longtext: 3WL6-pdb

content
HEADER    HYDROLASE                               08-NOV-13   3WL6
TITLE     CRYSTAL STRUCTURE OF POPH NATIVE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: OXIDIZED POLYVINYL ALCOHOL HYDROLASE;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: UNP RESIDUES 30-379;
COMPND   5 SYNONYM: OPH, OXIDIZED PVA HYDROLASE, BETA-DIKETONE HYDROLASE;
COMPND   6 EC: 3.7.1.7;
COMPND   7 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOMONAS SP.;
SOURCE   3 ORGANISM_TAXID: 306;
SOURCE   4 STRAIN: VM15C;
SOURCE   5 GENE: PVAB;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PET32A
KEYWDS    ALPHA/BETA-HYDROLASE, OXI-POLYVINYL ALCOHOL HYDROLASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Y.YANG,T.P.KO,J.H.LI,L.LIU,C.H.HUANG,H.C.CHAN,F.F.REN,D.X.JIA,A.H.-
AUTHOR   2 J.WANG,R.T.GUO,J.CHEN,G.C.DU
REVDAT   1   24-SEP-14 3WL6    0
JRNL        AUTH   Y.YANG,T.P.KO,L.LIU,J.LI,C.H.HUANG,H.C.CHAN,F.REN,D.JIA,
JRNL        AUTH 2 A.H.-J.WANG,R.T.GUO,J.CHEN,G.DU
JRNL        TITL   STRUCTURAL INSIGHTS INTO ENZYMATIC DEGRADATION OF OXIDIZED
JRNL        TITL 2 POLYVINYL ALCOHOL
JRNL        REF    CHEMBIOCHEM                   V.  15  1882 2014
JRNL        REFN                   ISSN 1439-4227
JRNL        PMID   25044912
JRNL        DOI    10.1002/CBIC.201402166
REMARK   2
REMARK   2 RESOLUTION.    1.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.21
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.60
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 25.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL
REMARK   3   NUMBER OF REFLECTIONS             : 83750
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.152
REMARK   3   FREE R VALUE                     : 0.174
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : 4254
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.60
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.66
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2090
REMARK   3   BIN FREE R VALUE                    : 0.2240
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 446
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 5002
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 13
REMARK   3   SOLVENT ATOMS            : 1049
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 19.44
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -7.82300
REMARK   3    B22 (A**2) : 2.87500
REMARK   3    B33 (A**2) : 4.94800
REMARK   3    B12 (A**2) : 5.13800
REMARK   3    B13 (A**2) : 0.10600
REMARK   3    B23 (A**2) : -1.38100
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.14
REMARK   3   ESD FROM SIGMAA              (A) : 0.10
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.17
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.12
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.013
REMARK   3   BOND ANGLES            (DEGREES) : 1.620
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.134 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 1.685 ; 2.000
REMARK   3   SIDE-CHAIN BOND              (A**2) : 2.032 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 2.894 ; 2.500
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : NULL
REMARK   3   KSOL        : NULL
REMARK   3   BSOL        : 41.33
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3WL6 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 26-NOV-13.
REMARK 100 THE RCSB ID CODE IS RCSB096473.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 02-MAY-13
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.6
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : NSRRC
REMARK 200  BEAMLINE                       : BL13B1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1
REMARK 200  MONOCHROMATOR                  : SI 111 CHANNEL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 85743
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.600
REMARK 200  RESOLUTION RANGE LOW       (A) : 25.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 95.6
REMARK 200  DATA REDUNDANCY                : 3.200
REMARK 200  R MERGE                    (I) : 0.05300
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 22.1000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.66
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.3
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.22000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 5.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 3WL5
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 45.13
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.24
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M TRI-SODIUM CITRATE, PH 5.6, 29%
REMARK 280  W/V POLYETHYLENE GLYCOL 4000, 3% W/V N-OCTYL-BETA-D-GLUCOSIDE,
REMARK 280  VAPOR DIFFUSION, SITTING DROP, TEMPERATURE 295K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ALA A    -4
REMARK 465     GLY A    -3
REMARK 465     ALA A    -2
REMARK 465     GLY A    -1
REMARK 465     ALA A     0
REMARK 465     ASP A     1
REMARK 465     ARG A     2
REMARK 465     THR A     3
REMARK 465     ALA A     4
REMARK 465     ALA A     5
REMARK 465     THR A     6
REMARK 465     PRO A     7
REMARK 465     ALA A     8
REMARK 465     ALA A     9
REMARK 465     ALA A    10
REMARK 465     ASN A    11
REMARK 465     PRO A    12
REMARK 465     ALA A    13
REMARK 465     ALA A    14
REMARK 465     THR A    15
REMARK 465     GLU A    16
REMARK 465     PRO A    17
REMARK 465     VAL A    18
REMARK 465     TRP A   352
REMARK 465     SER A   353
REMARK 465     HIS A   354
REMARK 465     PRO A   355
REMARK 465     GLN A   356
REMARK 465     PHE A   357
REMARK 465     GLU A   358
REMARK 465     LYS A   359
REMARK 465     ALA B    -4
REMARK 465     GLY B    -3
REMARK 465     ALA B    -2
REMARK 465     GLY B    -1
REMARK 465     ALA B     0
REMARK 465     ASP B     1
REMARK 465     ARG B     2
REMARK 465     THR B     3
REMARK 465     ALA B     4
REMARK 465     ALA B     5
REMARK 465     THR B     6
REMARK 465     PRO B     7
REMARK 465     ALA B     8
REMARK 465     ALA B     9
REMARK 465     ALA B    10
REMARK 465     ASN B    11
REMARK 465     PRO B    12
REMARK 465     ALA B    13
REMARK 465     ALA B    14
REMARK 465     THR B    15
REMARK 465     GLU B    16
REMARK 465     PRO B    17
REMARK 465     VAL B    18
REMARK 465     TRP B   255
REMARK 465     ASN B   256
REMARK 465     CYS B   257
REMARK 465     THR B   258
REMARK 465     ARG B   259
REMARK 465     PRO B   260
REMARK 465     ASP B   261
REMARK 465     GLY B   262
REMARK 465     SER B   263
REMARK 465     ARG B   264
REMARK 465     PHE B   265
REMARK 465     ALA B   351
REMARK 465     TRP B   352
REMARK 465     SER B   353
REMARK 465     HIS B   354
REMARK 465     PRO B   355
REMARK 465     GLN B   356
REMARK 465     PHE B   357
REMARK 465     GLU B   358
REMARK 465     LYS B   359
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH B   734     O    HOH B  1030              2.07
REMARK 500   N    LEU B   266     O    HOH B  1019              2.11
REMARK 500   O    HOH B   571     O    HOH B   581              2.12
REMARK 500   O    HOH A   870     O    HOH A   871              2.12
REMARK 500   O    HOH A   435     O    HOH A   498              2.14
REMARK 500   O    HOH A   866     O    HOH B   581              2.16
REMARK 500   O    HOH A   702     O    HOH A   910              2.17
REMARK 500   O    HOH A   693     O    HOH A   914              2.18
REMARK 500   O    HOH B   912     O    HOH B  1016              2.18
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    LEU B 168   CA  -  CB  -  CG  ANGL. DEV. =  14.4 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ALA A  81       55.44     38.13
REMARK 500    ARG A 107     -100.03   -100.99
REMARK 500    SER A 172     -124.49     61.33
REMARK 500    HIS A 298       78.01   -110.40
REMARK 500    ALA B  81       57.64     36.16
REMARK 500    ARG B 107      -95.97    -99.68
REMARK 500    SER B 172     -119.58     60.58
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 618        DISTANCE =  7.64 ANGSTROMS
REMARK 525    HOH A 635        DISTANCE =  5.40 ANGSTROMS
REMARK 525    HOH A 636        DISTANCE =  5.19 ANGSTROMS
REMARK 525    HOH A 650        DISTANCE =  6.31 ANGSTROMS
REMARK 525    HOH A 667        DISTANCE =  7.60 ANGSTROMS
REMARK 525    HOH A 678        DISTANCE =  5.12 ANGSTROMS
REMARK 525    HOH A 683        DISTANCE =  5.40 ANGSTROMS
REMARK 525    HOH A 689        DISTANCE =  5.54 ANGSTROMS
REMARK 525    HOH A 702        DISTANCE =  6.24 ANGSTROMS
REMARK 525    HOH A 713        DISTANCE =  5.54 ANGSTROMS
REMARK 525    HOH A 717        DISTANCE =  7.01 ANGSTROMS
REMARK 525    HOH A 730        DISTANCE =  5.26 ANGSTROMS
REMARK 525    HOH A 750        DISTANCE =  9.44 ANGSTROMS
REMARK 525    HOH A 763        DISTANCE =  6.72 ANGSTROMS
REMARK 525    HOH A 777        DISTANCE =  6.22 ANGSTROMS
REMARK 525    HOH A 784        DISTANCE =  9.80 ANGSTROMS
REMARK 525    HOH A 786        DISTANCE =  9.76 ANGSTROMS
REMARK 525    HOH A 792        DISTANCE =  6.95 ANGSTROMS
REMARK 525    HOH A 799        DISTANCE =  6.17 ANGSTROMS
REMARK 525    HOH A 804        DISTANCE =  7.07 ANGSTROMS
REMARK 525    HOH A 808        DISTANCE =  5.87 ANGSTROMS
REMARK 525    HOH A 814        DISTANCE =  6.23 ANGSTROMS
REMARK 525    HOH A 818        DISTANCE =  9.44 ANGSTROMS
REMARK 525    HOH A 820        DISTANCE =  6.65 ANGSTROMS
REMARK 525    HOH A 831        DISTANCE =  9.80 ANGSTROMS
REMARK 525    HOH A 844        DISTANCE =  6.92 ANGSTROMS
REMARK 525    HOH A 845        DISTANCE =  9.40 ANGSTROMS
REMARK 525    HOH A 885        DISTANCE =  5.90 ANGSTROMS
REMARK 525    HOH A 886        DISTANCE =  6.49 ANGSTROMS
REMARK 525    HOH A 888        DISTANCE =  7.61 ANGSTROMS
REMARK 525    HOH A 908        DISTANCE =  6.76 ANGSTROMS
REMARK 525    HOH A 909        DISTANCE =  8.30 ANGSTROMS
REMARK 525    HOH A 910        DISTANCE =  6.53 ANGSTROMS
REMARK 525    HOH A 918        DISTANCE =  9.74 ANGSTROMS
REMARK 525    HOH B 501        DISTANCE = 10.23 ANGSTROMS
REMARK 525    HOH B 502        DISTANCE = 11.34 ANGSTROMS
REMARK 525    HOH B 503        DISTANCE =  7.05 ANGSTROMS
REMARK 525    HOH B 504        DISTANCE = 11.89 ANGSTROMS
REMARK 525    HOH B 505        DISTANCE =  7.95 ANGSTROMS
REMARK 525    HOH B 506        DISTANCE = 10.86 ANGSTROMS
REMARK 525    HOH B 759        DISTANCE =  6.58 ANGSTROMS
REMARK 525    HOH B 765        DISTANCE =  6.78 ANGSTROMS
REMARK 525    HOH B 773        DISTANCE =  5.31 ANGSTROMS
REMARK 525    HOH B 788        DISTANCE =  6.46 ANGSTROMS
REMARK 525    HOH B 789        DISTANCE =  5.07 ANGSTROMS
REMARK 525    HOH B 812        DISTANCE =  7.26 ANGSTROMS
REMARK 525    HOH B 818        DISTANCE =  5.63 ANGSTROMS
REMARK 525    HOH B 820        DISTANCE =  6.51 ANGSTROMS
REMARK 525    HOH B 842        DISTANCE =  9.38 ANGSTROMS
REMARK 525    HOH B 849        DISTANCE =  6.13 ANGSTROMS
REMARK 525    HOH B 850        DISTANCE =  8.04 ANGSTROMS
REMARK 525    HOH B 856        DISTANCE =  6.19 ANGSTROMS
REMARK 525    HOH B 860        DISTANCE =  6.75 ANGSTROMS
REMARK 525    HOH B 872        DISTANCE =  5.45 ANGSTROMS
REMARK 525    HOH B 892        DISTANCE =  5.71 ANGSTROMS
REMARK 525    HOH B 893        DISTANCE =  7.27 ANGSTROMS
REMARK 525    HOH B 899        DISTANCE =  9.25 ANGSTROMS
REMARK 525    HOH B 907        DISTANCE =  6.51 ANGSTROMS
REMARK 525    HOH B 908        DISTANCE =  9.68 ANGSTROMS
REMARK 525    HOH B 910        DISTANCE =  5.39 ANGSTROMS
REMARK 525    HOH B 924        DISTANCE =  6.25 ANGSTROMS
REMARK 525    HOH B 925        DISTANCE =  5.42 ANGSTROMS
REMARK 525    HOH B 926        DISTANCE =  5.46 ANGSTROMS
REMARK 525    HOH B 931        DISTANCE =  5.21 ANGSTROMS
REMARK 525    HOH B 950        DISTANCE =  8.24 ANGSTROMS
REMARK 525    HOH B 951        DISTANCE =  6.28 ANGSTROMS
REMARK 525    HOH B 963        DISTANCE =  7.50 ANGSTROMS
REMARK 525    HOH B 984        DISTANCE =  5.46 ANGSTROMS
REMARK 525    HOH B 999        DISTANCE =  5.12 ANGSTROMS
REMARK 525    HOH B1001        DISTANCE =  8.23 ANGSTROMS
REMARK 525    HOH B1002        DISTANCE =  9.16 ANGSTROMS
REMARK 525    HOH B1009        DISTANCE =  6.70 ANGSTROMS
REMARK 525    HOH B1017        DISTANCE =  5.07 ANGSTROMS
REMARK 525    HOH B1021        DISTANCE =  6.72 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CIT B 401
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3WL5   RELATED DB: PDB
REMARK 900 RELATED ID: 3WL7   RELATED DB: PDB
REMARK 900 RELATED ID: 3WL8   RELATED DB: PDB
REMARK 900 RELATED ID: 3WLA   RELATED DB: PDB
DBREF  3WL6 A   -1   348  UNP    Q9LCQ7   OPH_PSESP       30    379
DBREF  3WL6 B   -1   348  UNP    Q9LCQ7   OPH_PSESP       30    379
SEQADV 3WL6 ALA A   -4  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 GLY A   -3  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA A   -2  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA A  349  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 SER A  350  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA A  351  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 TRP A  352  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 SER A  353  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 HIS A  354  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 PRO A  355  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 GLN A  356  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 PHE A  357  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 GLU A  358  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 LYS A  359  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA B   -4  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 GLY B   -3  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA B   -2  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA B  349  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 SER B  350  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 ALA B  351  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 TRP B  352  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 SER B  353  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 HIS B  354  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 PRO B  355  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 GLN B  356  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 PHE B  357  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 GLU B  358  UNP  Q9LCQ7              EXPRESSION TAG
SEQADV 3WL6 LYS B  359  UNP  Q9LCQ7              EXPRESSION TAG
SEQRES   1 A  364  ALA GLY ALA GLY ALA ASP ARG THR ALA ALA THR PRO ALA
SEQRES   2 A  364  ALA ALA ASN PRO ALA ALA THR GLU PRO VAL LYS TRP GLU
SEQRES   3 A  364  CYS PRO ALA GLY TYR GLU VAL LYS GLU GLY LEU ASN VAL
SEQRES   4 A  364  ASP PHE PRO HIS LYS GLY MET LYS ARG ALA PHE ILE VAL
SEQRES   5 A  364  TYR PRO ALA LYS ASN VAL SER GLY PRO ALA PRO VAL TRP
SEQRES   6 A  364  VAL PRO MET THR GLY SER VAL GLU SER THR ASN ASP ASN
SEQRES   7 A  364  LEU THR VAL ALA ARG SER GLY ALA ASN SER ILE LEU ALA
SEQRES   8 A  364  ASP HIS GLY TYR THR VAL ILE ALA PRO VAL ARG ALA CYS
SEQRES   9 A  364  ALA ASN GLN ASP PRO ASN ILE ARG GLY GLU ARG CYS ASN
SEQRES  10 A  364  GLY PRO GLY SER ASN GLY TRP ASN TRP ASN PRO TRP PHE
SEQRES  11 A  364  GLU GLY ARG ALA ALA ASP PRO SER GLY GLU HIS TRP LYS
SEQRES  12 A  364  ASN ASP GLU GLY PRO ASP SER SER PHE PHE VAL ALA MET
SEQRES  13 A  364  VAL GLN CYS VAL GLY THR LYS TYR LYS LEU ASP ALA ARG
SEQRES  14 A  364  ARG LEU PHE LEU GLY GLY ILE SER SER GLY GLY THR MET
SEQRES  15 A  364  THR ASN ARG ALA LEU LEU PHE ARG SER ASN PHE TRP ALA
SEQRES  16 A  364  GLY GLY LEU PRO ILE SER GLY GLU TRP TYR VAL THR SER
SEQRES  17 A  364  ASP ASP GLY THR PRO LEU SER PHE ASP ASP ALA ARG ALA
SEQRES  18 A  364  ALA VAL ALA ALA ALA PRO THR LYS ILE HIS GLN GLY ARG
SEQRES  19 A  364  VAL GLY PRO TYR PRO LEU PRO ALA LYS VAL GLY PRO LEU
SEQRES  20 A  364  ILE VAL MET THR VAL TRP GLY GLY GLU LYS ASP LEU TRP
SEQRES  21 A  364  ASN CYS THR ARG PRO ASP GLY SER ARG PHE LEU CYS ALA
SEQRES  22 A  364  ASP TYR ARG PRO SER THR GLN ALA GLY SER ASN PHE PHE
SEQRES  23 A  364  SER ALA GLN PRO ASP VAL VAL HIS VAL ALA CYS SER SER
SEQRES  24 A  364  THR HIS GLY HIS MET TRP PRO GLN LEU ASN THR GLN GLU
SEQRES  25 A  364  PHE ASN ARG TRP ALA LEU ASP THR LEU ALA SER HIS PRO
SEQRES  26 A  364  LYS GLY SER ASP PRO ARG SER PHE LYS LEU THR GLN PRO
SEQRES  27 A  364  PRO GLU GLY TYR THR CYS HIS VAL GLY PRO PHE THR GLY
SEQRES  28 A  364  LEU TYR ALA SER ALA TRP SER HIS PRO GLN PHE GLU LYS
SEQRES   1 B  364  ALA GLY ALA GLY ALA ASP ARG THR ALA ALA THR PRO ALA
SEQRES   2 B  364  ALA ALA ASN PRO ALA ALA THR GLU PRO VAL LYS TRP GLU
SEQRES   3 B  364  CYS PRO ALA GLY TYR GLU VAL LYS GLU GLY LEU ASN VAL
SEQRES   4 B  364  ASP PHE PRO HIS LYS GLY MET LYS ARG ALA PHE ILE VAL
SEQRES   5 B  364  TYR PRO ALA LYS ASN VAL SER GLY PRO ALA PRO VAL TRP
SEQRES   6 B  364  VAL PRO MET THR GLY SER VAL GLU SER THR ASN ASP ASN
SEQRES   7 B  364  LEU THR VAL ALA ARG SER GLY ALA ASN SER ILE LEU ALA
SEQRES   8 B  364  ASP HIS GLY TYR THR VAL ILE ALA PRO VAL ARG ALA CYS
SEQRES   9 B  364  ALA ASN GLN ASP PRO ASN ILE ARG GLY GLU ARG CYS ASN
SEQRES  10 B  364  GLY PRO GLY SER ASN GLY TRP ASN TRP ASN PRO TRP PHE
SEQRES  11 B  364  GLU GLY ARG ALA ALA ASP PRO SER GLY GLU HIS TRP LYS
SEQRES  12 B  364  ASN ASP GLU GLY PRO ASP SER SER PHE PHE VAL ALA MET
SEQRES  13 B  364  VAL GLN CYS VAL GLY THR LYS TYR LYS LEU ASP ALA ARG
SEQRES  14 B  364  ARG LEU PHE LEU GLY GLY ILE SER SER GLY GLY THR MET
SEQRES  15 B  364  THR ASN ARG ALA LEU LEU PHE ARG SER ASN PHE TRP ALA
SEQRES  16 B  364  GLY GLY LEU PRO ILE SER GLY GLU TRP TYR VAL THR SER
SEQRES  17 B  364  ASP ASP GLY THR PRO LEU SER PHE ASP ASP ALA ARG ALA
SEQRES  18 B  364  ALA VAL ALA ALA ALA PRO THR LYS ILE HIS GLN GLY ARG
SEQRES  19 B  364  VAL GLY PRO TYR PRO LEU PRO ALA LYS VAL GLY PRO LEU
SEQRES  20 B  364  ILE VAL MET THR VAL TRP GLY GLY GLU LYS ASP LEU TRP
SEQRES  21 B  364  ASN CYS THR ARG PRO ASP GLY SER ARG PHE LEU CYS ALA
SEQRES  22 B  364  ASP TYR ARG PRO SER THR GLN ALA GLY SER ASN PHE PHE
SEQRES  23 B  364  SER ALA GLN PRO ASP VAL VAL HIS VAL ALA CYS SER SER
SEQRES  24 B  364  THR HIS GLY HIS MET TRP PRO GLN LEU ASN THR GLN GLU
SEQRES  25 B  364  PHE ASN ARG TRP ALA LEU ASP THR LEU ALA SER HIS PRO
SEQRES  26 B  364  LYS GLY SER ASP PRO ARG SER PHE LYS LEU THR GLN PRO
SEQRES  27 B  364  PRO GLU GLY TYR THR CYS HIS VAL GLY PRO PHE THR GLY
SEQRES  28 B  364  LEU TYR ALA SER ALA TRP SER HIS PRO GLN PHE GLU LYS
HET    CIT  B 401      13
HETNAM     CIT CITRIC ACID
FORMUL   3  CIT    C6 H8 O7
FORMUL   4  HOH   *1049(H2 O)
HELIX    1   1 CYS A   22  TYR A   26  5                                   5
HELIX    2   2 SER A   69  VAL A   76  1                                   8
HELIX    3   3 ALA A   77  GLY A   80  5                                   4
HELIX    4   4 ALA A   81  GLY A   89  5                                   9
HELIX    5   5 ARG A   97  ASN A  101  5                                   5
HELIX    6   6 ASP A  131  ASN A  139  5                                   9
HELIX    7   7 GLY A  142  GLY A  156  1                                  15
HELIX    8   8 SER A  172  ARG A  185  1                                  14
HELIX    9   9 SER A  210  ALA A  221  1                                  12
HELIX   10  10 TYR A  270  GLN A  284  1                                  15
HELIX   11  11 ASN A  304  SER A  318  1                                  15
HELIX   12  12 ASP A  324  PHE A  328  5                                   5
HELIX   13  13 SER B   69  VAL B   76  1                                   8
HELIX   14  14 ALA B   77  GLY B   80  5                                   4
HELIX   15  15 ALA B   81  GLY B   89  5                                   9
HELIX   16  16 ARG B   97  ASN B  101  5                                   5
HELIX   17  17 ASP B  131  ASN B  139  5                                   9
HELIX   18  18 GLY B  142  TYR B  159  1                                  18
HELIX   19  19 SER B  172  ARG B  185  1                                  14
HELIX   20  20 SER B  210  ALA B  221  1                                  12
HELIX   21  21 ASP B  269  GLN B  284  1                                  16
HELIX   22  22 ASN B  304  SER B  318  1                                  15
HELIX   23  23 ASP B  324  PHE B  328  5                                   5
SHEET    1   A 9 GLY A  31  HIS A  38  0
SHEET    2   A 9 MET A  41  TYR A  48 -1  O  PHE A  45   N  ASN A  33
SHEET    3   A 9 THR A  91  PRO A  95 -1  O  VAL A  92   N  TYR A  48
SHEET    4   A 9 ALA A  57  MET A  63  1  N  PRO A  58   O  THR A  91
SHEET    5   A 9 LEU A 161  ILE A 171  1  O  ARG A 165   N  VAL A  59
SHEET    6   A 9 GLY A 191  ILE A 195  1  O  ILE A 195   N  GLY A 170
SHEET    7   A 9 LEU A 242  TRP A 248  1  O  MET A 245   N  GLY A 192
SHEET    8   A 9 VAL A 287  SER A 293  1  O  VAL A 290   N  VAL A 244
SHEET    9   A 9 THR A 338  VAL A 341 -1  O  HIS A 340   N  ALA A 291
SHEET    1   B 2 LEU A 254  THR A 258  0
SHEET    2   B 2 ARG A 264  ASP A 269 -1  O  CYS A 267   N  TRP A 255
SHEET    1   C 9 GLY B  31  HIS B  38  0
SHEET    2   C 9 MET B  41  TYR B  48 -1  O  ARG B  43   N  PHE B  36
SHEET    3   C 9 THR B  91  PRO B  95 -1  O  ALA B  94   N  ILE B  46
SHEET    4   C 9 ALA B  57  MET B  63  1  N  PRO B  58   O  THR B  91
SHEET    5   C 9 LEU B 161  ILE B 171  1  O  ASP B 162   N  ALA B  57
SHEET    6   C 9 GLY B 191  ILE B 195  1  O  ILE B 195   N  GLY B 170
SHEET    7   C 9 LEU B 242  TRP B 248  1  O  MET B 245   N  GLY B 192
SHEET    8   C 9 VAL B 287  SER B 293  1  O  VAL B 290   N  VAL B 244
SHEET    9   C 9 THR B 338  VAL B 341 -1  O  HIS B 340   N  ALA B 291
SSBOND   1 CYS A   22    CYS A  154                          1555   1555  2.08
SSBOND   2 CYS A   99    CYS A  111                          1555   1555  2.07
SSBOND   3 CYS A  257    CYS A  267                          1555   1555  2.06
SSBOND   4 CYS A  292    CYS A  339                          1555   1555  2.05
SSBOND   5 CYS B   22    CYS B  154                          1555   1555  2.07
SSBOND   6 CYS B   99    CYS B  111                          1555   1555  2.08
SSBOND   7 CYS B  292    CYS B  339                          1555   1555  2.06
CISPEP   1 TYR A  233    PRO A  234          0        -0.43
CISPEP   2 TYR B  233    PRO B  234          0        -0.43
SITE     1 AC1  9 SER B  66  VAL B  67  GLU B  68  ILE B 171
SITE     2 AC1  9 SER B 172  HIS B 298  HOH B 547  HOH B 909
SITE     3 AC1  9 HOH B1027
CRYST1   49.934   58.285   65.350  89.63  73.17  75.76 P 1           2
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.020026 -0.005082 -0.006428        0.00000
SCALE2      0.000000  0.017701  0.001239        0.00000
SCALE3      0.000000  0.000000  0.016026        0.00000
TER    2551      ALA A 351
TER    5004      SER B 350
MASTER      447    0    1   23   20    0    3    6 6064    2   27   56
END