longtext: 3WQH-pdb

content
HEADER    HYDROLASE/HYDROLASE INHIBITOR           27-JAN-14   3WQH
TITLE     CRYSTAL STRUCTURE OF HUMAN DPP-IV IN COMPLEX WITH ANAGLIPTIN
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: DIPEPTIDYL PEPTIDASE 4;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: UNP RESIDUES 39-766;
COMPND   5 SYNONYM: ADABP, ADENOSINE DEAMINASE COMPLEXING PROTEIN 2, ADCP-2,
COMPND   6 DIPEPTIDYL PEPTIDASE IV, DPP IV, T-CELL ACTIVATION ANTIGEN CD26,
COMPND   7 TP103, DIPEPTIDYL PEPTIDASE 4 MEMBRANE FORM, DIPEPTIDYL PEPTIDASE IV
COMPND   8 MEMBRANE FORM, DIPEPTIDYL PEPTIDASE 4 SOLUBLE FORM, DIPEPTIDYL
COMPND   9 PEPTIDASE IV SOLUBLE FORM;
COMPND  10 EC: 3.4.14.5;
COMPND  11 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: DPP4, ADCP2, CD26;
SOURCE   6 EXPRESSION_SYSTEM: BOMBYX MORI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 7091;
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: BACULOVIRUS
KEYWDS    DIABETES, BETA-PROPELLER, AMINOPEPTIDASE, MEMBRANE, HYDROLASE-
KEYWDS   2 HYDROLASE INHIBITOR COMPLEX
EXPDTA    X-RAY DIFFRACTION
AUTHOR    Y.S.WATANABE,S.OKADA,T.MOTOYAMA,R.TAKAHASHI,H.ADACHI,M.OKA
REVDAT   1   15-JUL-15 3WQH    0
JRNL        AUTH   Y.S.WATANABE,Y.YASUDA,Y.KOJIMA,S.OKADA,T.MOTOYAMA,
JRNL        AUTH 2 R.TAKAHASHI,M.OKA
JRNL        TITL   ANAGLIPTIN, A POTENT DIPEPTIDYL PEPTIDASE IV INHIBITOR: ITS
JRNL        TITL 2 SINGLE-CRYSTAL STRUCTURE AND ENZYME INTERACTIONS.
JRNL        REF    J ENZYME INHIB MED CHEM                  1 2015
JRNL        REFN                   ESSN 1475-6374
JRNL        PMID   26147347
JRNL        DOI    10.3109/14756366.2014.1002402
REMARK   2
REMARK   2 RESOLUTION.    2.85 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PRIMEX 2.2
REMARK   3   AUTHORS     : SCHRODINGER,LLC
REMARK   3
REMARK   3  REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.85
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 47.67
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   DATA CUTOFF HIGH  (ABS(F)/RMS(F)) : NULL
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : NULL
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 100.0
REMARK   3   NUMBER OF REFLECTIONS             : 45508
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : NULL
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.227
REMARK   3   FREE R VALUE                     : 0.288
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 2275
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.003
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 10
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.85
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.95
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.00
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 4550
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2920
REMARK   3   BIN FREE R VALUE                    : 0.3960
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 5.00
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 228
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.009
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 11926
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 239
REMARK   3   SOLVENT ATOMS            : 104
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 54.87
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : FLAT MODEL
REMARK   3   KSOL        : 0.32
REMARK   3   BSOL        : 56.52
REMARK   3   SOLVENT VOLUME (%) : NULL
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : NULL
REMARK   3   ESD FROM SIGMAA              (A) : NULL
REMARK   3   LOW RESOLUTION CUTOFF        (A) : NULL
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : NULL
REMARK   3   ESD FROM C-V SIGMAA          (A) : NULL
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.012
REMARK   3   BOND ANGLES            (DEGREES) : 1.853
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : NULL
REMARK   3   IMPROPER ANGLES        (DEGREES) : 0.376
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN BOND              (A**2) : NULL  ; NULL
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETERS AND TOPOLOGY.
REMARK   3   FORCE FIELD       : NULL
REMARK   3   PLANARITY WEIGHT  : NULL
REMARK   3   SOLVENT MODEL     : NULL
REMARK   3
REMARK   3  CLASH COUNT PER 100 RESIDUES.
REMARK   3   SEVERE : NULL
REMARK   3   OTHER  : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 3WQH COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 07-FEB-14.
REMARK 100 THE RCSB ID CODE IS RCSB096664.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 14-NOV-13
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SPRING-8
REMARK 200  BEAMLINE                       : BL41XU
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : RAYONIX MX225HE
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 45509
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.850
REMARK 200  RESOLUTION RANGE LOW       (A) : 49.110
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : NULL
REMARK 200  R MERGE                    (I) : 0.14900
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 12.1000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.85
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.92
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: 3BJM
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 55.86
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.79
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 18% PEG 4000, 15% PEG 1000, 0.15M NA/K
REMARK 280  -PHOSPHATE, PH 6.5, VAPOR DIFFUSION, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       32.58000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      209.80000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       34.49500
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      209.80000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       32.58000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       34.49500
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 8150 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 58210 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: 33.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLY A    37
REMARK 465     PRO A    38
REMARK 465     GLY B    37
REMARK 465     PRO B    38
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A  40      -20.72     87.85
REMARK 500    ASN A  51       47.95     77.34
REMARK 500    GLU A  73     -117.15     77.13
REMARK 500    ASN A  92        6.58     94.05
REMARK 500    SER A  93     -112.45     88.78
REMARK 500    THR A  94       26.61   -151.48
REMARK 500    PHE A  95      -57.30   -151.90
REMARK 500    PHE A  98      -16.13   -144.65
REMARK 500    ASN A 119       41.59     74.45
REMARK 500    GLN A 123     -112.76   -101.68
REMARK 500    TRP A 124     -144.68    -87.21
REMARK 500    ARG A 140       43.29     77.52
REMARK 500    ASN A 151        1.68     94.70
REMARK 500    HIS A 162       25.80   -149.03
REMARK 500    ILE A 185      -30.52   -133.89
REMARK 500    GLU A 191      115.79    -38.27
REMARK 500    ILE A 193      -74.57   -126.98
REMARK 500    ALA A 213       41.54   -141.08
REMARK 500    SER A 242     -162.14     71.92
REMARK 500    GLN A 320       42.91    -88.07
REMARK 500    PHE A 357      -41.45   -137.79
REMARK 500    ASP A 393     -171.21     74.42
REMARK 500    TRP A 402      169.54    170.46
REMARK 500    ASP A 413       -6.53   -141.51
REMARK 500    LYS A 423       -3.62     74.50
REMARK 500    LYS A 441       75.59   -107.37
REMARK 500    ASN A 450       75.14   -160.59
REMARK 500    SER A 473       28.25   -146.80
REMARK 500    ASN A 520     -107.50   -133.03
REMARK 500    TYR A 547      -68.59   -124.05
REMARK 500    ALA A 548        1.48     81.15
REMARK 500    CYS A 551       18.59     80.23
REMARK 500    ASN A 572       42.18     70.84
REMARK 500    TYR A 585       16.51     83.49
REMARK 500    ARG A 596       32.68     74.91
REMARK 500    ARG A 597       31.65   -164.70
REMARK 500    THR A 600      -87.17   -107.16
REMARK 500    SER A 630     -104.62     83.62
REMARK 500    ALA A 654       53.85     35.97
REMARK 500    PRO A 674       35.12    -75.44
REMARK 500    ASP A 678      -94.34   -120.00
REMARK 500    ASN A 679       26.16   -142.16
REMARK 500    ASN A 710      -86.31   -101.93
REMARK 500    ASP A 739     -157.66    -85.66
REMARK 500    ASN B  51       42.99     75.36
REMARK 500    GLU B  73       11.04     81.30
REMARK 500    PHE B  95       14.06     92.43
REMARK 500    GLN B 123     -104.50   -100.24
REMARK 500    TRP B 124     -141.65    -94.45
REMARK 500    ASN B 151       34.30     79.77
REMARK 500
REMARK 500 THIS ENTRY HAS      79 RAMACHANDRAN OUTLIERS.
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 ASN A   92     SER A   93                  137.56
REMARK 500 PHE A   95     ASP A   96                 -136.30
REMARK 500 ILE A  742     ALA A  743                 -148.56
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    ASN A  51        24.2      L          L   OUTSIDE RANGE
REMARK 500    ASN A  92        19.8      L          L   OUTSIDE RANGE
REMARK 500    SER A  93        21.4      L          L   OUTSIDE RANGE
REMARK 500    ARG A 140        24.9      L          L   OUTSIDE RANGE
REMARK 500    ASN A 151        22.3      L          L   OUTSIDE RANGE
REMARK 500    ASN A 170        24.8      L          L   OUTSIDE RANGE
REMARK 500    ASP A 192        23.5      L          L   OUTSIDE RANGE
REMARK 500    ASN A 497        25.0      L          L   OUTSIDE RANGE
REMARK 500    ALA A 548        23.6      L          L   OUTSIDE RANGE
REMARK 500    SER A 630        24.6      L          L   OUTSIDE RANGE
REMARK 500    ASP A 737        24.0      L          L   OUTSIDE RANGE
REMARK 500    ASN B  74        24.0      L          L   OUTSIDE RANGE
REMARK 500    PHE B  95        22.1      L          L   OUTSIDE RANGE
REMARK 500    ASN B 170        24.9      L          L   OUTSIDE RANGE
REMARK 500    ASP B 192        25.0      L          L   OUTSIDE RANGE
REMARK 500    GLU B 464        24.5      L          L   OUTSIDE RANGE
REMARK 500    ASP B 488        24.6      L          L   OUTSIDE RANGE
REMARK 500    GLU B 521        23.7      L          L   OUTSIDE RANGE
REMARK 500    ALA B 548        23.6      L          L   OUTSIDE RANGE
REMARK 500    TYR B 585        24.9      L          L   OUTSIDE RANGE
REMARK 500    SER B 630        23.0      L          L   OUTSIDE RANGE
REMARK 500    ASP B 737        22.6      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 807
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SKK A 808
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SKK B 807
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG A 801 BOUND
REMARK 800  TO ASN A 85
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG A 802 BOUND
REMARK 800  TO ASN A 150
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG A 803 BOUND
REMARK 800  TO ASN A 219
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG A 804 BOUND
REMARK 800  TO ASN A 229
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG A 805 BOUND
REMARK 800  TO ASN A 281
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG A 806 BOUND
REMARK 800  TO ASN A 321
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG B 801 BOUND
REMARK 800  TO ASN B 85
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG B 802 BOUND
REMARK 800  TO ASN B 150
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG B 803 BOUND
REMARK 800  TO ASN B 219
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG B 804 BOUND
REMARK 800  TO ASN B 229
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG B 805 BOUND
REMARK 800  TO ASN B 281
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE NAG B 806 BOUND
REMARK 800  TO ASN B 321
DBREF  3WQH A   39   766  UNP    P27487   DPP4_HUMAN      39    766
DBREF  3WQH B   39   766  UNP    P27487   DPP4_HUMAN      39    766
SEQADV 3WQH GLY A   37  UNP  P27487              EXPRESSION TAG
SEQADV 3WQH PRO A   38  UNP  P27487              EXPRESSION TAG
SEQADV 3WQH GLY B   37  UNP  P27487              EXPRESSION TAG
SEQADV 3WQH PRO B   38  UNP  P27487              EXPRESSION TAG
SEQRES   1 A  730  GLY PRO SER ARG LYS THR TYR THR LEU THR ASP TYR LEU
SEQRES   2 A  730  LYS ASN THR TYR ARG LEU LYS LEU TYR SER LEU ARG TRP
SEQRES   3 A  730  ILE SER ASP HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN
SEQRES   4 A  730  ILE LEU VAL PHE ASN ALA GLU TYR GLY ASN SER SER VAL
SEQRES   5 A  730  PHE LEU GLU ASN SER THR PHE ASP GLU PHE GLY HIS SER
SEQRES   6 A  730  ILE ASN ASP TYR SER ILE SER PRO ASP GLY GLN PHE ILE
SEQRES   7 A  730  LEU LEU GLU TYR ASN TYR VAL LYS GLN TRP ARG HIS SER
SEQRES   8 A  730  TYR THR ALA SER TYR ASP ILE TYR ASP LEU ASN LYS ARG
SEQRES   9 A  730  GLN LEU ILE THR GLU GLU ARG ILE PRO ASN ASN THR GLN
SEQRES  10 A  730  TRP VAL THR TRP SER PRO VAL GLY HIS LYS LEU ALA TYR
SEQRES  11 A  730  VAL TRP ASN ASN ASP ILE TYR VAL LYS ILE GLU PRO ASN
SEQRES  12 A  730  LEU PRO SER TYR ARG ILE THR TRP THR GLY LYS GLU ASP
SEQRES  13 A  730  ILE ILE TYR ASN GLY ILE THR ASP TRP VAL TYR GLU GLU
SEQRES  14 A  730  GLU VAL PHE SER ALA TYR SER ALA LEU TRP TRP SER PRO
SEQRES  15 A  730  ASN GLY THR PHE LEU ALA TYR ALA GLN PHE ASN ASP THR
SEQRES  16 A  730  GLU VAL PRO LEU ILE GLU TYR SER PHE TYR SER ASP GLU
SEQRES  17 A  730  SER LEU GLN TYR PRO LYS THR VAL ARG VAL PRO TYR PRO
SEQRES  18 A  730  LYS ALA GLY ALA VAL ASN PRO THR VAL LYS PHE PHE VAL
SEQRES  19 A  730  VAL ASN THR ASP SER LEU SER SER VAL THR ASN ALA THR
SEQRES  20 A  730  SER ILE GLN ILE THR ALA PRO ALA SER MET LEU ILE GLY
SEQRES  21 A  730  ASP HIS TYR LEU CYS ASP VAL THR TRP ALA THR GLN GLU
SEQRES  22 A  730  ARG ILE SER LEU GLN TRP LEU ARG ARG ILE GLN ASN TYR
SEQRES  23 A  730  SER VAL MET ASP ILE CYS ASP TYR ASP GLU SER SER GLY
SEQRES  24 A  730  ARG TRP ASN CYS LEU VAL ALA ARG GLN HIS ILE GLU MET
SEQRES  25 A  730  SER THR THR GLY TRP VAL GLY ARG PHE ARG PRO SER GLU
SEQRES  26 A  730  PRO HIS PHE THR LEU ASP GLY ASN SER PHE TYR LYS ILE
SEQRES  27 A  730  ILE SER ASN GLU GLU GLY TYR ARG HIS ILE CYS TYR PHE
SEQRES  28 A  730  GLN ILE ASP LYS LYS ASP CYS THR PHE ILE THR LYS GLY
SEQRES  29 A  730  THR TRP GLU VAL ILE GLY ILE GLU ALA LEU THR SER ASP
SEQRES  30 A  730  TYR LEU TYR TYR ILE SER ASN GLU TYR LYS GLY MET PRO
SEQRES  31 A  730  GLY GLY ARG ASN LEU TYR LYS ILE GLN LEU SER ASP TYR
SEQRES  32 A  730  THR LYS VAL THR CYS LEU SER CYS GLU LEU ASN PRO GLU
SEQRES  33 A  730  ARG CYS GLN TYR TYR SER VAL SER PHE SER LYS GLU ALA
SEQRES  34 A  730  LYS TYR TYR GLN LEU ARG CYS SER GLY PRO GLY LEU PRO
SEQRES  35 A  730  LEU TYR THR LEU HIS SER SER VAL ASN ASP LYS GLY LEU
SEQRES  36 A  730  ARG VAL LEU GLU ASP ASN SER ALA LEU ASP LYS MET LEU
SEQRES  37 A  730  GLN ASN VAL GLN MET PRO SER LYS LYS LEU ASP PHE ILE
SEQRES  38 A  730  ILE LEU ASN GLU THR LYS PHE TRP TYR GLN MET ILE LEU
SEQRES  39 A  730  PRO PRO HIS PHE ASP LYS SER LYS LYS TYR PRO LEU LEU
SEQRES  40 A  730  LEU ASP VAL TYR ALA GLY PRO CYS SER GLN LYS ALA ASP
SEQRES  41 A  730  THR VAL PHE ARG LEU ASN TRP ALA THR TYR LEU ALA SER
SEQRES  42 A  730  THR GLU ASN ILE ILE VAL ALA SER PHE ASP GLY ARG GLY
SEQRES  43 A  730  SER GLY TYR GLN GLY ASP LYS ILE MET HIS ALA ILE ASN
SEQRES  44 A  730  ARG ARG LEU GLY THR PHE GLU VAL GLU ASP GLN ILE GLU
SEQRES  45 A  730  ALA ALA ARG GLN PHE SER LYS MET GLY PHE VAL ASP ASN
SEQRES  46 A  730  LYS ARG ILE ALA ILE TRP GLY TRP SER TYR GLY GLY TYR
SEQRES  47 A  730  VAL THR SER MET VAL LEU GLY SER GLY SER GLY VAL PHE
SEQRES  48 A  730  LYS CYS GLY ILE ALA VAL ALA PRO VAL SER ARG TRP GLU
SEQRES  49 A  730  TYR TYR ASP SER VAL TYR THR GLU ARG TYR MET GLY LEU
SEQRES  50 A  730  PRO THR PRO GLU ASP ASN LEU ASP HIS TYR ARG ASN SER
SEQRES  51 A  730  THR VAL MET SER ARG ALA GLU ASN PHE LYS GLN VAL GLU
SEQRES  52 A  730  TYR LEU LEU ILE HIS GLY THR ALA ASP ASP ASN VAL HIS
SEQRES  53 A  730  PHE GLN GLN SER ALA GLN ILE SER LYS ALA LEU VAL ASP
SEQRES  54 A  730  VAL GLY VAL ASP PHE GLN ALA MET TRP TYR THR ASP GLU
SEQRES  55 A  730  ASP HIS GLY ILE ALA SER SER THR ALA HIS GLN HIS ILE
SEQRES  56 A  730  TYR THR HIS MET SER HIS PHE ILE LYS GLN CYS PHE SER
SEQRES  57 A  730  LEU PRO
SEQRES   1 B  730  GLY PRO SER ARG LYS THR TYR THR LEU THR ASP TYR LEU
SEQRES   2 B  730  LYS ASN THR TYR ARG LEU LYS LEU TYR SER LEU ARG TRP
SEQRES   3 B  730  ILE SER ASP HIS GLU TYR LEU TYR LYS GLN GLU ASN ASN
SEQRES   4 B  730  ILE LEU VAL PHE ASN ALA GLU TYR GLY ASN SER SER VAL
SEQRES   5 B  730  PHE LEU GLU ASN SER THR PHE ASP GLU PHE GLY HIS SER
SEQRES   6 B  730  ILE ASN ASP TYR SER ILE SER PRO ASP GLY GLN PHE ILE
SEQRES   7 B  730  LEU LEU GLU TYR ASN TYR VAL LYS GLN TRP ARG HIS SER
SEQRES   8 B  730  TYR THR ALA SER TYR ASP ILE TYR ASP LEU ASN LYS ARG
SEQRES   9 B  730  GLN LEU ILE THR GLU GLU ARG ILE PRO ASN ASN THR GLN
SEQRES  10 B  730  TRP VAL THR TRP SER PRO VAL GLY HIS LYS LEU ALA TYR
SEQRES  11 B  730  VAL TRP ASN ASN ASP ILE TYR VAL LYS ILE GLU PRO ASN
SEQRES  12 B  730  LEU PRO SER TYR ARG ILE THR TRP THR GLY LYS GLU ASP
SEQRES  13 B  730  ILE ILE TYR ASN GLY ILE THR ASP TRP VAL TYR GLU GLU
SEQRES  14 B  730  GLU VAL PHE SER ALA TYR SER ALA LEU TRP TRP SER PRO
SEQRES  15 B  730  ASN GLY THR PHE LEU ALA TYR ALA GLN PHE ASN ASP THR
SEQRES  16 B  730  GLU VAL PRO LEU ILE GLU TYR SER PHE TYR SER ASP GLU
SEQRES  17 B  730  SER LEU GLN TYR PRO LYS THR VAL ARG VAL PRO TYR PRO
SEQRES  18 B  730  LYS ALA GLY ALA VAL ASN PRO THR VAL LYS PHE PHE VAL
SEQRES  19 B  730  VAL ASN THR ASP SER LEU SER SER VAL THR ASN ALA THR
SEQRES  20 B  730  SER ILE GLN ILE THR ALA PRO ALA SER MET LEU ILE GLY
SEQRES  21 B  730  ASP HIS TYR LEU CYS ASP VAL THR TRP ALA THR GLN GLU
SEQRES  22 B  730  ARG ILE SER LEU GLN TRP LEU ARG ARG ILE GLN ASN TYR
SEQRES  23 B  730  SER VAL MET ASP ILE CYS ASP TYR ASP GLU SER SER GLY
SEQRES  24 B  730  ARG TRP ASN CYS LEU VAL ALA ARG GLN HIS ILE GLU MET
SEQRES  25 B  730  SER THR THR GLY TRP VAL GLY ARG PHE ARG PRO SER GLU
SEQRES  26 B  730  PRO HIS PHE THR LEU ASP GLY ASN SER PHE TYR LYS ILE
SEQRES  27 B  730  ILE SER ASN GLU GLU GLY TYR ARG HIS ILE CYS TYR PHE
SEQRES  28 B  730  GLN ILE ASP LYS LYS ASP CYS THR PHE ILE THR LYS GLY
SEQRES  29 B  730  THR TRP GLU VAL ILE GLY ILE GLU ALA LEU THR SER ASP
SEQRES  30 B  730  TYR LEU TYR TYR ILE SER ASN GLU TYR LYS GLY MET PRO
SEQRES  31 B  730  GLY GLY ARG ASN LEU TYR LYS ILE GLN LEU SER ASP TYR
SEQRES  32 B  730  THR LYS VAL THR CYS LEU SER CYS GLU LEU ASN PRO GLU
SEQRES  33 B  730  ARG CYS GLN TYR TYR SER VAL SER PHE SER LYS GLU ALA
SEQRES  34 B  730  LYS TYR TYR GLN LEU ARG CYS SER GLY PRO GLY LEU PRO
SEQRES  35 B  730  LEU TYR THR LEU HIS SER SER VAL ASN ASP LYS GLY LEU
SEQRES  36 B  730  ARG VAL LEU GLU ASP ASN SER ALA LEU ASP LYS MET LEU
SEQRES  37 B  730  GLN ASN VAL GLN MET PRO SER LYS LYS LEU ASP PHE ILE
SEQRES  38 B  730  ILE LEU ASN GLU THR LYS PHE TRP TYR GLN MET ILE LEU
SEQRES  39 B  730  PRO PRO HIS PHE ASP LYS SER LYS LYS TYR PRO LEU LEU
SEQRES  40 B  730  LEU ASP VAL TYR ALA GLY PRO CYS SER GLN LYS ALA ASP
SEQRES  41 B  730  THR VAL PHE ARG LEU ASN TRP ALA THR TYR LEU ALA SER
SEQRES  42 B  730  THR GLU ASN ILE ILE VAL ALA SER PHE ASP GLY ARG GLY
SEQRES  43 B  730  SER GLY TYR GLN GLY ASP LYS ILE MET HIS ALA ILE ASN
SEQRES  44 B  730  ARG ARG LEU GLY THR PHE GLU VAL GLU ASP GLN ILE GLU
SEQRES  45 B  730  ALA ALA ARG GLN PHE SER LYS MET GLY PHE VAL ASP ASN
SEQRES  46 B  730  LYS ARG ILE ALA ILE TRP GLY TRP SER TYR GLY GLY TYR
SEQRES  47 B  730  VAL THR SER MET VAL LEU GLY SER GLY SER GLY VAL PHE
SEQRES  48 B  730  LYS CYS GLY ILE ALA VAL ALA PRO VAL SER ARG TRP GLU
SEQRES  49 B  730  TYR TYR ASP SER VAL TYR THR GLU ARG TYR MET GLY LEU
SEQRES  50 B  730  PRO THR PRO GLU ASP ASN LEU ASP HIS TYR ARG ASN SER
SEQRES  51 B  730  THR VAL MET SER ARG ALA GLU ASN PHE LYS GLN VAL GLU
SEQRES  52 B  730  TYR LEU LEU ILE HIS GLY THR ALA ASP ASP ASN VAL HIS
SEQRES  53 B  730  PHE GLN GLN SER ALA GLN ILE SER LYS ALA LEU VAL ASP
SEQRES  54 B  730  VAL GLY VAL ASP PHE GLN ALA MET TRP TYR THR ASP GLU
SEQRES  55 B  730  ASP HIS GLY ILE ALA SER SER THR ALA HIS GLN HIS ILE
SEQRES  56 B  730  TYR THR HIS MET SER HIS PHE ILE LYS GLN CYS PHE SER
SEQRES  57 B  730  LEU PRO
MODRES 3WQH ASN A  321  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN B  150  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN A  229  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN A   85  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN A  281  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN B  219  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN B   85  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN B  321  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN A  219  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN A  150  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN B  229  ASN  GLYCOSYLATION SITE
MODRES 3WQH ASN B  281  ASN  GLYCOSYLATION SITE
HET    NAG  A 801      14
HET    NAG  A 802      14
HET    NAG  A 803      14
HET    NAG  A 804      14
HET    NAG  A 805      14
HET    NAG  A 806      14
HET    NAG  A 807      15
HET    SKK  A 808      28
HET    NAG  B 801      14
HET    NAG  B 802      14
HET    NAG  B 803      14
HET    NAG  B 804      14
HET    NAG  B 805      14
HET    NAG  B 806      14
HET    SKK  B 807      28
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     SKK N-[2-({2-[(2S)-2-CYANOPYRROLIDIN-1-YL]-2-
HETNAM   2 SKK  OXOETHYL}AMINO)-2-METHYLPROPYL]-2-METHYLPYRAZOLO[1,5-
HETNAM   3 SKK  A]PYRIMIDINE-6-CARBOXAMIDE
HETSYN     SKK ANAGLIPTIN
FORMUL   3  NAG    13(C8 H15 N O6)
FORMUL  10  SKK    2(C19 H25 N7 O2)
FORMUL  18  HOH   *104(H2 O)
HELIX    1   1 THR A   44  ASN A   51  1                                   8
HELIX    2   2 ASP A  200  VAL A  207  1                                   8
HELIX    3   3 PRO A  290  ILE A  295  1                                   6
HELIX    4   4 LEU A  340  GLN A  344  5                                   5
HELIX    5   5 GLU A  421  MET A  425  5                                   5
HELIX    6   6 LYS A  463  ALA A  465  5                                   3
HELIX    7   7 ASN A  497  GLN A  505  1                                   9
HELIX    8   8 ASN A  562  THR A  570  1                                   9
HELIX    9   9 GLY A  587  HIS A  592  1                                   6
HELIX   10  10 ALA A  593  ASN A  595  5                                   3
HELIX   11  11 THR A  600  MET A  616  1                                  17
HELIX   12  12 SER A  630  GLY A  641  1                                  12
HELIX   13  13 ARG A  658  TYR A  662  5                                   5
HELIX   14  14 ASP A  663  GLY A  672  1                                  10
HELIX   15  15 ASN A  679  SER A  686  1                                   8
HELIX   16  16 VAL A  688  VAL A  698  5                                  11
HELIX   17  17 PHE A  713  ASP A  725  1                                  13
HELIX   18  18 SER A  744  PHE A  763  1                                  20
HELIX   19  19 THR B   44  ASN B   51  1                                   8
HELIX   20  20 ASP B  200  VAL B  207  1                                   8
HELIX   21  21 ASP B  274  LEU B  276  5                                   3
HELIX   22  22 PRO B  290  ILE B  295  1                                   6
HELIX   23  23 VAL B  341  GLN B  344  5                                   4
HELIX   24  24 GLU B  421  MET B  425  5                                   5
HELIX   25  25 LYS B  463  ALA B  465  5                                   3
HELIX   26  26 ASN B  497  GLN B  505  1                                   9
HELIX   27  27 ASN B  562  THR B  570  1                                   9
HELIX   28  28 GLY B  587  HIS B  592  1                                   6
HELIX   29  29 ALA B  593  ASN B  595  5                                   3
HELIX   30  30 THR B  600  MET B  616  1                                  17
HELIX   31  31 SER B  630  GLY B  641  1                                  12
HELIX   32  32 ARG B  658  TYR B  662  5                                   5
HELIX   33  33 ASP B  663  GLY B  672  1                                  10
HELIX   34  34 ASN B  679  SER B  686  1                                   8
HELIX   35  35 VAL B  688  VAL B  698  5                                  11
HELIX   36  36 HIS B  712  ASP B  725  1                                  14
HELIX   37  37 SER B  744  PHE B  763  1                                  20
SHEET    1   A 4 ARG A  61  TRP A  62  0
SHEET    2   A 4 GLU A  67  GLN A  72 -1  O  LEU A  69   N  ARG A  61
SHEET    3   A 4 ASN A  75  ASN A  80 -1  O  LEU A  77   N  TYR A  70
SHEET    4   A 4 SER A  86  LEU A  90 -1  O  SER A  87   N  VAL A  78
SHEET    1   B 4 ILE A 102  ILE A 107  0
SHEET    2   B 4 PHE A 113  LYS A 122 -1  O  LEU A 115   N  SER A 106
SHEET    3   B 4 TYR A 128  ASP A 136 -1  O  SER A 131   N  TYR A 118
SHEET    4   B 4 GLN A 141  LEU A 142 -1  O  GLN A 141   N  ASP A 136
SHEET    1   C 4 ILE A 102  ILE A 107  0
SHEET    2   C 4 PHE A 113  LYS A 122 -1  O  LEU A 115   N  SER A 106
SHEET    3   C 4 TYR A 128  ASP A 136 -1  O  SER A 131   N  TYR A 118
SHEET    4   C 4 ILE A 148  THR A 152 -1  O  THR A 152   N  ALA A 130
SHEET    1   D 4 TRP A 154  TRP A 157  0
SHEET    2   D 4 LEU A 164  TRP A 168 -1  O  ALA A 165   N  THR A 156
SHEET    3   D 4 ASP A 171  LYS A 175 -1  O  ASP A 171   N  TRP A 168
SHEET    4   D 4 TYR A 183  ARG A 184 -1  O  TYR A 183   N  VAL A 174
SHEET    1   E 3 ILE A 194  ASN A 196  0
SHEET    2   E 3 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3   E 3 LEU A 214  TRP A 216 -1  N  TRP A 215   O  ALA A 224
SHEET    1   F 4 ILE A 194  ASN A 196  0
SHEET    2   F 4 PHE A 222  ASN A 229 -1  O  PHE A 228   N  TYR A 195
SHEET    3   F 4 THR A 265  ASN A 272 -1  O  PHE A 269   N  TYR A 225
SHEET    4   F 4 ILE A 285  GLN A 286 -1  O  ILE A 285   N  VAL A 270
SHEET    1   G 2 LEU A 235  PHE A 240  0
SHEET    2   G 2 LYS A 250  PRO A 255 -1  O  LYS A 250   N  PHE A 240
SHEET    1   H 4 HIS A 298  TRP A 305  0
SHEET    2   H 4 ARG A 310  ARG A 317 -1  O  SER A 312   N  THR A 304
SHEET    3   H 4 TYR A 322  TYR A 330 -1  O  ASP A 326   N  LEU A 313
SHEET    4   H 4 TRP A 337  ASN A 338 -1  O  ASN A 338   N  ASP A 329
SHEET    1   I 4 HIS A 298  TRP A 305  0
SHEET    2   I 4 ARG A 310  ARG A 317 -1  O  SER A 312   N  THR A 304
SHEET    3   I 4 TYR A 322  TYR A 330 -1  O  ASP A 326   N  LEU A 313
SHEET    4   I 4 HIS A 345  MET A 348 -1  O  HIS A 345   N  MET A 325
SHEET    1   J 4 HIS A 363  PHE A 364  0
SHEET    2   J 4 SER A 370  SER A 376 -1  O  TYR A 372   N  HIS A 363
SHEET    3   J 4 ARG A 382  GLN A 388 -1  O  PHE A 387   N  PHE A 371
SHEET    4   J 4 THR A 395  PHE A 396 -1  O  THR A 395   N  TYR A 386
SHEET    1   K 4 VAL A 404  LEU A 410  0
SHEET    2   K 4 TYR A 414  SER A 419 -1  O  TYR A 416   N  ALA A 409
SHEET    3   K 4 ASN A 430  GLN A 435 -1  O  ILE A 434   N  LEU A 415
SHEET    4   K 4 VAL A 442  CYS A 444 -1  O  THR A 443   N  LYS A 433
SHEET    1   L 4 TYR A 457  PHE A 461  0
SHEET    2   L 4 TYR A 467  CYS A 472 -1  O  GLN A 469   N  SER A 460
SHEET    3   L 4 TYR A 480  SER A 484 -1  O  THR A 481   N  LEU A 470
SHEET    4   L 4 GLY A 490  GLU A 495 -1  O  GLU A 495   N  TYR A 480
SHEET    1   M 8 SER A 511  ILE A 518  0
SHEET    2   M 8 LYS A 523  LEU A 530 -1  O  TYR A 526   N  ASP A 515
SHEET    3   M 8 ILE A 574  PHE A 578 -1  O  VAL A 575   N  ILE A 529
SHEET    4   M 8 TYR A 540  VAL A 546  1  N  ASP A 545   O  ALA A 576
SHEET    5   M 8 VAL A 619  TRP A 629  1  O  TRP A 627   N  LEU A 544
SHEET    6   M 8 CYS A 649  VAL A 653  1  O  VAL A 653   N  GLY A 628
SHEET    7   M 8 GLU A 699  GLY A 705  1  O  ILE A 703   N  ALA A 652
SHEET    8   M 8 GLN A 731  TYR A 735  1  O  GLN A 731   N  TYR A 700
SHEET    1   N 4 LEU B  60  TRP B  62  0
SHEET    2   N 4 GLU B  67  GLN B  72 -1  O  LEU B  69   N  ARG B  61
SHEET    3   N 4 ASN B  75  ASN B  80 -1  O  ASN B  75   N  GLN B  72
SHEET    4   N 4 SER B  86  LEU B  90 -1  O  SER B  87   N  VAL B  78
SHEET    1   O 4 ASP B 104  ILE B 107  0
SHEET    2   O 4 PHE B 113  LYS B 122 -1  O  GLU B 117   N  ASP B 104
SHEET    3   O 4 TYR B 128  ASP B 136 -1  O  SER B 131   N  TYR B 118
SHEET    4   O 4 GLN B 141  LEU B 142 -1  O  GLN B 141   N  ASP B 136
SHEET    1   P 4 TRP B 154  TRP B 157  0
SHEET    2   P 4 LEU B 164  TRP B 168 -1  O  ALA B 165   N  THR B 156
SHEET    3   P 4 ASP B 171  LYS B 175 -1  O  TYR B 173   N  TYR B 166
SHEET    4   P 4 TYR B 183  ARG B 184 -1  O  TYR B 183   N  VAL B 174
SHEET    1   Q 3 ILE B 194  ASN B 196  0
SHEET    2   Q 3 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3   Q 3 LEU B 214  TRP B 216 -1  N  TRP B 215   O  ALA B 224
SHEET    1   R 4 ILE B 194  ASN B 196  0
SHEET    2   R 4 PHE B 222  ASN B 229 -1  O  PHE B 228   N  TYR B 195
SHEET    3   R 4 THR B 265  ASN B 272 -1  O  VAL B 271   N  LEU B 223
SHEET    4   R 4 SER B 284  ILE B 287 -1  O  ILE B 285   N  VAL B 270
SHEET    1   S 2 LEU B 235  PHE B 240  0
SHEET    2   S 2 LYS B 250  PRO B 255 -1  O  LYS B 250   N  PHE B 240
SHEET    1   T 4 HIS B 298  THR B 307  0
SHEET    2   T 4 ARG B 310  ARG B 317 -1  O  SER B 312   N  THR B 304
SHEET    3   T 4 TYR B 322  TYR B 330 -1  O  VAL B 324   N  TRP B 315
SHEET    4   T 4 TRP B 337  CYS B 339 -1  O  ASN B 338   N  ASP B 329
SHEET    1   U 4 HIS B 298  THR B 307  0
SHEET    2   U 4 ARG B 310  ARG B 317 -1  O  SER B 312   N  THR B 304
SHEET    3   U 4 TYR B 322  TYR B 330 -1  O  VAL B 324   N  TRP B 315
SHEET    4   U 4 HIS B 345  MET B 348 -1  O  HIS B 345   N  MET B 325
SHEET    1   V 4 HIS B 363  PHE B 364  0
SHEET    2   V 4 SER B 370  SER B 376 -1  O  TYR B 372   N  HIS B 363
SHEET    3   V 4 ARG B 382  GLN B 388 -1  O  PHE B 387   N  PHE B 371
SHEET    4   V 4 THR B 395  PHE B 396 -1  O  THR B 395   N  TYR B 386
SHEET    1   W 4 VAL B 404  LEU B 410  0
SHEET    2   W 4 TYR B 414  SER B 419 -1  O  ILE B 418   N  ILE B 405
SHEET    3   W 4 ASN B 430  GLN B 435 -1  O  TYR B 432   N  TYR B 417
SHEET    4   W 4 VAL B 442  CYS B 444 -1  O  THR B 443   N  LYS B 433
SHEET    1   X 4 TYR B 457  PHE B 461  0
SHEET    2   X 4 TYR B 467  CYS B 472 -1  O  GLN B 469   N  SER B 460
SHEET    3   X 4 LEU B 479  SER B 484 -1  O  HIS B 483   N  TYR B 468
SHEET    4   X 4 GLY B 490  GLU B 495 -1  O  GLU B 495   N  TYR B 480
SHEET    1   Y 8 SER B 511  LEU B 519  0
SHEET    2   Y 8 THR B 522  LEU B 530 -1  O  MET B 528   N  LYS B 513
SHEET    3   Y 8 ILE B 574  PHE B 578 -1  O  VAL B 575   N  ILE B 529
SHEET    4   Y 8 TYR B 540  VAL B 546  1  N  LEU B 543   O  ALA B 576
SHEET    5   Y 8 VAL B 619  TRP B 629  1  O  ALA B 625   N  LEU B 542
SHEET    6   Y 8 CYS B 649  VAL B 653  1  O  VAL B 653   N  GLY B 628
SHEET    7   Y 8 GLU B 699  GLY B 705  1  O  ILE B 703   N  ALA B 652
SHEET    8   Y 8 GLN B 731  TYR B 735  1  O  GLN B 731   N  TYR B 700
SSBOND   1 CYS A  328    CYS A  339                          1555   1555  2.04
SSBOND   2 CYS A  385    CYS A  394                          1555   1555  2.01
SSBOND   3 CYS A  444    CYS A  447                          1555   1555  2.07
SSBOND   4 CYS A  454    CYS A  472                          1555   1555  2.05
SSBOND   5 CYS A  649    CYS A  762                          1555   1555  2.04
SSBOND   6 CYS B  328    CYS B  339                          1555   1555  2.05
SSBOND   7 CYS B  385    CYS B  394                          1555   1555  2.04
SSBOND   8 CYS B  444    CYS B  447                          1555   1555  2.02
SSBOND   9 CYS B  454    CYS B  472                          1555   1555  2.08
SSBOND  10 CYS B  649    CYS B  762                          1555   1555  2.04
LINK         ND2 ASN A 321                 C1  NAG A 806     1555   1555  1.43
LINK         ND2 ASN B 150                 C1  NAG B 802     1555   1555  1.44
LINK         ND2 ASN A 229                 C1  NAG A 804     1555   1555  1.45
LINK         ND2 ASN A  85                 C1  NAG A 801     1555   1555  1.45
LINK         ND2 ASN A 281                 C1  NAG A 805     1555   1555  1.45
LINK         ND2 ASN B 219                 C1  NAG B 803     1555   1555  1.45
LINK         ND2 ASN B  85                 C1  NAG B 801     1555   1555  1.45
LINK         ND2 ASN B 321                 C1  NAG B 806     1555   1555  1.45
LINK         ND2 ASN A 219                 C1  NAG A 803     1555   1555  1.45
LINK         ND2 ASN A 150                 C1  NAG A 802     1555   1555  1.45
LINK         ND2 ASN B 229                 C1  NAG B 804     1555   1555  1.45
LINK         ND2 ASN B 281                 C1  NAG B 805     1555   1555  1.46
CISPEP   1 GLY A  474    PRO A  475          0         8.14
CISPEP   2 GLY B  474    PRO B  475          0        12.70
SITE     1 AC1  3 NAG A 804  SER B 690  ASN B 694
SITE     1 AC2 12 ARG A 125  GLU A 205  GLU A 206  PHE A 357
SITE     2 AC2 12 TYR A 547  TYR A 585  SER A 630  TYR A 631
SITE     3 AC2 12 TYR A 662  TYR A 666  ASN A 710  HOH A 913
SITE     1 AC3 11 GLU B 205  GLU B 206  PHE B 357  ARG B 358
SITE     2 AC3 11 TYR B 547  TYR B 585  SER B 630  TYR B 631
SITE     3 AC3 11 TYR B 662  TYR B 666  ASN B 710
SITE     1 AC4  6 VAL A  78  ASN A  85  SER A  86  SER A  87
SITE     2 AC4  6 GLN A 388  THR A 395
SITE     1 AC5  4 ASN A 119  ASN A 150  ILE A 518  HOH A 941
SITE     1 AC6  3 ASN A 219  THR A 221  GLU A 309
SITE     1 AC7  6 ILE A 194  ASN A 229  THR A 231  GLU A 232
SITE     2 AC7  6 LYS A 267  NAG A 807
SITE     1 AC8  2 TRP A 187  ASN A 281
SITE     1 AC9  2 ASN A 321  ARG A 596
SITE     1 BC1  6 VAL B  78  ASN B  85  SER B  86  SER B  87
SITE     2 BC1  6 GLN B 388  THR B 395
SITE     1 BC2  8 HIS B 100  TYR B 118  SER B 131  TYR B 132
SITE     2 BC2  8 ASP B 133  ARG B 147  ILE B 148  ASN B 150
SITE     1 BC3  5 ASN B 219  THR B 221  GLN B 308  GLU B 309
SITE     2 BC3  5 HOH B1026
SITE     1 BC4  4 ASN B 229  THR B 231  GLU B 232  LYS B 267
SITE     1 BC5  3 TRP B 187  VAL B 279  ASN B 281
SITE     1 BC6  3 ASN B 321  MET B 348  ARG B 596
CRYST1   65.160   68.990  419.600  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.015347  0.000000  0.000000        0.00000
SCALE2      0.000000  0.014495  0.000000        0.00000
SCALE3      0.000000  0.000000  0.002383        0.00000
TER    5964      PRO A 766
TER   11928      PRO B 766
MASTER      424    0   15   37  102    0   24    612269    2  271  114
END