longtext: 3WLA-pdb

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HEADER    HYDROLASE                               08-NOV-13   3WLA
TITLE     CRYSTAL STRUCTURE OF SOPH NATIVE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: OXIDIZED POLYVINYL ALCOHOL HYDROLASE;
COMPND   3 CHAIN: A, B, C;
COMPND   4 SYNONYM: OPH, OXIDIZED PVA HYDROLASE, BETA-DIKETONE HYDROLASE;
COMPND   5 EC: 3.7.1.7;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: SPHINGOPYXIS;
SOURCE   3 ORGANISM_TAXID: 292913;
SOURCE   4 STRAIN: 113P3;
SOURCE   5 GENE: OPH;
SOURCE   6 EXPRESSION_SYSTEM: PICHIA PASTORIS;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 4922;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: GS115;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PPIC9K
KEYWDS    ALPHA/BETA-HYDROLASE, OXI-POLYVINYL ALCOHOL HYDROLASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Y.YANG,T.P.KO,J.H.LI,L.LIU,C.H.HUANG,H.C.CHAN,F.F.REN,D.X.JIA,A.H.-
AUTHOR   2 J.WANG,R.T.GUO,J.CHEN,G.C.DU
REVDAT   1   24-SEP-14 3WLA    0
JRNL        AUTH   Y.YANG,T.P.KO,L.LIU,J.LI,C.H.HUANG,H.C.CHAN,F.REN,D.JIA,
JRNL        AUTH 2 A.H.-J.WANG,R.T.GUO,J.CHEN,G.DU
JRNL        TITL   STRUCTURAL INSIGHTS INTO ENZYMATIC DEGRADATION OF OXIDIZED
JRNL        TITL 2 POLYVINYL ALCOHOL
JRNL        REF    CHEMBIOCHEM                   V.  15  1882 2014
JRNL        REFN                   ISSN 1439-4227
JRNL        PMID   25044912
JRNL        DOI    10.1002/CBIC.201402166
REMARK   2
REMARK   2 RESOLUTION.    1.90 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.21
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : ENGH & HUBER
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.90
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 25.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : NULL
REMARK   3   NUMBER OF REFLECTIONS             : 88575
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.193
REMARK   3   FREE R VALUE                     : 0.239
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : NULL
REMARK   3   FREE R VALUE TEST SET COUNT      : 4475
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : NULL
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.90
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.97
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : NULL
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : NULL
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2990
REMARK   3   BIN FREE R VALUE                    : 0.3350
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : NULL
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 405
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 7542
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 20
REMARK   3   SOLVENT ATOMS            : 834
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 31.99
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 2.30100
REMARK   3    B22 (A**2) : -2.71900
REMARK   3    B33 (A**2) : 0.41800
REMARK   3    B12 (A**2) : 3.55300
REMARK   3    B13 (A**2) : 4.37800
REMARK   3    B23 (A**2) : 1.63100
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.24
REMARK   3   ESD FROM SIGMAA              (A) : 0.26
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.29
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.31
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.019
REMARK   3   BOND ANGLES            (DEGREES) : 1.840
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : NULL
REMARK   3   KSOL        : NULL
REMARK   3   BSOL        : 60.88
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : NULL
REMARK   3  TOPOLOGY FILE  1   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3WLA COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 27-NOV-13.
REMARK 100 THE RCSB ID CODE IS RCSB096477.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 13-JUL-12
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : NSRRC
REMARK 200  BEAMLINE                       : BL13B1
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1
REMARK 200  MONOCHROMATOR                  : SI 111 CHANNEL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 92945
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.900
REMARK 200  RESOLUTION RANGE LOW       (A) : 25.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.8
REMARK 200  DATA REDUNDANCY                : 3.000
REMARK 200  R MERGE                    (I) : 0.05600
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 26.2000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.90
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.97
REMARK 200  COMPLETENESS FOR SHELL     (%) : 96.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.00
REMARK 200  R MERGE FOR SHELL          (I) : 0.51000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.900
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 3WL5
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 56.48
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.83
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.2M LI2SO4, 0.1 M BIS-TRIS PH 6.5, 3%
REMARK 280  V/V POLYETHYLENE GLYCOL 500, 6% V/V POLYETHYLENE GLYCOL 600, 25%
REMARK 280  W/V POLYETHYLENE GLYCOL 3350, VAPOR DIFFUSION, SITTING DROP,
REMARK 280  TEMPERATURE 295K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A    -5
REMARK 465     ALA A    -4
REMARK 465     GLU A    -3
REMARK 465     ALA A    -2
REMARK 465     TYR A    -1
REMARK 465     VAL A     0
REMARK 465     LYS A     1
REMARK 465     GLU B    -5
REMARK 465     ALA B    -4
REMARK 465     GLU B    -3
REMARK 465     ALA B    -2
REMARK 465     TYR B    -1
REMARK 465     VAL B     0
REMARK 465     LYS B     1
REMARK 465     GLU C    -5
REMARK 465     ALA C    -4
REMARK 465     GLU C    -3
REMARK 465     ALA C    -2
REMARK 465     TYR C    -1
REMARK 465     VAL C     0
REMARK 465     LYS C     1
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH C   602     O    HOH C   670              2.15
REMARK 500   OD1  ASN C   285     O    HOH C   668              2.17
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A  37       68.63   -107.06
REMARK 500    ALA A  38       63.11    -39.61
REMARK 500    ALA A  92      -74.32   -109.54
REMARK 500    SER A 156     -122.35     57.31
REMARK 500    PRO A 194      163.62    -49.28
REMARK 500    SER B  37       67.10   -103.23
REMARK 500    ALA B  38       62.52    -39.23
REMARK 500    GLN B  86       17.05     55.90
REMARK 500    ALA B  92      -81.05   -121.07
REMARK 500    SER B 156     -123.30     55.20
REMARK 500    PHE B 172      -64.23    -93.85
REMARK 500    PRO B 282      123.05    -38.78
REMARK 500    ALA C   5      146.82   -176.70
REMARK 500    LYS C  35      -75.33    -38.48
REMARK 500    ASP C  36     -139.39   -103.60
REMARK 500    SER C  37       24.80   -159.33
REMARK 500    ALA C  38       79.53    -62.34
REMARK 500    ALA C  92      -72.68   -123.39
REMARK 500    SER C 156     -125.13     58.37
REMARK 500    PHE C 172      -61.45    -90.57
REMARK 500    PRO C 282      123.62    -38.32
REMARK 500    PRO C 315      171.53    -56.54
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    TYR A 318         0.08    SIDE CHAIN
REMARK 500    TYR C 318         0.07    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    ASN C 106        23.7      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A1179        DISTANCE =  5.93 ANGSTROMS
REMARK 525    HOH A1193        DISTANCE =  5.35 ANGSTROMS
REMARK 525    HOH A1247        DISTANCE =  5.34 ANGSTROMS
REMARK 525    HOH A1267        DISTANCE =  5.09 ANGSTROMS
REMARK 525    HOH A1281        DISTANCE =  7.36 ANGSTROMS
REMARK 525    HOH A1292        DISTANCE =  6.92 ANGSTROMS
REMARK 525    HOH A1295        DISTANCE =  7.92 ANGSTROMS
REMARK 525    HOH A1302        DISTANCE =  6.07 ANGSTROMS
REMARK 525    HOH A1303        DISTANCE =  7.83 ANGSTROMS
REMARK 525    HOH A1304        DISTANCE =  5.85 ANGSTROMS
REMARK 525    HOH A1312        DISTANCE =  8.00 ANGSTROMS
REMARK 525    HOH B 685        DISTANCE =  5.90 ANGSTROMS
REMARK 525    HOH B 748        DISTANCE =  7.25 ANGSTROMS
REMARK 525    HOH B 751        DISTANCE =  7.01 ANGSTROMS
REMARK 525    HOH B 752        DISTANCE =  7.93 ANGSTROMS
REMARK 525    HOH B 753        DISTANCE =  7.69 ANGSTROMS
REMARK 525    HOH B 756        DISTANCE =  6.60 ANGSTROMS
REMARK 525    HOH C 501        DISTANCE =  8.02 ANGSTROMS
REMARK 525    HOH C 665        DISTANCE =  7.90 ANGSTROMS
REMARK 525    HOH C 674        DISTANCE =  6.63 ANGSTROMS
REMARK 525    HOH C 679        DISTANCE =  5.82 ANGSTROMS
REMARK 525    HOH C 699        DISTANCE =  7.01 ANGSTROMS
REMARK 525    HOH C 718        DISTANCE =  6.43 ANGSTROMS
REMARK 525    HOH C 743        DISTANCE =  7.07 ANGSTROMS
REMARK 525    HOH C 747        DISTANCE =  5.21 ANGSTROMS
REMARK 525    HOH C 750        DISTANCE =  6.48 ANGSTROMS
REMARK 525    HOH C 754        DISTANCE =  5.66 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A 901
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 B 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 C 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 C 402
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 3WL5   RELATED DB: PDB
REMARK 900 RELATED ID: 3WL6   RELATED DB: PDB
REMARK 900 RELATED ID: 3WL7   RELATED DB: PDB
REMARK 900 RELATED ID: 3WL8   RELATED DB: PDB
DBREF  3WLA A    1   330  UNP    Q588Z2   OPH_SPHS1       35    364
DBREF  3WLA B    1   330  UNP    Q588Z2   OPH_SPHS1       35    364
DBREF  3WLA C    1   330  UNP    Q588Z2   OPH_SPHS1       35    364
SEQADV 3WLA GLU A   -5  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA ALA A   -4  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA GLU A   -3  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA ALA A   -2  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA TYR A   -1  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA VAL A    0  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA GLU B   -5  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA ALA B   -4  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA GLU B   -3  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA ALA B   -2  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA TYR B   -1  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA VAL B    0  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA GLU C   -5  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA ALA C   -4  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA GLU C   -3  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA ALA C   -2  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA TYR C   -1  UNP  Q588Z2              EXPRESSION TAG
SEQADV 3WLA VAL C    0  UNP  Q588Z2              EXPRESSION TAG
SEQRES   1 A  336  GLU ALA GLU ALA TYR VAL LYS SER GLU TRP ALA CYS PRO
SEQRES   2 A  336  GLU GLY PHE THR PRO LYS ALA GLY LEU ASN THR ASP PHE
SEQRES   3 A  336  PRO SER ASP GLY LYS LYS ARG ALA PHE VAL VAL VAL PRO
SEQRES   4 A  336  PRO LYS ASP SER ALA GLY GLY ALA PRO VAL TRP VAL PRO
SEQRES   5 A  336  MET VAL GLY THR VAL GLU ALA THR ASN TRP ASN LEU ASN
SEQRES   6 A  336  VAL PRO ARG SER GLY ASN ASN ALA LYS LEU ALA GLU HIS
SEQRES   7 A  336  GLY TYR MET VAL ILE SER PRO VAL ARG GLN CYS ALA GLU
SEQRES   8 A  336  GLN ASP PRO ASN LEU GLY ALA GLY ALA CYS ASN GLY VAL
SEQRES   9 A  336  GLY LYS ASP GLY TRP THR TRP ASN PRO TRP ASN ASP GLY
SEQRES  10 A  336  ARG ALA PRO ASP ALA SER GLY ASP LYS TYR LYS THR ASP
SEQRES  11 A  336  ALA GLY ASP ASP VAL ARG PHE LEU GLU ALA MET VAL ARG
SEQRES  12 A  336  CYS VAL GLY THR LYS TRP LYS LEU ASP ARG LYS ARG LEU
SEQRES  13 A  336  PHE LEU GLY GLY ILE SER ALA GLY GLY THR MET THR ASN
SEQRES  14 A  336  ARG ALA LEU LEU PHE ASP SER GLU PHE TRP ALA GLY GLY
SEQRES  15 A  336  MET PRO ILE SER GLY GLU TRP TYR SER THR LYS ASP ASP
SEQRES  16 A  336  GLY SER THR VAL PRO PHE GLN GLU THR ARG LYS MET VAL
SEQRES  17 A  336  ALA ALA ALA PRO ALA LYS ILE TRP GLN GLY ARG VAL GLY
SEQRES  18 A  336  PRO TYR PRO LEU PRO SER LYS LEU ASP PRO MET VAL VAL
SEQRES  19 A  336  ILE THR VAL TRP GLY GLY GLU LYS ASP LEU TRP ASP CYS
SEQRES  20 A  336  GLY PRO PRO LEU GLY LEU CYS SER ASP TYR ARG PRO THR
SEQRES  21 A  336  THR GLN ALA SER SER ASN TYR PHE SER SER ILE SER ASN
SEQRES  22 A  336  VAL VAL HIS VAL ALA CYS SER ALA THR HIS GLY HIS MET
SEQRES  23 A  336  TRP PRO GLN VAL ASN THR ASP ALA PHE ASN LEU TRP ALA
SEQRES  24 A  336  LEU ASN THR MET ALA SER HIS PRO LYS GLY SER SER PRO
SEQRES  25 A  336  LYS ASP PHE LYS LEU THR ALA PRO PRO GLU GLY TYR SER
SEQRES  26 A  336  CYS LYS ILE GLY ARG PHE THR ASP HIS TYR LYS
SEQRES   1 B  336  GLU ALA GLU ALA TYR VAL LYS SER GLU TRP ALA CYS PRO
SEQRES   2 B  336  GLU GLY PHE THR PRO LYS ALA GLY LEU ASN THR ASP PHE
SEQRES   3 B  336  PRO SER ASP GLY LYS LYS ARG ALA PHE VAL VAL VAL PRO
SEQRES   4 B  336  PRO LYS ASP SER ALA GLY GLY ALA PRO VAL TRP VAL PRO
SEQRES   5 B  336  MET VAL GLY THR VAL GLU ALA THR ASN TRP ASN LEU ASN
SEQRES   6 B  336  VAL PRO ARG SER GLY ASN ASN ALA LYS LEU ALA GLU HIS
SEQRES   7 B  336  GLY TYR MET VAL ILE SER PRO VAL ARG GLN CYS ALA GLU
SEQRES   8 B  336  GLN ASP PRO ASN LEU GLY ALA GLY ALA CYS ASN GLY VAL
SEQRES   9 B  336  GLY LYS ASP GLY TRP THR TRP ASN PRO TRP ASN ASP GLY
SEQRES  10 B  336  ARG ALA PRO ASP ALA SER GLY ASP LYS TYR LYS THR ASP
SEQRES  11 B  336  ALA GLY ASP ASP VAL ARG PHE LEU GLU ALA MET VAL ARG
SEQRES  12 B  336  CYS VAL GLY THR LYS TRP LYS LEU ASP ARG LYS ARG LEU
SEQRES  13 B  336  PHE LEU GLY GLY ILE SER ALA GLY GLY THR MET THR ASN
SEQRES  14 B  336  ARG ALA LEU LEU PHE ASP SER GLU PHE TRP ALA GLY GLY
SEQRES  15 B  336  MET PRO ILE SER GLY GLU TRP TYR SER THR LYS ASP ASP
SEQRES  16 B  336  GLY SER THR VAL PRO PHE GLN GLU THR ARG LYS MET VAL
SEQRES  17 B  336  ALA ALA ALA PRO ALA LYS ILE TRP GLN GLY ARG VAL GLY
SEQRES  18 B  336  PRO TYR PRO LEU PRO SER LYS LEU ASP PRO MET VAL VAL
SEQRES  19 B  336  ILE THR VAL TRP GLY GLY GLU LYS ASP LEU TRP ASP CYS
SEQRES  20 B  336  GLY PRO PRO LEU GLY LEU CYS SER ASP TYR ARG PRO THR
SEQRES  21 B  336  THR GLN ALA SER SER ASN TYR PHE SER SER ILE SER ASN
SEQRES  22 B  336  VAL VAL HIS VAL ALA CYS SER ALA THR HIS GLY HIS MET
SEQRES  23 B  336  TRP PRO GLN VAL ASN THR ASP ALA PHE ASN LEU TRP ALA
SEQRES  24 B  336  LEU ASN THR MET ALA SER HIS PRO LYS GLY SER SER PRO
SEQRES  25 B  336  LYS ASP PHE LYS LEU THR ALA PRO PRO GLU GLY TYR SER
SEQRES  26 B  336  CYS LYS ILE GLY ARG PHE THR ASP HIS TYR LYS
SEQRES   1 C  336  GLU ALA GLU ALA TYR VAL LYS SER GLU TRP ALA CYS PRO
SEQRES   2 C  336  GLU GLY PHE THR PRO LYS ALA GLY LEU ASN THR ASP PHE
SEQRES   3 C  336  PRO SER ASP GLY LYS LYS ARG ALA PHE VAL VAL VAL PRO
SEQRES   4 C  336  PRO LYS ASP SER ALA GLY GLY ALA PRO VAL TRP VAL PRO
SEQRES   5 C  336  MET VAL GLY THR VAL GLU ALA THR ASN TRP ASN LEU ASN
SEQRES   6 C  336  VAL PRO ARG SER GLY ASN ASN ALA LYS LEU ALA GLU HIS
SEQRES   7 C  336  GLY TYR MET VAL ILE SER PRO VAL ARG GLN CYS ALA GLU
SEQRES   8 C  336  GLN ASP PRO ASN LEU GLY ALA GLY ALA CYS ASN GLY VAL
SEQRES   9 C  336  GLY LYS ASP GLY TRP THR TRP ASN PRO TRP ASN ASP GLY
SEQRES  10 C  336  ARG ALA PRO ASP ALA SER GLY ASP LYS TYR LYS THR ASP
SEQRES  11 C  336  ALA GLY ASP ASP VAL ARG PHE LEU GLU ALA MET VAL ARG
SEQRES  12 C  336  CYS VAL GLY THR LYS TRP LYS LEU ASP ARG LYS ARG LEU
SEQRES  13 C  336  PHE LEU GLY GLY ILE SER ALA GLY GLY THR MET THR ASN
SEQRES  14 C  336  ARG ALA LEU LEU PHE ASP SER GLU PHE TRP ALA GLY GLY
SEQRES  15 C  336  MET PRO ILE SER GLY GLU TRP TYR SER THR LYS ASP ASP
SEQRES  16 C  336  GLY SER THR VAL PRO PHE GLN GLU THR ARG LYS MET VAL
SEQRES  17 C  336  ALA ALA ALA PRO ALA LYS ILE TRP GLN GLY ARG VAL GLY
SEQRES  18 C  336  PRO TYR PRO LEU PRO SER LYS LEU ASP PRO MET VAL VAL
SEQRES  19 C  336  ILE THR VAL TRP GLY GLY GLU LYS ASP LEU TRP ASP CYS
SEQRES  20 C  336  GLY PRO PRO LEU GLY LEU CYS SER ASP TYR ARG PRO THR
SEQRES  21 C  336  THR GLN ALA SER SER ASN TYR PHE SER SER ILE SER ASN
SEQRES  22 C  336  VAL VAL HIS VAL ALA CYS SER ALA THR HIS GLY HIS MET
SEQRES  23 C  336  TRP PRO GLN VAL ASN THR ASP ALA PHE ASN LEU TRP ALA
SEQRES  24 C  336  LEU ASN THR MET ALA SER HIS PRO LYS GLY SER SER PRO
SEQRES  25 C  336  LYS ASP PHE LYS LEU THR ALA PRO PRO GLU GLY TYR SER
SEQRES  26 C  336  CYS LYS ILE GLY ARG PHE THR ASP HIS TYR LYS
HET    SO4  A 901       5
HET    SO4  B 401       5
HET    SO4  C 401       5
HET    SO4  C 402       5
HETNAM     SO4 SULFATE ION
FORMUL   4  SO4    4(O4 S 2-)
FORMUL   8  HOH   *834(H2 O)
HELIX    1   1 ALA A   53  VAL A   60  1                                   8
HELIX    2   2 PRO A   61  GLY A   64  5                                   4
HELIX    3   3 ASN A   65  GLY A   73  5                                   9
HELIX    4   4 ARG A   81  GLU A   85  5                                   5
HELIX    5   5 ALA A   92  ASN A   96  5                                   5
HELIX    6   6 ASP A  115  THR A  123  5                                   9
HELIX    7   7 GLY A  126  GLY A  140  1                                  15
HELIX    8   8 SER A  156  ASP A  169  1                                  14
HELIX    9   9 PRO A  194  ALA A  205  1                                  12
HELIX   10  10 TYR A  251  SER A  264  1                                  14
HELIX   11  11 ASN A  285  SER A  299  1                                  15
HELIX   12  12 SER A  305  PHE A  309  5                                   5
HELIX   13  13 ALA B   53  VAL B   60  1                                   8
HELIX   14  14 PRO B   61  GLY B   64  5                                   4
HELIX   15  15 ASN B   65  GLY B   73  5                                   9
HELIX   16  16 ARG B   81  GLU B   85  5                                   5
HELIX   17  17 ALA B   92  ASN B   96  5                                   5
HELIX   18  18 ASP B  115  THR B  123  5                                   9
HELIX   19  19 GLY B  126  GLY B  140  1                                  15
HELIX   20  20 SER B  156  ASP B  169  1                                  14
HELIX   21  21 PRO B  194  ALA B  205  1                                  12
HELIX   22  22 TYR B  251  ILE B  265  1                                  15
HELIX   23  23 ASN B  285  SER B  299  1                                  15
HELIX   24  24 SER B  305  PHE B  309  5                                   5
HELIX   25  25 ALA C   53  VAL C   60  1                                   8
HELIX   26  26 PRO C   61  GLY C   64  5                                   4
HELIX   27  27 ASN C   65  GLY C   73  5                                   9
HELIX   28  28 ARG C   81  GLU C   85  5                                   5
HELIX   29  29 ALA C   92  ASN C   96  5                                   5
HELIX   30  30 ASP C  115  THR C  123  5                                   9
HELIX   31  31 GLY C  126  GLY C  140  1                                  15
HELIX   32  32 SER C  156  ASP C  169  1                                  14
HELIX   33  33 PRO C  194  ALA C  205  1                                  12
HELIX   34  34 TYR C  251  ILE C  265  1                                  15
HELIX   35  35 ASN C  285  SER C  299  1                                  15
HELIX   36  36 SER C  305  PHE C  309  5                                   5
SHEET    1   A 9 GLY A  15  SER A  22  0
SHEET    2   A 9 LYS A  25  VAL A  32 -1  O  ARG A  27   N  PHE A  20
SHEET    3   A 9 MET A  75  PRO A  79 -1  O  SER A  78   N  VAL A  30
SHEET    4   A 9 ALA A  41  MET A  47  1  N  PRO A  42   O  MET A  75
SHEET    5   A 9 LEU A 145  ILE A 155  1  O  ARG A 149   N  VAL A  43
SHEET    6   A 9 GLY A 175  ILE A 179  1  O  ILE A 179   N  GLY A 154
SHEET    7   A 9 MET A 226  TRP A 232  1  O  ILE A 229   N  GLY A 176
SHEET    8   A 9 VAL A 268  SER A 274  1  O  VAL A 271   N  VAL A 228
SHEET    9   A 9 SER A 319  ILE A 322 -1  O  LYS A 321   N  ALA A 272
SHEET    1   B 2 LEU A 238  ASP A 240  0
SHEET    2   B 2 LEU A 247  ASP A 250 -1  O  SER A 249   N  TRP A 239
SHEET    1   C 9 GLY B  15  SER B  22  0
SHEET    2   C 9 LYS B  25  VAL B  32 -1  O  ARG B  27   N  PHE B  20
SHEET    3   C 9 MET B  75  PRO B  79 -1  O  VAL B  76   N  VAL B  32
SHEET    4   C 9 ALA B  41  MET B  47  1  N  TRP B  44   O  MET B  75
SHEET    5   C 9 LEU B 145  ILE B 155  1  O  ARG B 149   N  VAL B  43
SHEET    6   C 9 GLY B 175  ILE B 179  1  O  ILE B 179   N  GLY B 154
SHEET    7   C 9 MET B 226  TRP B 232  1  O  ILE B 229   N  GLY B 176
SHEET    8   C 9 VAL B 268  SER B 274  1  O  CYS B 273   N  THR B 230
SHEET    9   C 9 SER B 319  ILE B 322 -1  O  LYS B 321   N  ALA B 272
SHEET    1   D 2 LEU B 238  ASP B 240  0
SHEET    2   D 2 LEU B 247  ASP B 250 -1  O  CYS B 248   N  TRP B 239
SHEET    1   E 9 GLY C  15  SER C  22  0
SHEET    2   E 9 LYS C  25  VAL C  32 -1  O  ARG C  27   N  PHE C  20
SHEET    3   E 9 MET C  75  PRO C  79 -1  O  VAL C  76   N  VAL C  32
SHEET    4   E 9 ALA C  41  MET C  47  1  N  PRO C  42   O  MET C  75
SHEET    5   E 9 LEU C 145  ILE C 155  1  O  ARG C 149   N  VAL C  43
SHEET    6   E 9 GLY C 175  ILE C 179  1  O  ILE C 179   N  GLY C 154
SHEET    7   E 9 MET C 226  TRP C 232  1  O  ILE C 229   N  GLY C 176
SHEET    8   E 9 VAL C 268  SER C 274  1  O  VAL C 271   N  VAL C 228
SHEET    9   E 9 SER C 319  ILE C 322 -1  O  SER C 319   N  SER C 274
SHEET    1   F 2 LEU C 238  ASP C 240  0
SHEET    2   F 2 LEU C 247  ASP C 250 -1  O  SER C 249   N  TRP C 239
SSBOND   1 CYS A    6    CYS A  138                          1555   1555  2.12
SSBOND   2 CYS A   83    CYS A   95                          1555   1555  2.07
SSBOND   3 CYS A  273    CYS A  320                          1555   1555  2.06
SSBOND   4 CYS B    6    CYS B  138                          1555   1555  2.10
SSBOND   5 CYS B   83    CYS B   95                          1555   1555  2.07
SSBOND   6 CYS B  273    CYS B  320                          1555   1555  2.07
SSBOND   7 CYS C    6    CYS C  138                          1555   1555  2.10
SSBOND   8 CYS C   83    CYS C   95                          1555   1555  2.06
SSBOND   9 CYS C  273    CYS C  320                          1555   1555  2.09
CISPEP   1 TYR A  217    PRO A  218          0        -0.52
CISPEP   2 PRO A  243    PRO A  244          0         0.37
CISPEP   3 TYR B  217    PRO B  218          0        -0.84
CISPEP   4 PRO B  243    PRO B  244          0        -0.58
CISPEP   5 TYR C  217    PRO C  218          0        -0.14
CISPEP   6 PRO C  243    PRO C  244          0         0.49
SITE     1 AC1  7 THR A  50  VAL A  51  SER A 156  TRP A 239
SITE     2 AC1  7 HOH A1012  HOH A1029  ARG C  62
SITE     1 AC2  6 THR B  50  VAL B  51  SER B 156  TRP B 239
SITE     2 AC2  6 HIS B 279  HOH B 523
SITE     1 AC3  7 ARG A  62  THR C  50  VAL C  51  SER C 156
SITE     2 AC3  7 HOH C 502  HOH C 520  HOH C 604
SITE     1 AC4 10 GLY A  91  ALA A  92  GLY A  93  GLY C  91
SITE     2 AC4 10 ALA C  92  GLY C  93  HOH C 559  HOH C 610
SITE     3 AC4 10 HOH C 624  HOH C 642
CRYST1   58.340   75.742   75.980  74.04  88.67  74.01 P 1           3
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.017141 -0.004912  0.001015        0.00000
SCALE2      0.000000  0.013734 -0.003997        0.00000
SCALE3      0.000000  0.000000  0.013711        0.00000
TER    2515      LYS A 330
TER    5030      LYS B 330
TER    7545      LYS C 330
MASTER      387    0    4   36   33    0    9    6 8396    3   38   78
END