEQRES  13 A  740  GLY LYS GLU ASP ILE ILE TYR ASN GLY ILE THR ASP TRP
SEQRES  14 A  740  VAL TYR GLU GLU GLU VAL PHE SER ALA TYR SER ALA LEU
SEQRES  15 A  740  TRP TRP SER PRO ASN GLY THR PHE LEU ALA TYR ALA GLN
SEQRES  16 A  740  PHE ASN ASP THR GLU VAL PRO LEU ILE GLU TYR SER PHE
SEQRES  17 A  740  TYR SER ASP GLU SER LEU GLN TYR PRO LYS THR VAL ARG
SEQRES  18 A  740  VAL PRO TYR PRO LYS ALA GLY ALA VAL ASN PRO THR VAL
SEQRES  19 A  740  LYS PHE PHE VAL VAL ASN THR ASP SER LEU SER SER VAL
SEQRES  20 A  740  THR ASN ALA THR SER ILE GLN ILE THR ALA PRO ALA SER
SEQRES  21 A  740  MET LEU ILE GLY ASP HIS TYR LEU CYS ASP VAL THR TRP
SEQRES  22 A  740  ALA THR GLN GLU ARG ILE SER LEU GLN TRP LEU ARG ARG
SEQRES  23 A  740  ILE GLN ASN TYR SER VAL MET ASP ILE CYS ASP TYR ASP
SEQRES  24 A  740  GLU SER SER GLY ARG TRP ASN CYS LEU VAL ALA ARG GLN
SEQRES  25 A  740  HIS ILE GLU MET SER THR THR GLY TRP VAL GLY ARG PHE
SEQRES  26 A  740  ARG PRO SER GLU PRO HIS PHE THR LEU ASP GLY ASN SER
SEQRES  27 A  740  PHE TYR LYS ILE ILE SER ASN GLU GLU GLY TYR ARG HIS
SEQRES  28 A  740  ILE CYS TYR PHE GLN ILE ASP LYS LYS ASP CYS THR PHE
SEQRES  29 A  740  ILE THR LYS GLY THR TRP GLU VAL ILE GLY ILE GLU ALA
SEQRES  30 A  740  LEU THR SER ASP TYR LEU TYR TYR ILE SER ASN GLU TYR
SEQRES  31 A  740  LYS GLY MET PRO GLY GLY ARG ASN LEU TYR LYS ILE GLN
SEQRES  32 A  740  LEU SER ASP TYR THR LYS VAL THR CYS LEU SER CYS GLU
SEQRES  33 A  740  LEU ASN PRO GLU ARG CYS GLN TYR TYR SER VAL SER PHE
SEQRES  34 A  740  SER LYS GLU ALA LYS TYR TYR GLN LEU ARG CYS SER GLY
SEQRES  35 A  740  PRO GLY LEU PRO LEU TYR THR LEU HIS SER SER VAL ASN
SEQRES  36 A  740  ASP LYS GLY LEU ARG VAL LEU GLU ASP ASN SER ALA LEU
SEQRES  37 A  740  ASP LYS MET LEU GLN ASN VAL GLN MET PRO SER LYS LYS
SEQRES  38 A  740  LEU ASP PHE ILE ILE LEU ASN GLU THR LYS PHE TRP TYR
SEQRES  39 A  740  GLN MET ILE LEU PRO PRO HIS PHE ASP LYS SER LYS LYS
SEQRES  40 A  740  TYR PRO LEU LEU LEU ASP VAL TYR ALA GLY PRO CYS SER
SEQRES  41 A  740  GLN LYS ALA ASP THR VAL PHE ARG LEU ASN TRP ALA THR
SEQRES  42 A  740  TYR LEU ALA SER THR GLU ASN ILE ILE VAL ALA SER PHE
SEQRES  43 A  740  ASP GLY ARG GLY SER GLY TYR GLN GLY ASP LYS ILE MET
SEQRES  44 A  740  HIS ALA ILE ASN ARG ARG LEU GLY THR PHE GLU VAL GLU
SEQRES  45 A  740  ASP GLN ILE GLU ALA ALA ARG GLN PHE SER LYS MET GLY
SEQRES  46 A  740  PHE VAL ASP ASN LYS ARG ILE ALA ILE TRP GLY TRP SER
SEQRES  47 A  740  TYR GLY GLY TYR VAL THR SER MET VAL LEU GLY SER GLY
SEQRES  48 A  740  SER GLY VAL PHE LYS CYS GLY ILE ALA VAL ALA PRO VAL
SEQRES  49 A  740  SER ARG TRP GLU TYR TYR ASP SER VAL TYR THR GLU ARG
SEQRES  50 A  740  TYR MET GLY LEU PRO THR PRO GLU ASP ASN LEU ASP HIS
SEQRES  51 A  740  TYR ARG ASN SER THR VAL MET SER ARG ALA GLU ASN PHE
SEQRES  52 A  740  LYS GLN VAL GLU TYR LEU LEU ILE HIS GLY THR ALA ASP
SEQRES  53 A  740  ASP ASN VAL HIS PHE GLN GLN SER ALA GLN ILE SER LYS
SEQRES  54 A  740  ALA LEU VAL ASP VAL GLY VAL ASP PHE GLN ALA MET TRP
SEQRES  55 A  740  TYR THR ASP GLU ASP HIS GLY ILE ALA SER SER THR ALA
SEQRES  56 A  740  HIS GLN HIS ILE TYR THR HIS MET SER HIS PHE ILE LYS
SEQRES  57 A  740  GLN CYS PHE SER LEU PRO HIS HIS HIS HIS HIS HIS
SEQRES   1 B  740  ASP ASP ALA THR ALA ASP SER ARG LYS THR TYR THR LEU
SEQRES   2 B  740  THR ASP TYR LEU LYS ASN THR TYR ARG LEU LYS LEU TYR
SEQRES   3 B  740  SER LEU ARG TRP ILE SER ASP HIS GLU TYR LEU TYR LYS
SEQRES   4 B  740  GLN GLU ASN ASN ILE LEU VAL PHE ASN ALA GLU TYR GLY
SEQRES   5 B  740  ASN SER SER VAL PHE LEU GLU ASN SER THR PHE ASP GLU
SEQRES   6 B  740  PHE GLY HIS SER ILE ASN ASP TYR SER ILE SER PRO ASP
SEQRES   7 B  740  GLY GLN PHE ILE LEU LEU GLU TYR ASN TYR VAL LYS GLN
SEQRES   8 B  740  TRP ARG HIS SER TYR THR ALA SER TYR ASP ILE TYR ASP
SEQRES   9 B  740  LEU ASN LYS ARG GLN LEU ILE THR GLU GLU ARG ILE PRO
SEQRES  10 B  740  ASN ASN THR GLN TRP VAL THR TRP SER PRO VAL GLY HIS
SEQRES  11 B  740  LYS LEU ALA TYR VAL TRP ASN ASN ASP ILE TYR VAL LYS
SEQRES  12 B  740  ILE GLU PRO ASN LEU PRO SER TYR ARG ILE THR TRP THR
SEQRES  13 B  740  GLY LYS GLU ASP ILE ILE TYR ASN GLY ILE THR ASP TRP
SEQRES  14 B  740  VAL TYR GLU GLU GLU VAL PHE SER ALA TYR SER ALA LEU
SEQRES  15 B  740  TRP TRP SER PRO ASN GLY THR PHE LEU ALA TYR ALA GLN
SEQRES  16 B  740  PHE ASN ASP THR GLU VAL PRO LEU ILE GLU TYR SER PHE
SEQRES  17 B  740  TYR SER ASP GLU SER LEU GLN TYR PRO LYS THR VAL ARG
SEQRES  18 B  740  VAL PRO TYR PRO LYS ALA GLY ALA VAL ASN PRO THR VAL
SEQRES  19 B  740  LYS PHE PHE VAL VAL ASN THR ASP SER LEU SER SER VAL
SEQRES  20 B  740  THR ASN ALA THR SER ILE GLN ILE THR ALA PRO ALA SER
SEQRES  21 B  740  MET LEU ILE GLY ASP HIS TYR LEU CYS ASP VAL THR TRP
SEQRES  22 B  740  ALA THR GLN GLU ARG ILE SER LEU GLN TRP LEU ARG ARG
SEQRES  23 B  740  ILE GLN ASN TYR SER VAL MET ASP ILE CYS ASP TYR ASP
SEQRES  24 B  740  GLU SER SER GLY ARG TRP ASN CYS LEU VAL ALA ARG GLN
SEQRES  25 B  740  HIS ILE GLU MET SER THR THR GLY TRP VAL GLY ARG PHE
SEQRES  26 B  740  ARG PRO SER GLU PRO HIS PHE THR LEU ASP GLY ASN SER
SEQRES  27 B  740  PHE TYR LYS ILE ILE SER ASN GLU GLU GLY TYR ARG HIS
SEQRES  28 B  740  ILE CYS TYR PHE GLN ILE ASP LYS LYS ASP CYS THR PHE
SEQRES  29 B  740  ILE THR LYS GLY THR TRP GLU VAL ILE GLY ILE GLU ALA
SEQRES  30 B  740  LEU THR SER ASP TYR LEU TYR TYR ILE SER ASN GLU TYR
SEQRES  31 B  740  LYS GLY MET PRO GLY GLY ARG ASN LEU TYR LYS ILE GLN
SEQRES  32 B  740  LEU SER ASP TYR THR LYS VAL THR CYS LEU SER CYS GLU
SEQRES  33 B  740  LEU ASN PRO GLU ARG CYS GLN TYR TYR SER VAL SER PHE
SEQRES  34 B  740  SER LYS GLU ALA LYS TYR TYR GLN LEU ARG CYS SER GLY
SEQRES  35 B  740  PRO GLY LEU PRO LEU TYR THR LEU HIS SER SER VAL ASN
SEQRES  36 B  740  ASP LYS GLY LEU ARG VAL LEU GLU ASP ASN SER ALA LEU
SEQRES  37 B  740  ASP LYS MET LEU GLN ASN VAL GLN MET PRO SER LYS LYS
SEQRES  38 B  740  LEU ASP PHE ILE ILE LEU ASN GLU THR LYS PHE TRP TYR
SEQRES  39 B  740  GLN MET ILE LEU PRO PRO HIS PHE ASP LYS SER LYS LYS
SEQRES  40 B  740  TYR PRO LEU LEU LEU ASP VAL TYR ALA GLY PRO CYS SER
SEQRES  41 B  740  GLN LYS ALA ASP THR VAL PHE ARG LEU ASN TRP ALA THR
SEQRES  42 B  740  TYR LEU ALA SER THR GLU ASN ILE ILE VAL ALA SER PHE
SEQRES  43 B  740  ASP GLY ARG GLY SER GLY TYR GLN GLY ASP LYS ILE MET
SEQRES  44 B  740  HIS ALA ILE ASN ARG ARG LEU GLY THR PHE GLU VAL GLU
SEQRES  45 B  740  ASP GLN ILE GLU ALA ALA ARG GLN PHE SER LYS MET GLY
SEQRES  46 B  740  PHE VAL ASP ASN LYS ARG ILE ALA ILE TRP GLY TRP SER
SEQRES  47 B  740  TYR GLY GLY TYR VAL THR SER MET VAL LEU GLY SER GLY
SEQRES  48 B  740  SER GLY VAL PHE LYS CYS GLY ILE ALA VAL ALA PRO VAL
SEQRES  49 B  740  SER ARG TRP GLU TYR TYR ASP SER VAL TYR THR GLU ARG
SEQRES  50 B  740  TYR MET GLY LEU PRO THR PRO GLU ASP ASN LEU ASP HIS
SEQRES  51 B  740  TYR ARG ASN SER THR VAL MET SER ARG ALA GLU ASN PHE
SEQRES  52 B  740  LYS GLN VAL GLU TYR LEU LEU ILE HIS GLY THR ALA ASP
SEQRES  53 B  740  ASP ASN VAL HIS PHE GLN GLN SER ALA GLN ILE SER LYS
SEQRES  54 B  740  ALA LEU VAL ASP VAL GLY VAL ASP PHE GLN ALA MET TRP
SEQRES  55 B  740  TYR THR ASP GLU ASP HIS GLY ILE ALA SER SER THR ALA
SEQRES  56 B  740  HIS GLN HIS ILE TYR THR HIS MET SER HIS PHE ILE LYS
SEQRES  57 B  740  GLN CYS PHE SER LEU PRO HIS HIS HIS HIS HIS HIS
MODRES 3VJK ASN B  219  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN A  219  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN B   85  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN A  229  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN A   85  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN A  281  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN A  321  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN B  321  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN B  229  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN A   92  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN B  150  ASN  GLYCOSYLATION SITE
MODRES 3VJK ASN B  281  ASN  GLYCOSYLATION SITE
HET    M51  A1000      30
HET    NAG  A1085      14
HET    NAG  A1092      14
HET    NAG  A1219      14
HET    NAG  A1220      14
HET    NAG  A1229      14
HET    NAG  A1230      14
HET    NAG  A1281      14
HET    NAG  A1282      14
HET    NAG  A1321      14
HET    M51  B1000      30
HET    NAG  B1085      14
HET    NAG  B1150      14
HET    NAG  B1219      14
HET    NAG  B1220      14
HET    NAG  B1229      14
HET    NAG  B1230      14
HET    NAG  B1281      14
HET    NAG  B1321      14
HETNAM     M51 {(2S,4S)-4-[4-(3-METHYL-1-PHENYL-1H-PYRAZOL-5-YL)
HETNAM   2 M51  PIPERAZIN-1-YL]PYRROLIDIN-2-YL}(1,3-THIAZOLIDIN-3-YL)
HETNAM   3 M51  METHANONE
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   3  M51    2(C22 H30 N6 O S)
FORMUL   4  NAG    17(C8 H15 N O6)
FORMUL  17  HOH   *456(H2 O)
HELIX    1   1 THR A   44  ASN A   51  1                                   8
HELIX    2   2 PHE A   95  GLY A   99  5                                   5
HELIX    3   3 ASP A  200  GLU A  206  1                                   7
HELIX    4   4 ASP A  274  LEU A  276  5                                   3
HELIX    5   5 PRO A  290  ILE A  295  1                                   6
HELIX    6   6 VAL A  341  GLN A  344  5                                   4
HELIX    7   7 GLU A  421  MET A  425  5                                   5
HELIX    8   8 ASN A  497  ASN A  506  1                                  10
HELIX    9   9 ASN A  562  THR A  570  1                                   9
HELIX   10  10 GLY A  587  HIS A  592  1                                   6
HELIX   11  11 ALA A  593  ASN A  595  5                                   3
HELIX   12  12 THR A  600  LYS A  615  1                                  16
HELIX   13  13 SER A  630  GLY A  641  1                                  12
HELIX   14  14 ARG A  658  TYR A  662  5                                   5
HELIX   15  15 ASP A  663  GLY A  672  1                                  10
HELIX   16  16 ASN A  679  ASN A  685  1                                   7
HELIX   17  17 SER A  686  THR A  687  5                                   2
HELIX   18  18 VAL A  688  VAL A  698  5                                  11
HELIX   19  19 PHE A  713  VAL A  726  1                                  14
HELIX   20  20 SER A  744  SER A  764  1                                  21
HELIX   21  21 THR B   44  LYS B   50  1                                   7
HELIX   22  22 GLU B   91  ASP B   96  5                                   6
HELIX   23  23 ASP B  200  GLU B  206  1                                   7
HELIX   24  24 ASP B  274  LEU B  276  5                                   3
HELIX   25  25 PRO B  290  ILE B  295  1                                   6
HELIX   26  26 VAL B  341  GLN B  344  5                                   4
HELIX   27  27 GLU B  421  MET B  425  5                                   5
HELIX   28  28 ASN B  497  GLN B  505  1                                   9
HELIX   29  29 ASN B  562  THR B  570  1                                   9
HELIX   30  30 GLY B  587  HIS B  592  1                                   6
HELIX   31  31 ALA B  593  ASN B  595  5                                   3
HELIX   32  32 THR B  600  MET B  616  1                                  17
HELIX   33  33 SER B  630  GLY B  641  1                                  12
HELIX   34  34 ARG B  658  TYR B  662  5                                   5
HELIX   35  35 ASP B  663  GLY B  672  1                                  10
HELIX   36  36 ASN B  679  SER B  686  1                                   8
HELIX   37  37 VAL B  688  VAL B  698  5                                  11
HELIX   38  38 PHE B  713  GLY B  727  1                                  15
HELIX   39  39 SER B  744  SER B  764  1                                  21
SHEET    1   A 4 LEU A  60  TRP A  62  0
SHEET    2   A 4 GLU A  67  GLN A  72 -1  O  LEU A  69   N  ARG A  61
SHEET    3   A 4 ASN A  75  ASN A  80 -1  O  PHE A  79   N  TYR A  68
SHEET    4   A 4 SER A  86  LEU A  90 -1  O  PHE A  89   N  ILE A  76
SHEET    1   B 4 ILE A 102  ILE A 107  0
SHEET    2   B 4 PHE A 113  LYS A 122 -1  O  LEU A 115   N  SER A 106
SHEET    3   B 4 TYR A 128  ASP A 136 -1  O  SER A 131   N  TYR A 118
SHEET    4   B 4 GLN A 141  LEU A 142 -1  O  GLN A 141   N  ASP A 136
SHEET    1   C 4 TRP A 154  TRP A 157  0
SHEET    2   C 4 LEU A 164  TRP A 168 -1  O  ALA A 165   N  THR A 156
SHEET    3   C 4 ASP A 171  LYS A 175 -1  O  TYR A 173   N  TYR A 166
SHEET    4   C 4 TYR A 183  ARG A 184 -1  O  TYR A 183   N  VAL A 174
SHEET    1   D 3 ILE A 194  ASN A 196  0
SHEET    2   D 3 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3   D 3 LEU A 214  TRP A 216 -1  N  TRP A 215   O  ALA A 224
SHEET    1   E 4 ILE A 194  ASN A 196  0
SHEET    2   E 4 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3   E 4 THR A 265  ASN A 272 -1  O  THR A 265   N  ASN A 229
SHEET    4   E 4 ILE A 285  GLN A 286 -1  O  ILE A 285   N  VAL A 270
SHEET    1   F 2 LEU A 235  PHE A 240  0
SHEET    2   F 2 LYS A 250  PRO A 255 -1  O  VAL A 252   N  TYR A 238
SHEET    1   G 4 HIS A 298  THR A 307  0
SHEET    2   G 4 ARG A 310  ARG A 317 -1  O  ARG A 310   N  ALA A 306
SHEET    3   G 4 TYR A 322  ASP A 331 -1  O  CYS A 328   N  ILE A 311
SHEET    4   G 4 ARG A 336  CYS A 339 -1  O  ARG A 336   N  ASP A 331
SHEET    1   H 4 HIS A 298  THR A 307  0
SHEET    2   H 4 ARG A 310  ARG A 317 -1  O  ARG A 310   N  ALA A 306
SHEET    3   H 4 TYR A 322  ASP A 331 -1  O  CYS A 328   N  ILE A 311
SHEET    4   H 4 HIS A 345  MET A 348 -1  O  HIS A 345   N  MET A 325
SHEET    1   I 4 HIS A 363  PHE A 364  0
SHEET    2   I 4 SER A 370  SER A 376 -1  O  TYR A 372   N  HIS A 363
SHEET    3   I 4 ARG A 382  GLN A 388 -1  O  CYS A 385   N  LYS A 373
SHEET    4   I 4 THR A 395  PHE A 396 -1  O  THR A 395   N  TYR A 386
SHEET    1   J 4 VAL A 404  LEU A 410  0
SHEET    2   J 4 TYR A 414  SER A 419 -1  O  TYR A 416   N  ALA A 409
SHEET    3   J 4 ASN A 430  GLN A 435 -1  O  TYR A 432   N  TYR A 417
SHEET    4   J 4 VAL A 442  CYS A 444 -1  O  THR A 443   N  LYS A 433
SHEET    1   K 4 TYR A 457  PHE A 461  0
SHEET    2   K 4 TYR A 467  CYS A 472 -1  O  GLN A 469   N  SER A 460
SHEET    3   K 4 LEU A 479  SER A 484 -1  O  LEU A 479   N  CYS A 472
SHEET    4   K 4 LYS A 489  GLU A 495 -1  O  LEU A 491   N  LEU A 482
SHEET    1   L 8 SER A 511  LEU A 519  0
SHEET    2   L 8 THR A 522  LEU A 530 -1  O  LEU A 530   N  SER A 511
SHEET    3   L 8 ILE A 574  PHE A 578 -1  O  VAL A 575   N  ILE A 529
SHEET    4   L 8 TYR A 540  VAL A 546  1  N  ASP A 545   O  ALA A 576
SHEET    5   L 8 VAL A 619  TRP A 629  1  O  ALA A 625   N  LEU A 544
SHEET    6   L 8 CYS A 649  VAL A 653  1  O  VAL A 653   N  GLY A 628
SHEET    7   L 8 GLU A 699  GLY A 705  1  O  LEU A 701   N  GLY A 650
SHEET    8   L 8 GLN A 731  TYR A 735  1  O  GLN A 731   N  TYR A 700
SHEET    1   M 2 LYS B  41  THR B  42  0
SHEET    2   M 2 VAL B 507  GLN B 508  1  O  GLN B 508   N  LYS B  41
SHEET    1   N 4 LEU B  60  TRP B  62  0
SHEET    2   N 4 GLU B  67  GLN B  72 -1  O  LEU B  69   N  ARG B  61
SHEET    3   N 4 ASN B  75  ASN B  80 -1  O  PHE B  79   N  TYR B  68
SHEET    4   N 4 SER B  86  LEU B  90 -1  O  PHE B  89   N  ILE B  76
SHEET    1   O 4 ASP B 104  ILE B 107  0
SHEET    2   O 4 PHE B 113  LYS B 122 -1  O  LEU B 115   N  SER B 106
SHEET    3   O 4 TYR B 128  ASP B 136 -1  O  SER B 131   N  TYR B 118
SHEET    4   O 4 GLN B 141  LEU B 142 -1  O  GLN B 141   N  ASP B 136
SHEET    1   P 4 TRP B 154  TRP B 157  0
SHEET    2   P 4 LEU B 164  TRP B 168 -1  O  VAL B 167   N  TRP B 154
SHEET    3   P 4 ASP B 171  LYS B 175 -1  O  LYS B 175   N  LEU B 164
SHEET    4   P 4 TYR B 183  ARG B 184 -1  O  TYR B 183   N  VAL B 174
SHEET    1   Q 3 ILE B 194  ASN B 196  0
SHEET    2   Q 3 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3   Q 3 LEU B 214  TRP B 216 -1  N  TRP B 215   O  ALA B 224
SHEET    1   R 4 ILE B 194  ASN B 196  0
SHEET    2   R 4 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3   R 4 THR B 265  ASN B 272 -1  O  THR B 265   N  ASN B 229
SHEET    4   R 4 ILE B 285  GLN B 286 -1  O  ILE B 285   N  VAL B 270
SHEET    1   S 2 LEU B 235  PHE B 240  0
SHEET    2   S 2 LYS B 250  PRO B 255 -1  O  LYS B 250   N  PHE B 240
SHEET    1   T 4 HIS B 298  TRP B 305  0
SHEET    2   T 4 ARG B 310  ARG B 317 -1  O  LEU B 316   N  TYR B 299
SHEET    3   T 4 TYR B 322  ASP B 331 -1  O  CYS B 328   N  ILE B 311
SHEET    4   T 4 ARG B 336  CYS B 339 -1  O  ARG B 336   N  ASP B 331
SHEET    1   U 4 HIS B 298  TRP B 305  0
SHEET    2   U 4 ARG B 310  ARG B 317 -1  O  LEU B 316   N  TYR B 299
SHEET    3   U 4 TYR B 322  ASP B 331 -1  O  CYS B 328   N  ILE B 311
SHEET    4   U 4 HIS B 345  MET B 348 -1  O  GLU B 347   N  SER B 323
SHEET    1   V 4 HIS B 363  PHE B 364  0
SHEET    2   V 4 SER B 370  SER B 376 -1  O  TYR B 372   N  HIS B 363
SHEET    3   V 4 ARG B 382  GLN B 388 -1  O  PHE B 387   N  PHE B 371
SHEET    4   V 4 THR B 395  PHE B 396 -1  O  THR B 395   N  TYR B 386
SHEET    1   W 4 VAL B 404  LEU B 410  0
SHEET    2   W 4 TYR B 414  SER B 419 -1  O  TYR B 416   N  ALA B 409
SHEET    3   W 4 ASN B 430  GLN B 435 -1  O  TYR B 432   N  TYR B 417
SHEET    4   W 4 ASP B 438  CYS B 444 -1  O  THR B 443   N  LYS B 433
SHEET    1   X 4 TYR B 457  PHE B 461  0
SHEET    2   X 4 TYR B 467  CYS B 472 -1  O  ARG B 471   N  SER B 458
SHEET    3   X 4 LEU B 479  SER B 484 -1  O  LEU B 479   N  CYS B 472
SHEET    4   X 4 LYS B 489  GLU B 495 -1  O  GLU B 495   N  TYR B 480
SHEET    1   Y 8 SER B 511  LEU B 519  0
SHEET    2   Y 8 THR B 522  LEU B 530 -1  O  LEU B 530   N  SER B 511
SHEET    3   Y 8 ILE B 574  PHE B 578 -1  O  VAL B 575   N  ILE B 529
SHEET    4   Y 8 TYR B 540  VAL B 546  1  N  LEU B 543   O  ILE B 574
SHEET    5   Y 8 VAL B 619  TRP B 629  1  O  ASP B 620   N  TYR B 540
SHEET    6   Y 8 CYS B 649  VAL B 653  1  O  VAL B 653   N  GLY B 628
SHEET    7   Y 8 GLU B 699  GLY B 705  1  O  LEU B 701   N  ALA B 652
SHEET    8   Y 8 GLN B 731  TYR B 735  1  O  GLN B 731   N  TYR B 700
SSBOND   1 CYS A  328    CYS A  339                          1555   1555  2.06
SSBOND   2 CYS A  385    CYS A  394                          1555   1555  2.06
SSBOND   3 CYS A  444    CYS A  447                          1555   1555  2.03
SSBOND   4 CYS A  454    CYS A  472                          1555   1555  2.07
SSBOND   5 CYS A  649    CYS A  762                          1555   1555  2.05
SSBOND   6 CYS B  328    CYS B  339                          1555   1555  2.05
SSBOND   7 CYS B  385    CYS B  394                          1555   1555  2.06
SSBOND   8 CYS B  444    CYS B  447                          1555   1555  2.03
SSBOND   9 CYS B  454    CYS B  472                          1555   1555  2.04
SSBOND  10 CYS B  649    CYS B  762                          1555   1555  2.04
LINK         ND2 ASN B 219                 C1  NAG B1219     1555   1555  1.43
LINK         ND2 ASN A 219                 C1  NAG A1219     1555   1555  1.43
LINK         ND2 ASN B  85                 C1  NAG B1085     1555   1555  1.44
LINK         ND2 ASN A 229                 C1  NAG A1229     1555   1555  1.44
LINK         O4  NAG A1281                 C1  NAG A1282     1555   1555  1.44
LINK         O4  NAG B1229                 C1  NAG B1230     1555   1555  1.44
LINK         ND2 ASN A  85                 C1  NAG A1085     1555   1555  1.44
LINK         ND2 ASN A 281                 C1  NAG A1281     1555   1555  1.44
LINK         O4  NAG A1219                 C1  NAG A1220     1555   1555  1.45
LINK         ND2 ASN A 321                 C1  NAG A1321     1555   1555  1.45
LINK         ND2 ASN B 321                 C1  NAG B1321     1555   1555  1.45
LINK         ND2 ASN B 229                 C1  NAG B1229     1555   1555  1.45
LINK         O4  NAG B1219                 C1  NAG B1220     1555   1555  1.45
LINK         O4  NAG A1229                 C1  NAG A1230     1555   1555  1.45
LINK         ND2 ASN A  92                 C1  NAG A1092     1555   1555  1.46
LINK         ND2 ASN B 150                 C1  NAG B1150     1555   1555  1.46
LINK         ND2 ASN B 281                 C1  NAG B1281     1555   1555  1.46
CISPEP   1 GLY A  474    PRO A  475          0         7.31
CISPEP   2 GLY B  474    PRO B  475          0         9.48
SITE     1 AC1 14 ARG A 125  GLU A 205  GLU A 206  VAL A 207
SITE     2 AC1 14 SER A 209  PHE A 357  ARG A 358  SER A 630
SITE     3 AC1 14 TYR A 631  VAL A 656  TYR A 662  TYR A 666
SITE     4 AC1 14 ASN A 710  HOH A2063
SITE     1 AC2  3 ASN A  85  SER A  86  SER A  87
SITE     1 AC3  3 GLU A  73  ASN A  75  ASN A  92
SITE     1 AC4  5 ASN A 219  THR A 221  GLN A 308  GLU A 309
SITE     2 AC4  5 NAG A1220
SITE     1 AC5  3 ASN A 272  GLU A 332  NAG A1219
SITE     1 AC6  5 ILE A 194  ASN A 229  THR A 231  GLU A 232
SITE     2 AC6  5 NAG A1230
SITE     1 AC7  1 NAG A1229
SITE     1 AC8  4 TRP A 187  ASN A 281  NAG A1282  HOH A2090
SITE     1 AC9  4 THR A 188  NAG A1281  HOH A2084  HOH A2203
SITE     1 BC1  4 ASN A 321  MET A 348  SER A 349  ARG A 596
SITE     1 BC2 14 ARG B 125  GLU B 205  GLU B 206  VAL B 207
SITE     2 BC2 14 SER B 209  PHE B 357  ARG B 358  SER B 630
SITE     3 BC2 14 TYR B 631  TYR B 662  TYR B 666  ASN B 710
SITE     4 BC2 14 VAL B 711  HOH B2265
SITE     1 BC3  5 GLU B  67  VAL B  78  ASN B  85  SER B  86
SITE     2 BC3  5 SER B  87
SITE     1 BC4  2 ILE B 148  ASN B 150
SITE     1 BC5  5 ASN B 219  THR B 221  GLN B 308  GLU B 309
SITE     2 BC5  5 NAG B1220
SITE     1 BC6  2 TYR B 330  NAG B1219
SITE     1 BC7  5 ILE B 194  ASN B 229  THR B 231  GLU B 232
SITE     2 BC7  5 NAG B1230
SITE     1 BC8  1 NAG B1229
SITE     1 BC9  1 ASN B 281
SITE     1 CC1  4 ILE B 319  ASN B 321  SER B 349  ARG B 596
CRYST1  117.955  126.413  138.011  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008478  0.000000  0.000000        0.00000
SCALE2      0.000000  0.007911  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007246        0.00000
TER    5972      PRO A 766
TER   11944      PRO B 766
MASTER      406    0   19   39  100    0   30    612696    2  330  114
END