longtext: 3WIB-pdb

content
HEADER    HYDROLASE                               09-SEP-13   3WIB
TITLE     CRYSTAL STRUCTURE OF Y109W MUTANT HALOALKANE DEHALOGENASE DATA FROM
TITLE    2 AGROBACTERIUM TUMEFACIENS C58
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: HALOALKANE DEHALOGENASE;
COMPND   3 CHAIN: A, B;
COMPND   4 EC: 3.8.1.5;
COMPND   5 ENGINEERED: YES;
COMPND   6 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: AGROBACTERIUM TUMEFACIENS;
SOURCE   3 ORGANISM_TAXID: 176299;
SOURCE   4 STRAIN: C58 / ATCC 33970;
SOURCE   5 GENE: DHAA, DHA, ATU6064, AGR_PTI_130;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: ROSETTA(DE3)
KEYWDS    HALOALKANE DEHALOGENASE, HYDROLASE FOLD FAMILY, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    L.J.GUAN,H.YABUKI,M.OKAI,J.OHTSUKA,M.TANOKURA
REVDAT   1   23-JUL-14 3WIB    0
JRNL        AUTH   L.GUAN,H.YABUKI,M.OKAI,J.OHTSUKA,M.TANOKURA
JRNL        TITL   CRYSTAL STRUCTURE OF THE NOVEL HALOALKANE DEHALOGENASE DATA
JRNL        TITL 2 FROM AGROBACTERIUM TUMEFACIENS C58 REVEALS A SPECIAL
JRNL        TITL 3 HALIDE-STABILIZING PAIR AND ENANTIOSELECTIVITY MECHANISM.
JRNL        REF    APPL.MICROBIOL.BIOTECHNOL.                 2014
JRNL        REFN                   ESSN 1432-0614
JRNL        PMID   24770384
JRNL        DOI    10.1007/S00253-014-5751-2
REMARK   2
REMARK   2 RESOLUTION.    1.95 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.7.0032
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.95
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.80
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.8
REMARK   3   NUMBER OF REFLECTIONS             : 47662
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.196
REMARK   3   R VALUE            (WORKING SET) : 0.194
REMARK   3   FREE R VALUE                     : 0.235
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 2552
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.95
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.00
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 3449
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.00
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2420
REMARK   3   BIN FREE R VALUE SET COUNT          : 204
REMARK   3   BIN FREE R VALUE                    : 0.2950
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4655
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 39
REMARK   3   SOLVENT ATOMS            : 179
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 25.88
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.20000
REMARK   3    B22 (A**2) : 0.20000
REMARK   3    B33 (A**2) : -0.40000
REMARK   3    B12 (A**2) : -0.00000
REMARK   3    B13 (A**2) : -0.00000
REMARK   3    B23 (A**2) : -0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.160
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.150
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.101
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 3.511
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.950
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.926
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4848 ; 0.019 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  4569 ; 0.001 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  6618 ; 1.786 ; 1.951
REMARK   3   BOND ANGLES OTHERS          (DEGREES): 10467 ; 0.876 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   580 ; 6.305 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   241 ;34.014 ;23.112
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   731 ;12.960 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    41 ;16.739 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   716 ; 0.111 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  5516 ; 0.010 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  1213 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2323 ; 2.386 ; 2.471
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  2322 ; 2.385 ; 2.470
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  2902 ; 3.149 ; 3.690
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  2903 ; 3.149 ; 3.691
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  2525 ; 2.824 ; 2.690
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  2526 ; 2.823 ; 2.690
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  3717 ; 4.211 ; 3.952
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  5588 ; 5.337 ;19.935
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  5529 ; 5.321 ;19.899
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS
REMARK   4
REMARK   4 3WIB COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 26-SEP-13.
REMARK 100 THE RCSB ID CODE IS RCSB096370.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 06-FEB-11
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : PHOTON FACTORY
REMARK 200  BEAMLINE                       : AR-NW12A
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0000
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 210R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL
REMARK 200  DATA SCALING SOFTWARE          : NULL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 50218
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.950
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : NULL
REMARK 200  DATA REDUNDANCY                : NULL
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 49.63
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.44
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M N-CYCLOHEXYL-2-
REMARK 280  AMINOETHANESULFONIC ACID (CHES), 1.0M POTASSIUM SODIUM TARTRATE,
REMARK 280  0.2M LITHIUM SULFATE, 0.1M MANGANESE CHLORIDE, 1MM DTT, PH 8.5,
REMARK 280  EVAPORATION, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 4 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   -Y,X,Z
REMARK 290       4555   Y,-X,Z
REMARK 290       5555   -X,Y,-Z
REMARK 290       6555   X,-Y,-Z
REMARK 290       7555   Y,X,-Z
REMARK 290       8555   -Y,-X,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A    -5
REMARK 465     LYS A    -4
REMARK 465     GLU A    -3
REMARK 465     HIS A    -2
REMARK 465     ARG A    -1
REMARK 465     HIS A     0
REMARK 465     MET A     1
REMARK 465     THR A     2
REMARK 465     GLU A     3
REMARK 465     LYS A     4
REMARK 465     SER A     5
REMARK 465     ILE A   295
REMARK 465     ALA A   296
REMARK 465     GLY A   297
REMARK 465     ASN A   298
REMARK 465     MET B    -5
REMARK 465     LYS B    -4
REMARK 465     GLU B    -3
REMARK 465     HIS B    -2
REMARK 465     ARG B    -1
REMARK 465     HIS B     0
REMARK 465     MET B     1
REMARK 465     THR B     2
REMARK 465     GLU B     3
REMARK 465     LYS B     4
REMARK 465     SER B     5
REMARK 465     GLY B   297
REMARK 465     ASN B   298
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OD1  ASP A   235     NH2  ARG A   262              1.96
REMARK 500   OD2  ASP A   139     O    HOH A   567              2.18
REMARK 500   O    HOH B   476     O    HOH B   484              2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ASP A  18   CB  -  CG  -  OD1 ANGL. DEV. =   7.4 DEGREES
REMARK 500    ASP A  18   CB  -  CG  -  OD2 ANGL. DEV. =  -5.6 DEGREES
REMARK 500    ARG A 136   NE  -  CZ  -  NH2 ANGL. DEV. =  -4.0 DEGREES
REMARK 500    ARG A 150   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.0 DEGREES
REMARK 500    ASP B  67   CB  -  CG  -  OD1 ANGL. DEV. =   6.4 DEGREES
REMARK 500    ARG B 155   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.1 DEGREES
REMARK 500    ASP B 214   CB  -  CG  -  OD1 ANGL. DEV. =   5.7 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PRO A  44       57.09   -113.05
REMARK 500    THR A  45     -155.47   -110.18
REMARK 500    ASP A  98       72.82     41.46
REMARK 500    ASP A 108     -135.86     53.51
REMARK 500    GLU A 132       57.85     38.74
REMARK 500    LEU A 135      -41.69   -137.86
REMARK 500    GLU A 173      -59.50   -122.73
REMARK 500    PHE A 178       39.56    -90.07
REMARK 500    PHE A 247      -71.80   -152.13
REMARK 500    ILE A 273      -77.54   -103.65
REMARK 500    HIS A 274      -54.01   -134.46
REMARK 500    PHE B  22       41.35     35.17
REMARK 500    PRO B  44       56.14   -110.28
REMARK 500    THR B  45     -154.57   -113.01
REMARK 500    ASP B 108     -136.51     54.72
REMARK 500    GLU B 132       57.34     36.03
REMARK 500    LEU B 135      -40.74   -144.73
REMARK 500    GLU B 173      -65.18   -127.73
REMARK 500    PHE B 247      -74.64   -152.90
REMARK 500    ILE B 273      -77.08   -109.63
REMARK 500    HIS B 274      -54.23   -133.12
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NHE A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NHE A 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NHE B 301
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3WI7   RELATED DB: PDB
DBREF  3WIB A   -5   298  UNP    Q8U671   DHAA_AGRT5       1    304
DBREF  3WIB B   -5   298  UNP    Q8U671   DHAA_AGRT5       1    304
SEQADV 3WIB TRP A  109  UNP  Q8U671    TYR   115 ENGINEERED MUTATION
SEQADV 3WIB TRP B  109  UNP  Q8U671    TYR   115 ENGINEERED MUTATION
SEQRES   1 A  304  MET LYS GLU HIS ARG HIS MET THR GLU LYS SER PRO HIS
SEQRES   2 A  304  SER ALA PHE GLY ASP GLY ALA LYS ALA TYR ASP VAL PRO
SEQRES   3 A  304  ALA PHE GLY LEU GLN ILE HIS THR VAL GLU HIS GLY SER
SEQRES   4 A  304  GLY ALA PRO ILE VAL PHE LEU HIS GLY ASN PRO THR SER
SEQRES   5 A  304  SER TYR LEU TRP ARG HIS ILE PHE ARG ARG LEU HIS GLY
SEQRES   6 A  304  HIS GLY ARG LEU LEU ALA VAL ASP LEU ILE GLY TYR GLY
SEQRES   7 A  304  GLN SER SER LYS PRO ASP ILE GLU TYR THR LEU GLU ASN
SEQRES   8 A  304  GLN GLN ARG TYR VAL ASP ALA TRP PHE ASP ALA LEU ASP
SEQRES   9 A  304  LEU ARG ASN VAL THR LEU VAL LEU GLN ASP TRP GLY ALA
SEQRES  10 A  304  ALA PHE GLY LEU ASN TRP ALA SER ARG ASN PRO ASP ARG
SEQRES  11 A  304  VAL ARG ALA VAL ALA PHE PHE GLU PRO VAL LEU ARG ASN
SEQRES  12 A  304  ILE ASP SER VAL ASP LEU SER PRO GLU PHE VAL THR ARG
SEQRES  13 A  304  ARG ALA LYS LEU ARG GLN PRO GLY GLU GLY GLU ILE PHE
SEQRES  14 A  304  VAL GLN GLN GLU ASN ARG PHE LEU THR GLU LEU PHE PRO
SEQRES  15 A  304  TRP PHE PHE LEU THR PRO LEU ALA PRO GLU ASP LEU ARG
SEQRES  16 A  304  GLN TYR GLN THR PRO PHE PRO THR PRO HIS SER ARG LYS
SEQRES  17 A  304  ALA ILE LEU ALA GLY PRO ARG ASN LEU PRO VAL ASP GLY
SEQRES  18 A  304  GLU PRO ALA SER THR VAL ALA PHE LEU GLU GLN ALA VAL
SEQRES  19 A  304  ASN TRP LEU ASN THR SER ASP THR PRO LYS LEU LEU LEU
SEQRES  20 A  304  THR PHE LYS PRO GLY PHE LEU LEU THR ASP ALA ILE LEU
SEQRES  21 A  304  LYS TRP SER GLN VAL THR ILE ARG ASN LEU GLU ILE GLU
SEQRES  22 A  304  ALA ALA GLY ALA GLY ILE HIS PHE VAL GLN GLU GLU GLN
SEQRES  23 A  304  PRO GLU THR ILE ALA ARG LEU LEU ASP ALA TRP LEU THR
SEQRES  24 A  304  ARG ILE ALA GLY ASN
SEQRES   1 B  304  MET LYS GLU HIS ARG HIS MET THR GLU LYS SER PRO HIS
SEQRES   2 B  304  SER ALA PHE GLY ASP GLY ALA LYS ALA TYR ASP VAL PRO
SEQRES   3 B  304  ALA PHE GLY LEU GLN ILE HIS THR VAL GLU HIS GLY SER
SEQRES   4 B  304  GLY ALA PRO ILE VAL PHE LEU HIS GLY ASN PRO THR SER
SEQRES   5 B  304  SER TYR LEU TRP ARG HIS ILE PHE ARG ARG LEU HIS GLY
SEQRES   6 B  304  HIS GLY ARG LEU LEU ALA VAL ASP LEU ILE GLY TYR GLY
SEQRES   7 B  304  GLN SER SER LYS PRO ASP ILE GLU TYR THR LEU GLU ASN
SEQRES   8 B  304  GLN GLN ARG TYR VAL ASP ALA TRP PHE ASP ALA LEU ASP
SEQRES   9 B  304  LEU ARG ASN VAL THR LEU VAL LEU GLN ASP TRP GLY ALA
SEQRES  10 B  304  ALA PHE GLY LEU ASN TRP ALA SER ARG ASN PRO ASP ARG
SEQRES  11 B  304  VAL ARG ALA VAL ALA PHE PHE GLU PRO VAL LEU ARG ASN
SEQRES  12 B  304  ILE ASP SER VAL ASP LEU SER PRO GLU PHE VAL THR ARG
SEQRES  13 B  304  ARG ALA LYS LEU ARG GLN PRO GLY GLU GLY GLU ILE PHE
SEQRES  14 B  304  VAL GLN GLN GLU ASN ARG PHE LEU THR GLU LEU PHE PRO
SEQRES  15 B  304  TRP PHE PHE LEU THR PRO LEU ALA PRO GLU ASP LEU ARG
SEQRES  16 B  304  GLN TYR GLN THR PRO PHE PRO THR PRO HIS SER ARG LYS
SEQRES  17 B  304  ALA ILE LEU ALA GLY PRO ARG ASN LEU PRO VAL ASP GLY
SEQRES  18 B  304  GLU PRO ALA SER THR VAL ALA PHE LEU GLU GLN ALA VAL
SEQRES  19 B  304  ASN TRP LEU ASN THR SER ASP THR PRO LYS LEU LEU LEU
SEQRES  20 B  304  THR PHE LYS PRO GLY PHE LEU LEU THR ASP ALA ILE LEU
SEQRES  21 B  304  LYS TRP SER GLN VAL THR ILE ARG ASN LEU GLU ILE GLU
SEQRES  22 B  304  ALA ALA GLY ALA GLY ILE HIS PHE VAL GLN GLU GLU GLN
SEQRES  23 B  304  PRO GLU THR ILE ALA ARG LEU LEU ASP ALA TRP LEU THR
SEQRES  24 B  304  ARG ILE ALA GLY ASN
HET    NHE  A 401      13
HET    NHE  A 402      13
HET    NHE  B 301      13
HETNAM     NHE 2-[N-CYCLOHEXYLAMINO]ETHANE SULFONIC ACID
HETSYN     NHE N-CYCLOHEXYLTAURINE; CHES
FORMUL   3  NHE    3(C8 H17 N O3 S)
FORMUL   6  HOH   *179(H2 O)
HELIX    1   1 HIS A    7  GLY A   11  5                                   5
HELIX    2   2 SER A   46  LEU A   49  5                                   4
HELIX    3   3 TRP A   50  LEU A   57  1                                   8
HELIX    4   4 THR A   82  ASP A   98  1                                  17
HELIX    5   5 TRP A  109  ASN A  121  1                                  13
HELIX    6   6 ASP A  139  LEU A  143  5                                   5
HELIX    7   7 SER A  144  ARG A  155  1                                  12
HELIX    8   8 GLY A  158  GLN A  165  1                                   8
HELIX    9   9 ASN A  168  GLU A  173  1                                   6
HELIX   10  10 GLU A  173  PHE A  178  1                                   6
HELIX   11  11 ALA A  184  THR A  193  1                                  10
HELIX   12  12 PRO A  198  SER A  200  5                                   3
HELIX   13  13 ARG A  201  LEU A  211  1                                  11
HELIX   14  14 PRO A  217  GLN A  226  1                                  10
HELIX   15  15 ALA A  227  SER A  234  1                                   8
HELIX   16  16 THR A  250  ILE A  261  1                                  12
HELIX   17  17 PHE A  275  GLN A  280  1                                   6
HELIX   18  18 GLN A  280  ARG A  294  1                                  15
HELIX   19  19 HIS B    7  GLY B   11  5                                   5
HELIX   20  20 SER B   46  LEU B   49  5                                   4
HELIX   21  21 TRP B   50  LEU B   57  1                                   8
HELIX   22  22 THR B   82  ASP B   98  1                                  17
HELIX   23  23 TRP B  109  ASN B  121  1                                  13
HELIX   24  24 ASP B  139  LEU B  143  5                                   5
HELIX   25  25 SER B  144  GLN B  156  1                                  13
HELIX   26  26 GLY B  158  GLN B  165  1                                   8
HELIX   27  27 ASN B  168  GLU B  173  1                                   6
HELIX   28  28 GLU B  173  PHE B  178  1                                   6
HELIX   29  29 ALA B  184  THR B  193  1                                  10
HELIX   30  30 PRO B  198  SER B  200  5                                   3
HELIX   31  31 ARG B  201  LEU B  211  1                                  11
HELIX   32  32 PRO B  217  GLN B  226  1                                  10
HELIX   33  33 ALA B  227  SER B  234  1                                   8
HELIX   34  34 THR B  250  ILE B  261  1                                  12
HELIX   35  35 PHE B  275  GLU B  279  5                                   5
HELIX   36  36 GLN B  280  ALA B  296  1                                  17
SHEET    1   A 8 LYS A  15  ALA A  21  0
SHEET    2   A 8 LEU A  24  HIS A  31 -1  O  GLU A  30   N  LYS A  15
SHEET    3   A 8 ARG A  62  VAL A  66 -1  O  LEU A  63   N  HIS A  31
SHEET    4   A 8 PRO A  36  LEU A  40  1  N  ILE A  37   O  LEU A  64
SHEET    5   A 8 VAL A 102  ASP A 108  1  O  VAL A 105   N  VAL A  38
SHEET    6   A 8 VAL A 125  PRO A 133  1  O  ALA A 129   N  LEU A 104
SHEET    7   A 8 LYS A 238  PRO A 245  1  O  LEU A 239   N  PHE A 130
SHEET    8   A 8 LEU A 264  GLY A 272  1  O  GLU A 265   N  LYS A 238
SHEET    1   B 8 LYS B  15  ALA B  21  0
SHEET    2   B 8 LEU B  24  HIS B  31 -1  O  LEU B  24   N  ALA B  21
SHEET    3   B 8 ARG B  62  VAL B  66 -1  O  ALA B  65   N  VAL B  29
SHEET    4   B 8 PRO B  36  LEU B  40  1  N  ILE B  37   O  LEU B  64
SHEET    5   B 8 VAL B 102  ASP B 108  1  O  VAL B 105   N  VAL B  38
SHEET    6   B 8 VAL B 125  PRO B 133  1  O  ALA B 129   N  LEU B 104
SHEET    7   B 8 LYS B 238  PRO B 245  1  O  LEU B 239   N  PHE B 130
SHEET    8   B 8 LEU B 264  GLY B 272  1  O  ALA B 269   N  THR B 242
CISPEP   1 ASN A   43    PRO A   44          0         2.05
CISPEP   2 GLU A  216    PRO A  217          0        -1.37
CISPEP   3 LYS A  244    PRO A  245          0        -3.07
CISPEP   4 ASN B   43    PRO B   44          0        -0.35
CISPEP   5 GLU B  216    PRO B  217          0        -4.30
CISPEP   6 LYS B  244    PRO B  245          0         0.68
SITE     1 AC1  9 ASN A  43  ASP A 108  TRP A 109  PHE A 147
SITE     2 AC1  9 LEU A 174  GLY A 207  PRO A 208  PHE A 247
SITE     3 AC1  9 HIS A 274
SITE     1 AC2  8 TRP A 177  PHE A 179  PRO A 182  ILE A 273
SITE     2 AC2  8 HOH A 547  SER B 144  TRP B 177  PHE B 179
SITE     1 AC3  8 ASN B  43  ASP B 108  TRP B 109  PHE B 147
SITE     2 AC3  8 GLY B 207  PRO B 208  PHE B 247  HIS B 274
CRYST1  123.620  123.620   88.080  90.00  90.00  90.00 P 4 2 2      16
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008089  0.000000  0.000000        0.00000
SCALE2      0.000000  0.008089  0.000000        0.00000
SCALE3      0.000000  0.000000  0.011353        0.00000
TER    2322      ARG A 294
TER    4668      ALA B 296
MASTER      388    0    3   36   16    0    7    6 4873    2   39   48
END