longtext: 3WA6-pdb

content
HEADER    HYDROLASE                               27-APR-13   3WA6
TITLE     CRYSTAL STRUCTURE OF TANNASE FROM LACTOBACILLUS PLANTARUM IN THE
TITLE    2 ORTHORHOMBIC CRYSTAL
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: TANNASE;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.1.1.20;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: LACTOBACILLUS PLANTARUM;
SOURCE   3 ORGANISM_TAXID: 1590;
SOURCE   4 STRAIN: SN35N;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PET-21A(+)
KEYWDS    ALPHA/BETA-HYDROLASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Y.MATOBA,N.TANAKA,M.SUGIYAMA
REVDAT   1   24-JUL-13 3WA6    0
JRNL        AUTH   Y.MATOBA,N.TANAKA,M.NODA,F.HIGASHIKAWA,T.KUMAGAI,M.SUGIYAMA
JRNL        TITL   CRYSTALLOGRAPHIC AND MUTATIONAL ANALYSES OF TANNASE FROM
JRNL        TITL 2 LACTOBACILLUS PLANTARUM
JRNL        REF    PROTEINS                                   2013
JRNL        REFN                   ESSN 1097-0134
JRNL        PMID   23836494
JRNL        DOI    10.1002/PROT.24355
REMARK   2
REMARK   2 RESOLUTION.    1.80 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.2
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.80
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 23.63
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 2.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 3617886.270
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 96.8
REMARK   3   NUMBER OF REFLECTIONS             : 39790
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.178
REMARK   3   FREE R VALUE                     : 0.207
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 2023
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.005
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.80
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.91
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 93.30
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 6007
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2450
REMARK   3   BIN FREE R VALUE                    : 0.2800
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 4.80
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 303
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.016
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 3560
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 10
REMARK   3   SOLVENT ATOMS            : 331
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 18.70
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 20.40
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -1.35000
REMARK   3    B22 (A**2) : 3.92000
REMARK   3    B33 (A**2) : -2.57000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.18
REMARK   3   ESD FROM SIGMAA              (A) : 0.16
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.22
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.21
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.005
REMARK   3   BOND ANGLES            (DEGREES) : 1.30
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 23.10
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.79
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.120 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 1.660 ; 2.000
REMARK   3   SIDE-CHAIN BOND              (A**2) : 1.890 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 2.710 ; 2.500
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : FLAT MODEL
REMARK   3   KSOL        : 0.40
REMARK   3   BSOL        : 62.48
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM
REMARK   3  PARAMETER FILE  2  : WATER_REP.PARAM
REMARK   3  PARAMETER FILE  3  : ION.PARAM
REMARK   3  PARAMETER FILE  4  : NULL
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP
REMARK   3  TOPOLOGY FILE  2   : WATER.TOP
REMARK   3  TOPOLOGY FILE  3   : ION.TOP
REMARK   3  TOPOLOGY FILE  4   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: BULK SOLVENT MODEL USED
REMARK   4
REMARK   4 3WA6 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 09-MAY-13.
REMARK 100 THE RCSB ID CODE IS RCSB096079.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 22-NOV-11
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SPRING-8
REMARK 200  BEAMLINE                       : BL26B2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0000
REMARK 200  MONOCHROMATOR                  : SI 111 CHANNEL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : RAYONIX MX225HE
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 41361
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.800
REMARK 200  RESOLUTION RANGE LOW       (A) : 100.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.9
REMARK 200  DATA REDUNDANCY                : 5.500
REMARK 200  R MERGE                    (I) : 0.09900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 17.2000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.80
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.86
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.2
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.90
REMARK 200  R MERGE FOR SHELL          (I) : 0.43500
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.800
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SAD
REMARK 200 SOFTWARE USED: SHARP
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 41.66
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.11
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: PEG8000, SODIUM ACETATE, AMMONIUM
REMARK 280  SULFATE, PH 5, VAPOR DIFFUSION, SITTING DROP, TEMPERATURE 297K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 2 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -X,Y,-Z+1/2
REMARK 290       4555   X,-Y,-Z
REMARK 290       5555   X+1/2,Y+1/2,Z
REMARK 290       6555   -X+1/2,-Y+1/2,Z+1/2
REMARK 290       7555   -X+1/2,Y+1/2,-Z+1/2
REMARK 290       8555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      100.00500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      100.00500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000       27.93500
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       39.12000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   6 -1.000000  0.000000  0.000000       27.93500
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       39.12000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000      100.00500
REMARK 290   SMTRY1   7 -1.000000  0.000000  0.000000       27.93500
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       39.12000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000      100.00500
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000       27.93500
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000       39.12000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1440 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 35740 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -10.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   2  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000 -1.000000      100.00500
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A 863  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     SER A     2
REMARK 465     ASN A     3
REMARK 465     LEU A   470
REMARK 465     GLU A   471
REMARK 465     HIS A   472
REMARK 465     HIS A   473
REMARK 465     HIS A   474
REMARK 465     HIS A   475
REMARK 465     HIS A   476
REMARK 465     HIS A   477
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A 163     -121.98     60.85
REMARK 500    CYS A 204       58.79     30.75
REMARK 500    THR A 236     -141.35    -93.27
REMARK 500    THR A 340      151.84     84.00
REMARK 500    ALA A 344     -158.66    -81.15
REMARK 500    HIS A 451       99.63    -69.27
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A 502
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 A SEQUENCE DATABASE REFERENCE FOR THIS PROTEIN DOES NOT CURRENTLY
REMARK 999 EXIST.
DBREF  3WA6 A    1   477  PDB    3WA6     3WA6             1    477
SEQRES   1 A  477  MET SER ASN ARG LEU ILE PHE ASP ALA ASP TRP LEU VAL
SEQRES   2 A  477  PRO GLU GLN VAL GLN VAL ALA GLY GLN ALA ILE GLN TYR
SEQRES   3 A  477  TYR ALA ALA ARG ASN ILE GLN TYR VAL GLN HIS PRO VAL
SEQRES   4 A  477  ALA ALA ILE GLN VAL LEU ASN VAL PHE VAL PRO ALA ALA
SEQRES   5 A  477  TYR LEU HIS GLY SER SER VAL ASN GLY TYR GLN ARG ALA
SEQRES   6 A  477  THR ALA PRO ILE LEU MET PRO ASN THR VAL GLY GLY TYR
SEQRES   7 A  477  LEU PRO GLY PRO ALA ASP ASP PRO GLN ARG VAL THR TRP
SEQRES   8 A  477  PRO THR ASN ALA GLY THR ILE GLN GLN ALA LEU LYS ARG
SEQRES   9 A  477  GLY TYR VAL VAL VAL ALA ALA GLY ILE ARG GLY ARG THR
SEQRES  10 A  477  THR VAL ASP LYS SER GLY GLN ARG VAL GLY GLN ALA PRO
SEQRES  11 A  477  ALA PHE ILE VAL ASP MET LYS ALA ALA ILE ARG TYR VAL
SEQRES  12 A  477  LYS TYR ASN GLN GLY ARG LEU PRO GLY ASP THR ASN ARG
SEQRES  13 A  477  ILE ILE THR ASN GLY THR SER ALA GLY GLY ALA THR SER
SEQRES  14 A  477  ALA LEU ALA GLY ALA SER GLY ASN SER ALA TYR PHE GLU
SEQRES  15 A  477  PRO ALA LEU THR ALA LEU GLY ALA ALA PRO ALA THR ASP
SEQRES  16 A  477  ASP ILE PHE ALA VAL SER ALA TYR CYS PRO ILE HIS ASN
SEQRES  17 A  477  LEU GLU HIS ALA ASP MET ALA TYR GLU TRP GLN PHE ASN
SEQRES  18 A  477  GLY ILE ASN ASP TRP HIS ARG TYR GLN PRO VAL ALA GLY
SEQRES  19 A  477  THR THR LYS ASN GLY ARG PRO LYS PHE GLU PRO VAL SER
SEQRES  20 A  477  GLY GLN LEU THR VAL GLU GLU GLN ALA LEU SER LEU ALA
SEQRES  21 A  477  LEU LYS ALA GLN PHE SER THR TYR LEU ASN GLN LEU GLN
SEQRES  22 A  477  LEU THR ALA SER ASP GLY THR HIS LEU THR LEU ASN GLU
SEQRES  23 A  477  ALA GLY MET GLY SER PHE ARG ASP VAL VAL ARG GLN LEU
SEQRES  24 A  477  LEU ILE SER SER ALA GLN THR ALA PHE ASP GLN GLY THR
SEQRES  25 A  477  ASP ILE HIS LYS TYR ALA GLY PHE VAL VAL THR GLY ASN
SEQRES  26 A  477  GLN VAL THR ASP LEU ASP LEU SER ALA TYR LEU LYS SER
SEQRES  27 A  477  LEU THR ARG MET LYS ALA VAL PRO ALA PHE ASP GLN LEU
SEQRES  28 A  477  ASP LEU THR SER PRO GLU ASN ASN LEU PHE GLY ASP ALA
SEQRES  29 A  477  THR ALA LYS ALA LYS HIS PHE THR ALA LEU ALA GLN THR
SEQRES  30 A  477  ARG SER THR VAL THR ALA GLN LEU ALA ASP ALA GLU LEU
SEQRES  31 A  477  ILE GLN ALA ILE ASN PRO LEU SER TYR LEU THR THR THR
SEQRES  32 A  477  SER SER GLN VAL ALA LYS HIS TRP ARG ILE ARG HIS GLY
SEQRES  33 A  477  ALA ALA ASP ARG ASP THR SER PHE ALA ILE PRO ILE ILE
SEQRES  34 A  477  LEU ALA ILE MET LEU GLU ASN HIS GLY TYR GLY ILE ASP
SEQRES  35 A  477  PHE ALA LEU PRO TRP ASP ILE PRO HIS SER GLY ASP TYR
SEQRES  36 A  477  ASP LEU GLY ASP LEU PHE SER TRP ILE ASP GLY LEU CYS
SEQRES  37 A  477  GLN LEU GLU HIS HIS HIS HIS HIS HIS
HET    SO4  A 501       5
HET    SO4  A 502       5
HETNAM     SO4 SULFATE ION
FORMUL   2  SO4    2(O4 S 2-)
FORMUL   4  HOH   *331(H2 O)
HELIX    1   1 ASP A    8  LEU A   12  5                                   5
HELIX    2   2 ALA A   52  HIS A   55  5                                   4
HELIX    3   3 ASN A   94  GLY A  105  1                                  12
HELIX    4   4 PRO A  130  ASN A  146  1                                  17
HELIX    5   5 SER A  163  SER A  175  1                                  13
HELIX    6   6 SER A  178  TYR A  180  5                                   3
HELIX    7   7 PHE A  181  GLY A  189  1                                   9
HELIX    8   8 ASN A  208  ASN A  221  1                                  14
HELIX    9   9 LYS A  237  GLY A  239  5                                   3
HELIX   10  10 THR A  251  LEU A  272  1                                  22
HELIX   11  11 GLY A  290  GLN A  310  1                                  21
HELIX   12  12 ASP A  331  THR A  340  1                                  10
HELIX   13  13 SER A  355  PHE A  361  1                                   7
HELIX   14  14 THR A  372  ARG A  378  1                                   7
HELIX   15  15 ASP A  387  ILE A  394  1                                   8
HELIX   16  16 ASN A  395  LEU A  400  5                                   6
HELIX   17  17 PHE A  424  HIS A  437  1                                  14
HELIX   18  18 ASP A  456  GLN A  469  1                                  14
SHEET    1   A 9 VAL A  13  VAL A  19  0
SHEET    2   A 9 GLN A  22  GLN A  33 -1  O  GLN A  22   N  VAL A  19
SHEET    3   A 9 VAL A  44  PRO A  50 -1  O  VAL A  47   N  ALA A  29
SHEET    4   A 9 VAL A 107  ALA A 111 -1  O  VAL A 108   N  PHE A  48
SHEET    5   A 9 ILE A  69  PRO A  72  1  N  LEU A  70   O  VAL A 107
SHEET    6   A 9 ILE A 157  ASN A 160  1  O  ILE A 158   N  MET A  71
SHEET    7   A 9 ALA A 199  TYR A 203  1  O  SER A 201   N  THR A 159
SHEET    8   A 9 HIS A 410  ARG A 414  1  O  HIS A 410   N  VAL A 200
SHEET    9   A 9 GLY A 440  ALA A 444  1  O  ASP A 442   N  ILE A 413
SHEET    1   B 2 ASP A 225  THR A 235  0
SHEET    2   B 2 PRO A 241  GLN A 249 -1  O  VAL A 246   N  ARG A 228
SHEET    1   C 2 PHE A 320  THR A 323  0
SHEET    2   C 2 GLN A 326  LEU A 330 -1  O  GLN A 326   N  THR A 323
CISPEP   1 TRP A   91    PRO A   92          0        -0.04
CISPEP   2 ALA A  129    PRO A  130          0         0.32
CISPEP   3 VAL A  345    PRO A  346          0        -0.43
SITE     1 AC1  9 MET A 289  ARG A 293  ARG A 297  LEU A 330
SITE     2 AC1  9 ASP A 331  LEU A 332  SER A 333  HOH A 781
SITE     3 AC1  9 HOH A 920
SITE     1 AC2  7 GLY A  76  GLY A  77  SER A 163  HIS A 451
SITE     2 AC2  7 HOH A 701  HOH A 766  HOH A 805
CRYST1   55.870   78.240  200.010  90.00  90.00  90.00 C 2 2 21      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017899  0.000000  0.000000        0.00000
SCALE2      0.000000  0.012781  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005000        0.00000
TER    3572      GLN A 469
MASTER      306    0    2   18   13    0    5    6 3901    1   10   37
END