longtext: 4ARA-pdb

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HEADER    HYDROLASE                               23-APR-12   4ARA
TITLE     MUS MUSCULUS ACETYLCHOLINESTERASE IN COMPLEX WITH (R)-C5685 AT 2.5 A
TITLE    2 RESOLUTION.
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ACETYLCHOLINESTERASE;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: CATALYTIC DOMAIN, RESIDUES 32-574;
COMPND   5 SYNONYM: ACHE;
COMPND   6 EC: 3.1.1.7;
COMPND   7 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MUS MUSCULUS;
SOURCE   3 ORGANISM_COMMON: HOUSE MOUSE;
SOURCE   4 ORGANISM_TAXID: 10090;
SOURCE   5 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE   6 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE   8 EXPRESSION_SYSTEM_VARIANT: HEK293F
KEYWDS    HYDROLASE, ENATIOMERS, INHIBITOR, CHEMICAL LEAD
EXPDTA    X-RAY DIFFRACTION
AUTHOR    L.BERG,M.S.NIEMIEC,W.QIAN,C.D.ANDERSSON,P.WITTUNGSTAFSHEDE,F.EKSTROM,
AUTHOR   2 A.LINUSSON
REVDAT   1   28-NOV-12 4ARA    0
JRNL        AUTH   L.BERG,M.S.NIEMIEC,W.QIAN,C.D.ANDERSSON,P.WITTUNG-STAFSHEDE,
JRNL        AUTH 2 F.EKSTROM,A.LINUSSON
JRNL        TITL   SIMILAR BUT DIFFERENT: THERMODYNAMIC AND STRUCTURAL
JRNL        TITL 2 CHARACTERIZATION OF A PAIR OF ENANTIOMERS BINDING TO
JRNL        TITL 3 ACETYLCHOLINESTERASE.
JRNL        REF    ANGEW.CHEM.INT.ED.ENGL.                    2012
JRNL        REFN                   ESSN 1521-3773
JRNL        PMID   23161758
JRNL        DOI    10.1002/ANIE.201205113
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   L.BERG,C.D.ANDERSSON,E.ARTURSSON,A.HORNBERG,A.TUNEMALM,
REMARK   1  AUTH 2 A.LINUSSON,F.EKSTROM
REMARK   1  TITL   TARGETING ACETYLCHOLINESTERASE: IDENTIFICATION OF CHEMICAL
REMARK   1  TITL 2 LEADS BY HIGH THROUGHPUT SCREENING, STRUCTURE DETERMINATION
REMARK   1  TITL 3 AND MOLECULAR MODELING.
REMARK   1  REF    PLOS ONE                      V.   6 26039 2011
REMARK   1  REFN                   ISSN 1932-6203
REMARK   1  PMID   22140425
REMARK   1  DOI    10.1371/JOURNAL.PONE.0026039
REMARK   2
REMARK   2 RESOLUTION.    2.50 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VICENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.50
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 28.855
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.34
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.79
REMARK   3   NUMBER OF REFLECTIONS             : 70252
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.1834
REMARK   3   R VALUE            (WORKING SET) : 0.1826
REMARK   3   FREE R VALUE                     : 0.2220
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 2.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 1416
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 28.8570 -  5.3757    0.99     7189   143  0.1885 0.2164
REMARK   3     2  5.3757 -  4.2713    1.00     6985   132  0.1405 0.1623
REMARK   3     3  4.2713 -  3.7327    1.00     6914   140  0.1496 0.1868
REMARK   3     4  3.7327 -  3.3920    1.00     6867   141  0.1836 0.2211
REMARK   3     5  3.3920 -  3.1492    1.00     6883   130  0.2051 0.2611
REMARK   3     6  3.1492 -  2.9637    1.00     6812   152  0.2038 0.2251
REMARK   3     7  2.9637 -  2.8154    1.00     6822   136  0.2017 0.2763
REMARK   3     8  2.8154 -  2.6929    1.00     6819   134  0.2188 0.2521
REMARK   3     9  2.6929 -  2.5893    1.00     6773   154  0.2283 0.2913
REMARK   3    10  2.5893 -  2.5000    1.00     6772   154  0.2516 0.3072
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.00
REMARK   3   SHRINKAGE RADIUS   : 0.73
REMARK   3   K_SOL              : 0.346
REMARK   3   B_SOL              : 60.440
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.25
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 22.96
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 39.74
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 10.8219
REMARK   3    B22 (A**2) : 12.8430
REMARK   3    B33 (A**2) : -23.6649
REMARK   3    B12 (A**2) : 0.0000
REMARK   3    B13 (A**2) : 0.0000
REMARK   3    B23 (A**2) : 0.0000
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.004           8837
REMARK   3   ANGLE     :  0.898          12028
REMARK   3   CHIRALITY :  0.062           1284
REMARK   3   PLANARITY :  0.005           1569
REMARK   3   DIHEDRAL  : 15.764           3242
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 17
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 1:142)
REMARK   3    ORIGIN FOR THE GROUP (A):  30.3331  15.4227  32.1785
REMARK   3    T TENSOR
REMARK   3      T11:   0.2680 T22:   0.2377
REMARK   3      T33:   0.2733 T12:   0.0043
REMARK   3      T13:  -0.0062 T23:   0.0216
REMARK   3    L TENSOR
REMARK   3      L11:   0.1653 L22:   0.1515
REMARK   3      L33:   1.1039 L12:  -0.0646
REMARK   3      L13:  -0.0023 L23:  -0.0362
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0523 S12:  -0.0982 S13:   0.0348
REMARK   3      S21:   0.1247 S22:   0.0154 S23:  -0.0733
REMARK   3      S31:  -0.0659 S32:   0.0513 S33:  -0.0000
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 143:223)
REMARK   3    ORIGIN FOR THE GROUP (A):  33.2478   4.1202  24.0344
REMARK   3    T TENSOR
REMARK   3      T11:   0.2888 T22:   0.1961
REMARK   3      T33:   0.2787 T12:   0.0377
REMARK   3      T13:  -0.0043 T23:   0.0419
REMARK   3    L TENSOR
REMARK   3      L11:   0.1483 L22:   0.5967
REMARK   3      L33:   0.8150 L12:   0.2277
REMARK   3      L13:   0.3633 L23:   0.1429
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0073 S12:  -0.0490 S13:  -0.0278
REMARK   3      S21:   0.1101 S22:   0.0576 S23:  -0.0461
REMARK   3      S31:   0.1922 S32:   0.0234 S33:   0.0000
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 224:341)
REMARK   3    ORIGIN FOR THE GROUP (A):  37.7919  10.9019  10.3854
REMARK   3    T TENSOR
REMARK   3      T11:   0.1622 T22:   0.1245
REMARK   3      T33:   0.2217 T12:   0.0429
REMARK   3      T13:   0.0319 T23:   0.0192
REMARK   3    L TENSOR
REMARK   3      L11:   0.8001 L22:   0.0000
REMARK   3      L33:   1.3220 L12:   0.3229
REMARK   3      L13:   0.2284 L23:  -0.3981
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0884 S12:   0.1377 S13:   0.0056
REMARK   3      S21:  -0.0313 S22:   0.0118 S23:  -0.0087
REMARK   3      S31:   0.1742 S32:   0.1599 S33:  -0.0002
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 342:486)
REMARK   3    ORIGIN FOR THE GROUP (A):  18.5229  17.1741   6.3422
REMARK   3    T TENSOR
REMARK   3      T11:   0.1834 T22:   0.2424
REMARK   3      T33:   0.2619 T12:  -0.0244
REMARK   3      T13:  -0.0065 T23:   0.0320
REMARK   3    L TENSOR
REMARK   3      L11:   0.2268 L22:   0.5033
REMARK   3      L33:   1.4203 L12:  -0.0797
REMARK   3      L13:   0.3579 L23:   0.1415
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0342 S12:   0.1124 S13:   0.0048
REMARK   3      S21:  -0.1121 S22:   0.0327 S23:   0.0835
REMARK   3      S31:   0.0444 S32:  -0.2615 S33:  -0.0000
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 487:513)
REMARK   3    ORIGIN FOR THE GROUP (A):   6.8700   1.4649  13.7085
REMARK   3    T TENSOR
REMARK   3      T11:   0.3515 T22:   0.4830
REMARK   3      T33:   0.4191 T12:  -0.2953
REMARK   3      T13:   0.0134 T23:   0.0765
REMARK   3    L TENSOR
REMARK   3      L11:   0.2896 L22:   0.3384
REMARK   3      L33:   0.8203 L12:  -0.2812
REMARK   3      L13:  -0.1398 L23:   0.0446
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0970 S12:   0.4955 S13:  -0.1487
REMARK   3      S21:  -0.0701 S22:  -0.0042 S23:   0.5685
REMARK   3      S31:   0.3724 S32:  -1.0654 S33:   0.0012
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 514:542)
REMARK   3    ORIGIN FOR THE GROUP (A):  17.6550   6.3933  -1.1430
REMARK   3    T TENSOR
REMARK   3      T11:   0.2660 T22:   0.4620
REMARK   3      T33:   0.2711 T12:  -0.0665
REMARK   3      T13:  -0.0476 T23:  -0.0164
REMARK   3    L TENSOR
REMARK   3      L11:   0.0313 L22:   0.0090
REMARK   3      L33:   0.6534 L12:  -0.0093
REMARK   3      L13:   0.1462 L23:  -0.0442
REMARK   3    S TENSOR
REMARK   3      S11:   0.0674 S12:  -0.1576 S13:   0.0765
REMARK   3      S21:  -0.3743 S22:  -0.0490 S23:   0.1712
REMARK   3      S31:   0.2459 S32:   0.2076 S33:   0.0094
REMARK   3   TLS GROUP : 7
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 4:45)
REMARK   3    ORIGIN FOR THE GROUP (A):  -3.6357   6.1103 -61.5447
REMARK   3    T TENSOR
REMARK   3      T11:   0.2640 T22:   0.4445
REMARK   3      T33:   0.3153 T12:   0.0732
REMARK   3      T13:  -0.0824 T23:  -0.0849
REMARK   3    L TENSOR
REMARK   3      L11:   0.1856 L22:   0.7024
REMARK   3      L33:   0.9043 L12:  -0.1066
REMARK   3      L13:   0.2767 L23:  -0.0168
REMARK   3    S TENSOR
REMARK   3      S11:   0.0747 S12:   0.3060 S13:   0.0863
REMARK   3      S21:  -0.3115 S22:  -0.1366 S23:   0.0126
REMARK   3      S31:  -0.1399 S32:  -0.3327 S33:  -0.0008
REMARK   3   TLS GROUP : 8
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 46:86)
REMARK   3    ORIGIN FOR THE GROUP (A):   5.2890  -4.2856 -52.9246
REMARK   3    T TENSOR
REMARK   3      T11:   0.3597 T22:   0.3593
REMARK   3      T33:   0.3143 T12:  -0.0442
REMARK   3      T13:  -0.0313 T23:  -0.0629
REMARK   3    L TENSOR
REMARK   3      L11:  -0.0538 L22:   0.5770
REMARK   3      L33:   0.5359 L12:  -0.0542
REMARK   3      L13:  -0.1648 L23:   0.4534
REMARK   3    S TENSOR
REMARK   3      S11:   0.0781 S12:   0.2434 S13:  -0.0439
REMARK   3      S21:  -0.1155 S22:  -0.1693 S23:   0.0386
REMARK   3      S31:   0.5495 S32:   0.0695 S33:  -0.0074
REMARK   3   TLS GROUP : 9
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 87:142)
REMARK   3    ORIGIN FOR THE GROUP (A):  -1.3090   4.6399 -50.3098
REMARK   3    T TENSOR
REMARK   3      T11:   0.1961 T22:   0.3488
REMARK   3      T33:   0.2721 T12:   0.0221
REMARK   3      T13:  -0.0882 T23:  -0.0881
REMARK   3    L TENSOR
REMARK   3      L11:   0.5536 L22:   0.1622
REMARK   3      L33:   0.6006 L12:   0.3230
REMARK   3      L13:   0.2268 L23:   0.1461
REMARK   3    S TENSOR
REMARK   3      S11:   0.0392 S12:   0.1752 S13:  -0.1582
REMARK   3      S21:  -0.2557 S22:  -0.2172 S23:   0.0825
REMARK   3      S31:   0.3045 S32:  -0.4991 S33:  -0.0120
REMARK   3   TLS GROUP : 10
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 143:190)
REMARK   3    ORIGIN FOR THE GROUP (A):  12.3105   6.7368 -54.0317
REMARK   3    T TENSOR
REMARK   3      T11:   0.3099 T22:   0.3635
REMARK   3      T33:   0.2861 T12:   0.0174
REMARK   3      T13:  -0.0322 T23:  -0.0648
REMARK   3    L TENSOR
REMARK   3      L11:   0.1282 L22:   0.1281
REMARK   3      L33:   0.7286 L12:   0.1347
REMARK   3      L13:  -0.0386 L23:  -0.0379
REMARK   3    S TENSOR
REMARK   3      S11:   0.1592 S12:   0.3240 S13:  -0.1130
REMARK   3      S21:  -0.2351 S22:  -0.0886 S23:   0.0934
REMARK   3      S31:  -0.1263 S32:   0.2206 S33:   0.0001
REMARK   3   TLS GROUP : 11
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 191:255)
REMARK   3    ORIGIN FOR THE GROUP (A):  15.1798   8.2124 -44.1651
REMARK   3    T TENSOR
REMARK   3      T11:   0.2267 T22:   0.3075
REMARK   3      T33:   0.2643 T12:   0.0043
REMARK   3      T13:  -0.0305 T23:  -0.0326
REMARK   3    L TENSOR
REMARK   3      L11:   0.0075 L22:   0.2716
REMARK   3      L33:   0.9570 L12:  -0.1866
REMARK   3      L13:   0.0636 L23:   0.2967
REMARK   3    S TENSOR
REMARK   3      S11:   0.0677 S12:  -0.0016 S13:  -0.0485
REMARK   3      S21:   0.0236 S22:   0.0025 S23:  -0.0339
REMARK   3      S31:  -0.0771 S32:   0.2105 S33:   0.0001
REMARK   3   TLS GROUP : 12
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 256:297)
REMARK   3    ORIGIN FOR THE GROUP (A):  22.4756  -9.8867 -46.8130
REMARK   3    T TENSOR
REMARK   3      T11:   0.4227 T22:   0.3671
REMARK   3      T33:   0.3215 T12:   0.1473
REMARK   3      T13:  -0.0921 T23:  -0.0778
REMARK   3    L TENSOR
REMARK   3      L11:   0.2412 L22:   0.2535
REMARK   3      L33:   0.1356 L12:  -0.2569
REMARK   3      L13:   0.1066 L23:   0.0365
REMARK   3    S TENSOR
REMARK   3      S11:   0.0622 S12:   0.0922 S13:  -0.1681
REMARK   3      S21:   0.0286 S22:  -0.0414 S23:  -0.0218
REMARK   3      S31:   0.3512 S32:   0.5162 S33:  -0.0001
REMARK   3   TLS GROUP : 13
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 298:331)
REMARK   3    ORIGIN FOR THE GROUP (A):  18.2876  14.2963 -40.1912
REMARK   3    T TENSOR
REMARK   3      T11:   0.1739 T22:   0.2684
REMARK   3      T33:   0.2728 T12:  -0.0455
REMARK   3      T13:  -0.0219 T23:  -0.0379
REMARK   3    L TENSOR
REMARK   3      L11:   0.4274 L22:   0.3005
REMARK   3      L33:   0.7741 L12:  -0.2992
REMARK   3      L13:   0.0964 L23:   0.2516
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1014 S12:   0.2486 S13:   0.1285
REMARK   3      S21:   0.0067 S22:   0.1129 S23:  -0.0960
REMARK   3      S31:   0.0292 S32:   0.3509 S33:  -0.0001
REMARK   3   TLS GROUP : 14
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 332:382)
REMARK   3    ORIGIN FOR THE GROUP (A):  13.8485  -6.1946 -22.0912
REMARK   3    T TENSOR
REMARK   3      T11:   0.5538 T22:   0.2004
REMARK   3      T33:   0.3377 T12:   0.0191
REMARK   3      T13:  -0.0361 T23:  -0.0032
REMARK   3    L TENSOR
REMARK   3      L11:   0.1683 L22:   0.3401
REMARK   3      L33:   0.2821 L12:   0.0680
REMARK   3      L13:  -0.0702 L23:   0.0906
REMARK   3    S TENSOR
REMARK   3      S11:   0.1290 S12:  -0.1209 S13:   0.0046
REMARK   3      S21:   0.2211 S22:   0.0258 S23:  -0.0245
REMARK   3      S31:   0.5619 S32:   0.2245 S33:   0.0006
REMARK   3   TLS GROUP : 15
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 383:486)
REMARK   3    ORIGIN FOR THE GROUP (A):   0.7352   5.8360 -29.0308
REMARK   3    T TENSOR
REMARK   3      T11:   0.1865 T22:   0.2232
REMARK   3      T33:   0.2259 T12:  -0.0454
REMARK   3      T13:  -0.0048 T23:  -0.0358
REMARK   3    L TENSOR
REMARK   3      L11:   0.1448 L22:   0.6458
REMARK   3      L33:   1.4030 L12:   0.3314
REMARK   3      L13:  -0.1232 L23:   0.3301
REMARK   3    S TENSOR
REMARK   3      S11:   0.1272 S12:  -0.0905 S13:   0.0418
REMARK   3      S21:   0.1870 S22:  -0.1625 S23:   0.0317
REMARK   3      S31:   0.0814 S32:  -0.1277 S33:  -0.0000
REMARK   3   TLS GROUP : 16
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 487:513)
REMARK   3    ORIGIN FOR THE GROUP (A):  -1.0503  22.4945 -28.6338
REMARK   3    T TENSOR
REMARK   3      T11:   0.3733 T22:   0.3490
REMARK   3      T33:   0.3594 T12:   0.0523
REMARK   3      T13:  -0.0191 T23:  -0.0593
REMARK   3    L TENSOR
REMARK   3      L11:   0.1012 L22:   0.1330
REMARK   3      L33:   0.3725 L12:  -0.0023
REMARK   3      L13:  -0.1959 L23:   0.0712
REMARK   3    S TENSOR
REMARK   3      S11:   0.0429 S12:  -0.5460 S13:   0.2598
REMARK   3      S21:   0.0021 S22:  -0.0191 S23:   0.3205
REMARK   3      S31:  -0.3827 S32:  -0.4171 S33:   0.0010
REMARK   3   TLS GROUP : 17
REMARK   3    SELECTION: CHAIN B AND (RESSEQ 514:543)
REMARK   3    ORIGIN FOR THE GROUP (A):  13.3573  11.4043 -21.5981
REMARK   3    T TENSOR
REMARK   3      T11:   0.3900 T22:   0.4073
REMARK   3      T33:   0.2502 T12:  -0.0125
REMARK   3      T13:  -0.0043 T23:  -0.0539
REMARK   3    L TENSOR
REMARK   3      L11:  -0.0023 L22:  -0.0104
REMARK   3      L33:   0.4414 L12:  -0.0212
REMARK   3      L13:   0.0037 L23:  -0.0913
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0336 S12:   0.0358 S13:  -0.0356
REMARK   3      S21:   0.1517 S22:  -0.0224 S23:   0.1543
REMARK   3      S31:   0.0937 S32:   0.2484 S33:  -0.0000
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4ARA COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 23-APR-12.
REMARK 100 THE PDBE ID CODE IS EBI-52170.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 14-FEB-12
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : MAX II
REMARK 200  BEAMLINE                       : I911-5
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.0
REMARK 200  MONOCHROMATOR                  : RH-COATED SI MIRROR, BENT FOR
REMARK 200                                   VERTICAL COLLIMATION
REMARK 200  OPTICS                         : MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD (MX-225)
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 70472
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.50
REMARK 200  RESOLUTION RANGE LOW       (A) : 28.98
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 5.4
REMARK 200  R MERGE                    (I) : 0.08
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 17.00
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.50
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.64
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.4
REMARK 200  R MERGE FOR SHELL          (I) : 0.56
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 4.00
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: REFMAC
REMARK 200 STARTING MODEL: PDB ENTRY 1J06
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 65
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.6
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M HEPES, 30 % (V/V)
REMARK 280  POLYETHELENEGLYCOLEMONOMETHYLETHER, PH 7
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       39.38200
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      113.67000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       56.00600
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      113.67000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       39.38200
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       56.00600
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 6440 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 38240 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -14.2 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     PRO A   258
REMARK 465     PRO A   259
REMARK 465     GLY A   260
REMARK 465     GLY A   261
REMARK 465     ALA A   262
REMARK 465     GLY A   263
REMARK 465     GLY A   264
REMARK 465     THR A   543
REMARK 465     ALA A   544
REMARK 465     THR A   545
REMARK 465     GLU A   546
REMARK 465     ALA A   547
REMARK 465     PRO A   548
REMARK 465     GLU B     1
REMARK 465     GLY B     2
REMARK 465     ARG B     3
REMARK 465     PRO B   258
REMARK 465     PRO B   259
REMARK 465     GLY B   260
REMARK 465     GLY B   261
REMARK 465     ALA B   262
REMARK 465     GLY B   263
REMARK 465     GLY B   264
REMARK 465     ALA B   544
REMARK 465     THR B   545
REMARK 465     GLU B   546
REMARK 465     ALA B   547
REMARK 465     PRO B   548
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ARG A   3    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS A  23    CG   CD   CE   NZ
REMARK 470     GLU A  81    CG   CD   OE1  OE2
REMARK 470     GLU A 268    CG   CD   OE1  OE2
REMARK 470     ARG A 493    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS A 496    CG   CD   CE   NZ
REMARK 470     GLU B   4    CG   CD   OE1  OE2
REMARK 470     ARG B 107    CG   CD   NE   CZ   NH1  NH2
REMARK 470     GLU B 268    CG   CD   OE1  OE2
REMARK 470     ARG B 493    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS B 496    CG   CD   CE   NZ
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OE2  GLU A   285     O    HOH A  2223              2.19
REMARK 500   NH1  ARG B   274     O    HOH B  2037              2.19
REMARK 500   ND2  ASN B   350     O5   NAG B  1552              1.32
REMARK 500   OH7  1PE B  1546     O    HOH B  2266              2.15
REMARK 500   O    HOH B  2026     O    HOH B  2089              2.11
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   NH1  ARG A    13     O    SER B    57     2555     2.16
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PHE A  47      -10.11     71.89
REMARK 500    ALA A  62       52.11   -119.48
REMARK 500    ALA A 167       72.36   -155.87
REMARK 500    SER A 203     -118.29     59.00
REMARK 500    PRO A 217        2.48    -67.65
REMARK 500    ASP A 306      -85.30   -133.61
REMARK 500    HIS A 387       57.31   -143.74
REMARK 500    VAL A 407      -66.15   -126.45
REMARK 500    ARG A 493       50.43   -115.59
REMARK 500    ASP A 494       91.17   -160.67
REMARK 500    SER A 541       46.35    -89.92
REMARK 500    ALA B  62       52.18   -118.38
REMARK 500    ALA B 167       70.08   -150.67
REMARK 500    SER B 203     -119.33     55.59
REMARK 500    ASP B 306      -78.00   -121.57
REMARK 500    ASN B 350     -167.19   -128.83
REMARK 500    VAL B 407      -65.39   -127.24
REMARK 500    SER B 497      125.90     65.05
REMARK 500    SER B 541       31.90    -82.58
REMARK 500    ALA B 542      -13.97   -163.55
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    TYR B 337        24.9      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 600
REMARK 600 HETEROGEN
REMARK 600 4-(DIMETHYLAMINO)-N-{[(2S)-1-ETHYLPYRROLIDIN-
REMARK 600  2-YL]METHYL}-2-METHOXY-5-NITROBENZAMIDE (568):
REMARK 600  CORRESPONDING TO LIGAND C56 IN THE RACEMIC STRUCTURE 4A23
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE C56 A 600
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A1544
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1545
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A1546
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG A1547
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1548
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1550
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1551
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A1552
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE C56 B 600
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B1545
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE 1PE B1546
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B1547
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B1548
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO B1549
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B1550
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B1551
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO B1553
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B1554
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL B1555
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO B1556
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800   NAG A1549 BOUND TO ASN A 350
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800   NAG A1553 BOUND TO ASN A 464
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800   NAG B1552 BOUND TO ASN B 350
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 1C2B   RELATED DB: PDB
REMARK 900  ELECTROPHORUS ELECTRICUS ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1C2O   RELATED DB: PDB
REMARK 900  ELECTROPHORUS ELECTRICUS ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 1J06   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE IN THE
REMARK 900  APOFORM
REMARK 900 RELATED ID: 1J07   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MOUSE ACETYLCHOLINESTERASE-
REMARK 900  DECIDIUM COMPLEX
REMARK 900 RELATED ID: 1KU6   RELATED DB: PDB
REMARK 900  FASCICULIN 2-MOUSE ACETYLCHOLINESTERASE COMPLEX
REMARK 900 RELATED ID: 1MAA   RELATED DB: PDB
REMARK 900  MOUSE ACETYLCHOLINESTERASE CATALYTIC DOMAIN,
REMARK 900  GLYCOSYLATEDPROTEIN
REMARK 900 RELATED ID: 1MAH   RELATED DB: PDB
REMARK 900  FASCICULIN2 - MOUSE ACETYLCHOLINESTERASE COMPLEX
REMARK 900 RELATED ID: 1N5M   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MOUSE ACETYLCHOLINESTERASE-
REMARK 900  GALLAMINE COMPLEX
REMARK 900 RELATED ID: 1N5R   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MOUSE ACETYLCHOLINESTERASE-
REMARK 900  PROPIDIUM COMPLEX
REMARK 900 RELATED ID: 1Q83   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MOUSE ACETYLCHOLINESTERASE-
REMARK 900  TZ2PA6SYN COMPLEX
REMARK 900 RELATED ID: 1Q84   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MOUSE ACETYLCHOLINESTERASE-
REMARK 900  TZ2PA6ANTI COMPLEX
REMARK 900 RELATED ID: 2C0P   RELATED DB: PDB
REMARK 900  AGED FORM OF MOUSE ACETYLCHOLINESTERASE INHIBITED BY
REMARK 900  TABUN
REMARK 900 RELATED ID: 2C0Q   RELATED DB: PDB
REMARK 900  NON-AGED FORM OF MOUSE ACETYLCHOLINESTERASE INHIBITED BY
REMARK 900   TABUN
REMARK 900 RELATED ID: 2H9Y   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE
REMARK 900  COMPLEXEDWITH M-(N,N,N-TRIMETHYLAMMONIO)
REMARK 900  TRIFLUOROACETOPHENONE
REMARK 900 RELATED ID: 2HA0   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE
REMARK 900  COMPLEXEDWITH 4-KETOAMYLTRIMETHYLAMMONIUM
REMARK 900 RELATED ID: 2HA2   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE
REMARK 900  COMPLEXEDWITH SUCCINYLCHOLINE
REMARK 900 RELATED ID: 2HA3   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE
REMARK 900  COMPLEXEDWITH CHOLINE
REMARK 900 RELATED ID: 2HA4   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MUTANT S203A OF
REMARK 900  MOUSEACETYLCHOLINESTERASE COMPLEXED WITH ACETYLCHOLINE
REMARK 900 RELATED ID: 2HA5   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MUTANT S203A OF
REMARK 900  ACETYLCHOLINESTERASECOMPLEXED WITH ACETYLTHIOCHOLINE
REMARK 900 RELATED ID: 2HA6   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MUTANT S203A OF
REMARK 900  MOUSEACETYLCHOLINESTERASE COMPLEXED WITH SUCCINYLCHOLINE
REMARK 900 RELATED ID: 2HA7   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MUTANT S203A OF
REMARK 900  MOUSEACETYLCHOLINESTERASE COMPLEXED WITH BUTYRYLTHIOCHOLINE
REMARK 900 RELATED ID: 2JEY   RELATED DB: PDB
REMARK 900  MUS MUSCULUS ACETYLCHOLINESTERASE IN COMPLEX WITH HLO-7
REMARK 900 RELATED ID: 2JEZ   RELATED DB: PDB
REMARK 900  MUS MUSCULUS ACETYLCHOLINESTERASE IN COMPLEX WITH TABUN
REMARK 900  AND HLO-7
REMARK 900 RELATED ID: 2JF0   RELATED DB: PDB
REMARK 900  MUS MUSCULUS ACETYLCHOLINESTERASE IN COMPLEX WITH TABUN
REMARK 900  AND ORTHO-7
REMARK 900 RELATED ID: 2JGE   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY NON-AGED METHAMIDOPHOS
REMARK 900 RELATED ID: 2JGF   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY NON-AGED FENAMIPHOS
REMARK 900 RELATED ID: 2JGG   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY NON-AGED SARIN
REMARK 900 RELATED ID: 2JGH   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY NON-AGED VX
REMARK 900 RELATED ID: 2JGI   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY NON-AGED DIISOPROPYL FLUOROPHOSPHATE (DFP)
REMARK 900 RELATED ID: 2JGJ   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY AGED METHAMIDOPHOS
REMARK 900 RELATED ID: 2JGK   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY AGED FENAMIPHOS
REMARK 900 RELATED ID: 2JGL   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY AGED VX AND SARIN
REMARK 900 RELATED ID: 2JGM   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE INHIBITED
REMARK 900  BY AGED DIISOPROPYL FLUOROPHOSPHATE (DFP)
REMARK 900 RELATED ID: 2WHP   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF ACETYLCHOLINESTERASE, PHOSPHONYLATED BY
REMARK 900   SARIN AND IN COMPLEX WITH HI-6
REMARK 900 RELATED ID: 2WHQ   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF ACETYLCHOLINESTERASE, PHOSPHONYLATED BY
REMARK 900   SARIN (AGED) IN COMPLEX WITH HI-6
REMARK 900 RELATED ID: 2WHR   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF ACETYLCHOLINESTERASE IN COMPLEX WITH
REMARK 900   K027
REMARK 900 RELATED ID: 2WLS   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MUS MUSCULUS ACETYLCHOLINESTERASE IN
REMARK 900   COMPLEX WITH AMTS13
REMARK 900 RELATED ID: 2WU3   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE IN COMPLEX
REMARK 900   WITH FENAMIPHOS AND HI-6
REMARK 900 RELATED ID: 2WU4   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MOUSE ACETYLCHOLINESTERASE IN COMPLEX
REMARK 900   WITH FENAMIPHOS AND ORTHO-7
REMARK 900 RELATED ID: 2XUD   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE Y337A MUTANT OF MOUSE
REMARK 900  ACETYLCHOLINESTERASE
REMARK 900 RELATED ID: 2XUF   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A-TZ2PA6 ANTI COMPLEX (
REMARK 900  1 MTH)
REMARK 900 RELATED ID: 2XUG   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A-TZ2PA6 ANTI COMPLEX (
REMARK 900  1 WK)
REMARK 900 RELATED ID: 2XUH   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A-TZ2PA6 ANTI COMPLEX (
REMARK 900  10 MTH)
REMARK 900 RELATED ID: 2XUI   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A-TZ2PA6 SYN COMPLEX (1
REMARK 900   WK)
REMARK 900 RELATED ID: 2XUJ   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A-TZ2PA6 SYN COMPLEX (1
REMARK 900   MTH)
REMARK 900 RELATED ID: 2XUK   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A-TZ2PA6 SYN COMPLEX (
REMARK 900  10 MTH)
REMARK 900 RELATED ID: 2XUO   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF MACHE-Y337A MUTANT IN COMPLEX
REMARK 900  WITH SOAKED TZ2PA6 ANTI INHIBITOR
REMARK 900 RELATED ID: 2XUP   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MACHE-Y337A MUTANT IN COMPLEX
REMARK 900   WITH SOAKED TZ2PA6 SYN INHIBITOR
REMARK 900 RELATED ID: 2XUQ   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF THE MACHE-Y337A MUTANT IN COMPLEX
REMARK 900   WITH SOAKED TZ2PA6 ANTI-SYN INHIBITORS
REMARK 900 RELATED ID: 4A16   RELATED DB: PDB
REMARK 900  STRUCTURE OF MOUSE ACETYLCHOLINESTERASE COMPLEX WITH
REMARK 900  HUPRINE DERIVATIVE
REMARK 900 RELATED ID: 4A23   RELATED DB: PDB
REMARK 900  MUS MUSCULUS ACETYLCHOLINESTERASE IN COMPLEX WITH RACEMIC
REMARK 900   C5685
REMARK 900 RELATED ID: 4ARB   RELATED DB: PDB
REMARK 900  MUS MUSCULUS ACETYLCHOLINESTERASE IN COMPLEX WITH (S)-
REMARK 900  C5685 AT 2.25 A RESOLUTION.
DBREF  4ARA A    1   543  UNP    P21836   ACES_MOUSE      32    574
DBREF  4ARA B    1   543  UNP    P21836   ACES_MOUSE      32    574
SEQADV 4ARA ALA A  544  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA THR A  545  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA GLU A  546  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA ALA A  547  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA PRO A  548  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA ALA B  544  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA THR B  545  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA GLU B  546  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA ALA B  547  UNP  P21836              EXPRESSION TAG
SEQADV 4ARA PRO B  548  UNP  P21836              EXPRESSION TAG
SEQRES   1 A  548  GLU GLY ARG GLU ASP PRO GLN LEU LEU VAL ARG VAL ARG
SEQRES   2 A  548  GLY GLY GLN LEU ARG GLY ILE ARG LEU LYS ALA PRO GLY
SEQRES   3 A  548  GLY PRO VAL SER ALA PHE LEU GLY ILE PRO PHE ALA GLU
SEQRES   4 A  548  PRO PRO VAL GLY SER ARG ARG PHE MET PRO PRO GLU PRO
SEQRES   5 A  548  LYS ARG PRO TRP SER GLY VAL LEU ASP ALA THR THR PHE
SEQRES   6 A  548  GLN ASN VAL CYS TYR GLN TYR VAL ASP THR LEU TYR PRO
SEQRES   7 A  548  GLY PHE GLU GLY THR GLU MET TRP ASN PRO ASN ARG GLU
SEQRES   8 A  548  LEU SER GLU ASP CYS LEU TYR LEU ASN VAL TRP THR PRO
SEQRES   9 A  548  TYR PRO ARG PRO ALA SER PRO THR PRO VAL LEU ILE TRP
SEQRES  10 A  548  ILE TYR GLY GLY GLY PHE TYR SER GLY ALA ALA SER LEU
SEQRES  11 A  548  ASP VAL TYR ASP GLY ARG PHE LEU ALA GLN VAL GLU GLY
SEQRES  12 A  548  ALA VAL LEU VAL SER MET ASN TYR ARG VAL GLY THR PHE
SEQRES  13 A  548  GLY PHE LEU ALA LEU PRO GLY SER ARG GLU ALA PRO GLY
SEQRES  14 A  548  ASN VAL GLY LEU LEU ASP GLN ARG LEU ALA LEU GLN TRP
SEQRES  15 A  548  VAL GLN GLU ASN ILE ALA ALA PHE GLY GLY ASP PRO MET
SEQRES  16 A  548  SER VAL THR LEU PHE GLY GLU SER ALA GLY ALA ALA SER
SEQRES  17 A  548  VAL GLY MET HIS ILE LEU SER LEU PRO SER ARG SER LEU
SEQRES  18 A  548  PHE HIS ARG ALA VAL LEU GLN SER GLY THR PRO ASN GLY
SEQRES  19 A  548  PRO TRP ALA THR VAL SER ALA GLY GLU ALA ARG ARG ARG
SEQRES  20 A  548  ALA THR LEU LEU ALA ARG LEU VAL GLY CYS PRO PRO GLY
SEQRES  21 A  548  GLY ALA GLY GLY ASN ASP THR GLU LEU ILE ALA CYS LEU
SEQRES  22 A  548  ARG THR ARG PRO ALA GLN ASP LEU VAL ASP HIS GLU TRP
SEQRES  23 A  548  HIS VAL LEU PRO GLN GLU SER ILE PHE ARG PHE SER PHE
SEQRES  24 A  548  VAL PRO VAL VAL ASP GLY ASP PHE LEU SER ASP THR PRO
SEQRES  25 A  548  GLU ALA LEU ILE ASN THR GLY ASP PHE GLN ASP LEU GLN
SEQRES  26 A  548  VAL LEU VAL GLY VAL VAL LYS ASP GLU GLY SER TYR PHE
SEQRES  27 A  548  LEU VAL TYR GLY VAL PRO GLY PHE SER LYS ASP ASN GLU
SEQRES  28 A  548  SER LEU ILE SER ARG ALA GLN PHE LEU ALA GLY VAL ARG
SEQRES  29 A  548  ILE GLY VAL PRO GLN ALA SER ASP LEU ALA ALA GLU ALA
SEQRES  30 A  548  VAL VAL LEU HIS TYR THR ASP TRP LEU HIS PRO GLU ASP
SEQRES  31 A  548  PRO THR HIS LEU ARG ASP ALA MET SER ALA VAL VAL GLY
SEQRES  32 A  548  ASP HIS ASN VAL VAL CYS PRO VAL ALA GLN LEU ALA GLY
SEQRES  33 A  548  ARG LEU ALA ALA GLN GLY ALA ARG VAL TYR ALA TYR ILE
SEQRES  34 A  548  PHE GLU HIS ARG ALA SER THR LEU THR TRP PRO LEU TRP
SEQRES  35 A  548  MET GLY VAL PRO HIS GLY TYR GLU ILE GLU PHE ILE PHE
SEQRES  36 A  548  GLY LEU PRO LEU ASP PRO SER LEU ASN TYR THR THR GLU
SEQRES  37 A  548  GLU ARG ILE PHE ALA GLN ARG LEU MET LYS TYR TRP THR
SEQRES  38 A  548  ASN PHE ALA ARG THR GLY ASP PRO ASN ASP PRO ARG ASP
SEQRES  39 A  548  SER LYS SER PRO GLN TRP PRO PRO TYR THR THR ALA ALA
SEQRES  40 A  548  GLN GLN TYR VAL SER LEU ASN LEU LYS PRO LEU GLU VAL
SEQRES  41 A  548  ARG ARG GLY LEU ARG ALA GLN THR CYS ALA PHE TRP ASN
SEQRES  42 A  548  ARG PHE LEU PRO LYS LEU LEU SER ALA THR ALA THR GLU
SEQRES  43 A  548  ALA PRO
SEQRES   1 B  548  GLU GLY ARG GLU ASP PRO GLN LEU LEU VAL ARG VAL ARG
SEQRES   2 B  548  GLY GLY GLN LEU ARG GLY ILE ARG LEU LYS ALA PRO GLY
SEQRES   3 B  548  GLY PRO VAL SER ALA PHE LEU GLY ILE PRO PHE ALA GLU
SEQRES   4 B  548  PRO PRO VAL GLY SER ARG ARG PHE MET PRO PRO GLU PRO
SEQRES   5 B  548  LYS ARG PRO TRP SER GLY VAL LEU ASP ALA THR THR PHE
SEQRES   6 B  548  GLN ASN VAL CYS TYR GLN TYR VAL ASP THR LEU TYR PRO
SEQRES   7 B  548  GLY PHE GLU GLY THR GLU MET TRP ASN PRO ASN ARG GLU
SEQRES   8 B  548  LEU SER GLU ASP CYS LEU TYR LEU ASN VAL TRP THR PRO
SEQRES   9 B  548  TYR PRO ARG PRO ALA SER PRO THR PRO VAL LEU ILE TRP
SEQRES  10 B  548  ILE TYR GLY GLY GLY PHE TYR SER GLY ALA ALA SER LEU
SEQRES  11 B  548  ASP VAL TYR ASP GLY ARG PHE LEU ALA GLN VAL GLU GLY
SEQRES  12 B  548  ALA VAL LEU VAL SER MET ASN TYR ARG VAL GLY THR PHE
SEQRES  13 B  548  GLY PHE LEU ALA LEU PRO GLY SER ARG GLU ALA PRO GLY
SEQRES  14 B  548  ASN VAL GLY LEU LEU ASP GLN ARG LEU ALA LEU GLN TRP
SEQRES  15 B  548  VAL GLN GLU ASN ILE ALA ALA PHE GLY GLY ASP PRO MET
SEQRES  16 B  548  SER VAL THR LEU PHE GLY GLU SER ALA GLY ALA ALA SER
SEQRES  17 B  548  VAL GLY MET HIS ILE LEU SER LEU PRO SER ARG SER LEU
SEQRES  18 B  548  PHE HIS ARG ALA VAL LEU GLN SER GLY THR PRO ASN GLY
SEQRES  19 B  548  PRO TRP ALA THR VAL SER ALA GLY GLU ALA ARG ARG ARG
SEQRES  20 B  548  ALA THR LEU LEU ALA ARG LEU VAL GLY CYS PRO PRO GLY
SEQRES  21 B  548  GLY ALA GLY GLY ASN ASP THR GLU LEU ILE ALA CYS LEU
SEQRES  22 B  548  ARG THR ARG PRO ALA GLN ASP LEU VAL ASP HIS GLU TRP
SEQRES  23 B  548  HIS VAL LEU PRO GLN GLU SER ILE PHE ARG PHE SER PHE
SEQRES  24 B  548  VAL PRO VAL VAL ASP GLY ASP PHE LEU SER ASP THR PRO
SEQRES  25 B  548  GLU ALA LEU ILE ASN THR GLY ASP PHE GLN ASP LEU GLN
SEQRES  26 B  548  VAL LEU VAL GLY VAL VAL LYS ASP GLU GLY SER TYR PHE
SEQRES  27 B  548  LEU VAL TYR GLY VAL PRO GLY PHE SER LYS ASP ASN GLU
SEQRES  28 B  548  SER LEU ILE SER ARG ALA GLN PHE LEU ALA GLY VAL ARG
SEQRES  29 B  548  ILE GLY VAL PRO GLN ALA SER ASP LEU ALA ALA GLU ALA
SEQRES  30 B  548  VAL VAL LEU HIS TYR THR ASP TRP LEU HIS PRO GLU ASP
SEQRES  31 B  548  PRO THR HIS LEU ARG ASP ALA MET SER ALA VAL VAL GLY
SEQRES  32 B  548  ASP HIS ASN VAL VAL CYS PRO VAL ALA GLN LEU ALA GLY
SEQRES  33 B  548  ARG LEU ALA ALA GLN GLY ALA ARG VAL TYR ALA TYR ILE
SEQRES  34 B  548  PHE GLU HIS ARG ALA SER THR LEU THR TRP PRO LEU TRP
SEQRES  35 B  548  MET GLY VAL PRO HIS GLY TYR GLU ILE GLU PHE ILE PHE
SEQRES  36 B  548  GLY LEU PRO LEU ASP PRO SER LEU ASN TYR THR THR GLU
SEQRES  37 B  548  GLU ARG ILE PHE ALA GLN ARG LEU MET LYS TYR TRP THR
SEQRES  38 B  548  ASN PHE ALA ARG THR GLY ASP PRO ASN ASP PRO ARG ASP
SEQRES  39 B  548  SER LYS SER PRO GLN TRP PRO PRO TYR THR THR ALA ALA
SEQRES  40 B  548  GLN GLN TYR VAL SER LEU ASN LEU LYS PRO LEU GLU VAL
SEQRES  41 B  548  ARG ARG GLY LEU ARG ALA GLN THR CYS ALA PHE TRP ASN
SEQRES  42 B  548  ARG PHE LEU PRO LYS LEU LEU SER ALA THR ALA THR GLU
SEQRES  43 B  548  ALA PRO
HET    C56  A 600      50
HET    PEG  A1544       7
HET    EDO  A1545       4
HET    PEG  A1546       7
HET    PEG  A1547       7
HET    EDO  A1548       4
HET    NAG  A1549      14
HET    EDO  A1550       4
HET    EDO  A1551       4
HET    EDO  A1552       4
HET    NAG  A1553      14
HET    C56  B 600      50
HET    PEG  B1545       7
HET    1PE  B1546      16
HET    PEG  B1547       7
HET    PEG  B1548       7
HET    EDO  B1549       4
HET    PEG  B1550       7
HET    PEG  B1551       7
HET    NAG  B1552      14
HET    EDO  B1553       4
HET    PEG  B1554       7
HET     CL  B1555       1
HET    EDO  B1556       4
HETNAM     C56 4-(DIMETHYLAMINO)-N-{[(2R)-1-ETHYLPYRROLIDIN-
HETNAM   2 C56  2-YL]METHYL}-2-METHOXY-5-NITROBENZAMIDE
HETNAM     PEG DI(HYDROXYETHYL)ETHER
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     1PE PENTAETHYLENE GLYCOL
HETNAM      CL CHLORIDE ION
HETSYN     EDO ETHYLENE GLYCOL
HETSYN     1PE PEG400
FORMUL   3  C56    2(C17 H26 N4 O4)
FORMUL   4  PEG    9(C4 H10 O3)
FORMUL   5  EDO    8(C2 H6 O2)
FORMUL   6  NAG    3(C8 H15 N O6)
FORMUL   7  1PE    C10 H22 O6
FORMUL   8   CL    CL 1-
FORMUL   9  HOH   *628(H2 O)
HELIX    1   1 ASP A    5  GLN A    7  5                                   3
HELIX    2   2 VAL A   42  ARG A   46  5                                   5
HELIX    3   3 PHE A   80  MET A   85  1                                   6
HELIX    4   4 LEU A  130  ASP A  134  5                                   5
HELIX    5   5 GLY A  135  GLU A  142  1                                   8
HELIX    6   6 VAL A  153  LEU A  159  1                                   7
HELIX    7   7 ASN A  170  ILE A  187  1                                  18
HELIX    8   8 ALA A  188  PHE A  190  5                                   3
HELIX    9   9 SER A  203  SER A  215  1                                  13
HELIX   10  10 LEU A  216  ARG A  219  5                                   4
HELIX   11  11 SER A  240  GLY A  256  1                                  17
HELIX   12  12 ASN A  265  ARG A  274  1                                  10
HELIX   13  13 PRO A  277  HIS A  284  1                                   8
HELIX   14  14 GLU A  285  LEU A  289  5                                   5
HELIX   15  15 THR A  311  THR A  318  1                                   8
HELIX   16  16 GLY A  335  GLY A  342  5                                   8
HELIX   17  17 SER A  355  VAL A  367  1                                  13
HELIX   18  18 SER A  371  THR A  383  1                                  13
HELIX   19  19 ASP A  390  VAL A  407  1                                  18
HELIX   20  20 VAL A  407  GLN A  421  1                                  15
HELIX   21  21 PRO A  440  GLY A  444  5                                   5
HELIX   22  22 GLU A  450  PHE A  455  1                                   6
HELIX   23  23 GLY A  456  ASP A  460  5                                   5
HELIX   24  24 ASP A  460  ASN A  464  5                                   5
HELIX   25  25 THR A  466  GLY A  487  1                                  22
HELIX   26  26 ARG A  525  ARG A  534  1                                  10
HELIX   27  27 ARG A  534  SER A  541  1                                   8
HELIX   28  28 VAL B   42  ARG B   46  5                                   5
HELIX   29  29 PHE B   80  MET B   85  1                                   6
HELIX   30  30 LEU B  130  ASP B  134  5                                   5
HELIX   31  31 GLY B  135  GLU B  142  1                                   8
HELIX   32  32 VAL B  153  LEU B  159  1                                   7
HELIX   33  33 ASN B  170  ILE B  187  1                                  18
HELIX   34  34 ALA B  188  PHE B  190  5                                   3
HELIX   35  35 SER B  203  LEU B  214  1                                  12
HELIX   36  36 SER B  215  LEU B  221  5                                   7
HELIX   37  37 SER B  240  GLY B  256  1                                  17
HELIX   38  38 ASN B  265  THR B  275  1                                  11
HELIX   39  39 PRO B  277  ASP B  283  1                                   7
HELIX   40  40 HIS B  284  LEU B  289  5                                   6
HELIX   41  41 THR B  311  GLY B  319  1                                   9
HELIX   42  42 GLY B  335  GLY B  342  5                                   8
HELIX   43  43 SER B  355  VAL B  367  1                                  13
HELIX   44  44 SER B  371  THR B  383  1                                  13
HELIX   45  45 ASP B  390  VAL B  407  1                                  18
HELIX   46  46 VAL B  407  GLN B  421  1                                  15
HELIX   47  47 PRO B  440  GLY B  444  5                                   5
HELIX   48  48 GLU B  450  PHE B  455  1                                   6
HELIX   49  49 GLY B  456  ASP B  460  5                                   5
HELIX   50  50 ASP B  460  ASN B  464  5                                   5
HELIX   51  51 THR B  466  GLY B  487  1                                  22
HELIX   52  52 ARG B  525  ARG B  534  1                                  10
HELIX   53  53 PHE B  535  SER B  541  1                                   7
SHEET    1  AA 3 LEU A   9  VAL A  12  0
SHEET    2  AA 3 GLY A  15  ARG A  18 -1  O  GLY A  15   N  VAL A  12
SHEET    3  AA 3 VAL A  59  ASP A  61  1  O  LEU A  60   N  ARG A  18
SHEET    1  AB11 ILE A  20  ALA A  24  0
SHEET    2  AB11 GLY A  27  PRO A  36 -1  O  GLY A  27   N  ALA A  24
SHEET    3  AB11 TYR A  98  PRO A 104 -1  O  LEU A  99   N  ILE A  35
SHEET    4  AB11 VAL A 145  MET A 149 -1  O  LEU A 146   N  TRP A 102
SHEET    5  AB11 THR A 112  ILE A 118  1  O  PRO A 113   N  VAL A 145
SHEET    6  AB11 GLY A 192  GLU A 202  1  N  ASP A 193   O  THR A 112
SHEET    7  AB11 ARG A 224  GLN A 228  1  O  ARG A 224   N  LEU A 199
SHEET    8  AB11 GLN A 325  VAL A 331  1  O  GLN A 325   N  ALA A 225
SHEET    9  AB11 ARG A 424  PHE A 430  1  O  ARG A 424   N  VAL A 326
SHEET   10  AB11 GLN A 509  LEU A 513  1  O  VAL A 511   N  ILE A 429
SHEET   11  AB11 GLU A 519  ARG A 522 -1  O  GLU A 519   N  SER A 512
SHEET    1  AC 2 VAL A  68  CYS A  69  0
SHEET    2  AC 2 LEU A  92  SER A  93  1  N  SER A  93   O  VAL A  68
SHEET    1  BA 3 LEU B   9  VAL B  12  0
SHEET    2  BA 3 GLY B  15  ARG B  18 -1  O  GLY B  15   N  VAL B  12
SHEET    3  BA 3 VAL B  59  ASP B  61  1  O  LEU B  60   N  ARG B  18
SHEET    1  BB11 ILE B  20  ALA B  24  0
SHEET    2  BB11 GLY B  27  PRO B  36 -1  O  GLY B  27   N  ALA B  24
SHEET    3  BB11 TYR B  98  PRO B 104 -1  O  LEU B  99   N  ILE B  35
SHEET    4  BB11 VAL B 145  MET B 149 -1  O  LEU B 146   N  TRP B 102
SHEET    5  BB11 THR B 112  ILE B 118  1  O  PRO B 113   N  VAL B 145
SHEET    6  BB11 GLY B 192  GLU B 202  1  N  ASP B 193   O  THR B 112
SHEET    7  BB11 ARG B 224  GLN B 228  1  O  ARG B 224   N  LEU B 199
SHEET    8  BB11 GLN B 325  VAL B 331  1  O  GLN B 325   N  ALA B 225
SHEET    9  BB11 ARG B 424  PHE B 430  1  O  ARG B 424   N  VAL B 326
SHEET   10  BB11 GLN B 509  LEU B 513  1  O  VAL B 511   N  ILE B 429
SHEET   11  BB11 GLU B 519  ARG B 522 -1  O  GLU B 519   N  SER B 512
SHEET    1  BC 2 VAL B  68  CYS B  69  0
SHEET    2  BC 2 LEU B  92  SER B  93  1  N  SER B  93   O  VAL B  68
SSBOND   1 CYS A   69    CYS A   96                          1555   1555  2.04
SSBOND   2 CYS A  257    CYS A  272                          1555   1555  2.04
SSBOND   3 CYS A  409    CYS A  529                          1555   1555  2.03
SSBOND   4 CYS B   69    CYS B   96                          1555   1555  2.04
SSBOND   5 CYS B  257    CYS B  272                          1555   1555  2.04
SSBOND   6 CYS B  409    CYS B  529                          1555   1555  2.04
LINK         ND2 ASN A 350                 C1  NAG A1549     1555   1555  1.47
LINK         ND2 ASN A 464                 C1  NAG A1553     1555   1555  1.46
LINK         ND2 ASN B 350                 C1  NAG B1552     1555   1555  1.47
CISPEP   1 TYR A  105    PRO A  106          0        -0.99
CISPEP   2 TYR B  105    PRO B  106          0         1.55
CISPEP   3 SER B  497    PRO B  498          0        11.72
SITE     1 AC1 13 TYR A  72  ASP A  74  TRP A  86  TYR A 124
SITE     2 AC1 13 TRP A 286  ILE A 294  PHE A 295  ARG A 296
SITE     3 AC1 13 TYR A 337  PHE A 338  TYR A 341  HOH A2236
SITE     4 AC1 13 HOH A2258
SITE     1 AC2  3 ASP A 304  SER A 309  ASP A 310
SITE     1 AC3  1 GLU A 313
SITE     1 AC4  2 ASP A 323  PEG A1547
SITE     1 AC5  5 ILE A 213  ARG A 219  PHE A 222  ASP A 323
SITE     2 AC5  5 PEG A1546
SITE     1 AC6  1 GLU A 313
SITE     1 AC7  1 ASP A   5
SITE     1 AC8  3 SER A  57  VAL A  59  HOH A2346
SITE     1 AC9  2 THR A  63  HOH A2347
SITE     1 BC1 15 TYR B  72  ASP B  74  TRP B  86  GLY B 121
SITE     2 BC1 15 GLY B 122  TYR B 124  TRP B 286  SER B 293
SITE     3 BC1 15 ILE B 294  PHE B 295  ARG B 296  TYR B 337
SITE     4 BC1 15 PHE B 338  TYR B 341  HOH B2176
SITE     1 BC2  6 THR B 112  PRO B 113  GLU B 142  GLY B 143
SITE     2 BC2  6 ALA B 144  ARG B 485
SITE     1 BC3 11 LEU A 380  HIS A 381  GLN A 527  PHE A 535
SITE     2 BC3 11 ALA B 377  LEU B 380  HIS B 381  PHE B 531
SITE     3 BC3 11 PHE B 535  HOH B2203  HOH B2266
SITE     1 BC4  2 ASP B 323  EDO B1549
SITE     1 BC5  3 HIS B 381  TYR B 382  HOH B2268
SITE     1 BC6  2 ARG B 219  PEG B1547
SITE     1 BC7  3 ARG B  11  GLU B 185  HOH B2269
SITE     1 BC8  5 LYS B 332  ASP B 333  ASP B 396  LEU B 441
SITE     2 BC8  5 TRP B 442
SITE     1 BC9  1 GLU B 313
SITE     1 CC1  1 SER B 309
SITE     1 CC2  1 SER B 240
SITE     1 CC3  1 LYS B 516
SITE     1 CC4  5 SER A 347  ASN A 350  HOH A2261  HOH A2344
SITE     2 CC4  5 HOH A2345
SITE     1 CC5  1 ASN A 464
SITE     1 CC6  3 SER B 347  ASN B 350  HOH B2270
CRYST1   78.764  112.012  227.340  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.012696  0.000000  0.000000        0.00000
SCALE2      0.000000  0.008928  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004399        0.00000
TER    4174      ALA A 542
TER    8338      THR B 543
MASTER      867    0   24   53   32    0   36    6 9218    2  268   86
END