longtext: 4B6G-pdb

content
HEADER    HYDROLASE                               13-AUG-12   4B6G
TITLE     THE CRYSTAL STRUCTURE OF THE NEISSERIAL ESTERASE D.
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PUTATIVE ESTERASE;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: S-FORMYLGLUTATHIONE HYDROLASE PROTEIN;
COMPND   5 EC: 3.1.2.12;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: NEISSERIA MENINGITIDIS MC58;
SOURCE   3 ORGANISM_TAXID: 122586;
SOURCE   4 STRAIN: C311;
SOURCE   5 VARIANT: NUMBER 3;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PMSGC7
KEYWDS    HYDROLASE, FORMALDEHYDE DETOXIFICATION, ALPHA/BETA SERINE HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    R.M.COUNAGO,B.KOBE
REVDAT   1   14-NOV-12 4B6G    0
JRNL        AUTH   N.H.CHEN,R.M.COUNAGO,K.Y.DJOKO,M.P.JENNINGS,M.A.APICELLA,
JRNL        AUTH 2 B.KOBE,A.G.MCEWAN
JRNL        TITL   A GLUTATHIONE-DEPENDENT DETOXIFICATION SYSTEM IS REQUIRED
JRNL        TITL 2 FOR FORMALDEHYDE RESISTANCE AND OPTIMAL SURVIVAL OF
JRNL        TITL 3 NEISSERIA MENINGITIDIS IN BIOFILMS.
JRNL        REF    ANTIOXID.REDOX SIGNAL.                     2012
JRNL        REFN                   ESSN 1557-7716
JRNL        PMID   22937752
JRNL        DOI    10.1089/ARS.2012.4749
REMARK   2
REMARK   2 RESOLUTION.    1.40 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VICENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.400
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.842
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 0.00
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.65
REMARK   3   NUMBER OF REFLECTIONS             : 121215
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.1434
REMARK   3   R VALUE            (WORKING SET) : 0.1431
REMARK   3   FREE R VALUE                     : 0.1539
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 3.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 3610
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 19.8436 -  4.1355    0.96     4749   139  0.1429 0.1552
REMARK   3     2  4.1355 -  3.2873    0.98     4592   180  0.1337 0.1479
REMARK   3     3  3.2873 -  2.8732    0.99     4603   143  0.1428 0.1549
REMARK   3     4  2.8732 -  2.6111    1.00     4593   142  0.1383 0.1518
REMARK   3     5  2.6111 -  2.4243    1.00     4575   134  0.1329 0.1478
REMARK   3     6  2.4243 -  2.2816    1.00     4553   156  0.1230 0.1300
REMARK   3     7  2.2816 -  2.1675    1.00     4542   146  0.1288 0.1347
REMARK   3     8  2.1675 -  2.0732    1.00     4532   140  0.1281 0.1348
REMARK   3     9  2.0732 -  1.9935    1.00     4563   121  0.1310 0.1503
REMARK   3    10  1.9935 -  1.9248    1.00     4537   131  0.1295 0.1278
REMARK   3    11  1.9248 -  1.8646    1.00     4523   138  0.1320 0.1433
REMARK   3    12  1.8646 -  1.8114    1.00     4511   143  0.1324 0.1372
REMARK   3    13  1.8114 -  1.7637    1.00     4490   145  0.1379 0.1553
REMARK   3    14  1.7637 -  1.7207    1.00     4492   153  0.1462 0.1665
REMARK   3    15  1.7207 -  1.6816    1.00     4500   139  0.1489 0.1712
REMARK   3    16  1.6816 -  1.6458    1.00     4499   127  0.1482 0.1580
REMARK   3    17  1.6458 -  1.6129    1.00     4477   135  0.1442 0.1512
REMARK   3    18  1.6129 -  1.5825    1.00     4505   132  0.1544 0.1563
REMARK   3    19  1.5825 -  1.5542    1.00     4493   122  0.1589 0.1772
REMARK   3    20  1.5542 -  1.5279    1.00     4474   147  0.1645 0.1627
REMARK   3    21  1.5279 -  1.5033    1.00     4444   132  0.1783 0.1762
REMARK   3    22  1.5033 -  1.4801    1.00     4516   126  0.1840 0.1897
REMARK   3    23  1.4801 -  1.4584    1.00     4469   133  0.1915 0.1958
REMARK   3    24  1.4584 -  1.4379    1.00     4471   134  0.2018 0.2566
REMARK   3    25  1.4379 -  1.4184    1.00     4469   111  0.2190 0.2222
REMARK   3    26  1.4184 -  1.4000    1.00     4433   161  0.2209 0.2448
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.70
REMARK   3   K_SOL              : 0
REMARK   3   B_SOL              : 0
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.09
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 13.36
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 10.91
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 21.023
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.011           4651
REMARK   3   ANGLE     :  1.222           6315
REMARK   3   CHIRALITY :  0.077            641
REMARK   3   PLANARITY :  0.007            839
REMARK   3   DIHEDRAL  : 14.519           1690
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 14
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN A AND (RESID -7 THROUGH 11 )
REMARK   3    ORIGIN FOR THE GROUP (A):  21.0100 102.0605  30.4042
REMARK   3    T TENSOR
REMARK   3      T11:   0.1826 T22:   0.1404
REMARK   3      T33:   0.1437 T12:  -0.0092
REMARK   3      T13:   0.0013 T23:  -0.0038
REMARK   3    L TENSOR
REMARK   3      L11:   0.0347 L22:   0.1193
REMARK   3      L33:   0.0182 L12:   0.0627
REMARK   3      L13:  -0.0218 L23:  -0.0446
REMARK   3    S TENSOR
REMARK   3      S11:   0.0187 S12:  -0.1132 S13:   0.0164
REMARK   3      S21:   0.0339 S22:  -0.0025 S23:   0.0283
REMARK   3      S31:  -0.0328 S32:   0.0467 S33:   0.0000
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN A AND (RESID 12 THROUGH 34 )
REMARK   3    ORIGIN FOR THE GROUP (A):  15.4846  99.6806  21.7720
REMARK   3    T TENSOR
REMARK   3      T11:   0.1563 T22:   0.1252
REMARK   3      T33:   0.1543 T12:  -0.0006
REMARK   3      T13:  -0.0031 T23:  -0.0035
REMARK   3    L TENSOR
REMARK   3      L11:   0.0880 L22:   0.0649
REMARK   3      L33:   0.1368 L12:   0.0705
REMARK   3      L13:  -0.0655 L23:  -0.0250
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0017 S12:   0.0049 S13:   0.0635
REMARK   3      S21:   0.0427 S22:  -0.0324 S23:   0.0395
REMARK   3      S31:  -0.0623 S32:   0.0261 S33:  -0.0000
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN A AND (RESID 35 THROUGH 99 )
REMARK   3    ORIGIN FOR THE GROUP (A):  18.6055  90.4300  21.0711
REMARK   3    T TENSOR
REMARK   3      T11:   0.1286 T22:   0.1141
REMARK   3      T33:   0.1257 T12:  -0.0013
REMARK   3      T13:   0.0021 T23:   0.0067
REMARK   3    L TENSOR
REMARK   3      L11:   0.2427 L22:   0.4232
REMARK   3      L33:   0.7290 L12:   0.0107
REMARK   3      L13:   0.0643 L23:  -0.0662
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0272 S12:  -0.0215 S13:  -0.0122
REMARK   3      S21:   0.0290 S22:  -0.0096 S23:   0.0119
REMARK   3      S31:  -0.0387 S32:   0.0144 S33:  -0.0001
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN A AND (RESID 100 THROUGH 177 )
REMARK   3    ORIGIN FOR THE GROUP (A):  14.9932  91.7852   9.7404
REMARK   3    T TENSOR
REMARK   3      T11:   0.1362 T22:   0.1268
REMARK   3      T33:   0.1338 T12:   0.0017
REMARK   3      T13:   0.0015 T23:   0.0066
REMARK   3    L TENSOR
REMARK   3      L11:   0.2993 L22:   0.2539
REMARK   3      L33:   0.5666 L12:   0.0182
REMARK   3      L13:   0.0351 L23:   0.0412
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0221 S12:   0.0174 S13:  -0.0078
REMARK   3      S21:  -0.0355 S22:   0.0013 S23:   0.0401
REMARK   3      S31:  -0.0492 S32:  -0.0260 S33:   0.0000
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN A AND (RESID 178 THROUGH 229 )
REMARK   3    ORIGIN FOR THE GROUP (A):   9.4016  79.8831   5.6104
REMARK   3    T TENSOR
REMARK   3      T11:   0.1499 T22:   0.1645
REMARK   3      T33:   0.1645 T12:  -0.0047
REMARK   3      T13:  -0.0043 T23:  -0.0011
REMARK   3    L TENSOR
REMARK   3      L11:   0.1786 L22:   0.3604
REMARK   3      L33:   0.5099 L12:  -0.0880
REMARK   3      L13:  -0.0543 L23:   0.1719
REMARK   3    S TENSOR
REMARK   3      S11:   0.0078 S12:   0.0682 S13:  -0.0744
REMARK   3      S21:  -0.0645 S22:  -0.0354 S23:   0.0532
REMARK   3      S31:   0.0532 S32:  -0.0969 S33:  -0.0000
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN A AND (RESID 230 THROUGH 275 )
REMARK   3    ORIGIN FOR THE GROUP (A):  22.9752  75.7497  11.1054
REMARK   3    T TENSOR
REMARK   3      T11:   0.1649 T22:   0.1379
REMARK   3      T33:   0.1418 T12:   0.0061
REMARK   3      T13:   0.0077 T23:   0.0002
REMARK   3    L TENSOR
REMARK   3      L11:   0.2236 L22:   0.1147
REMARK   3      L33:   0.2849 L12:  -0.1251
REMARK   3      L13:   0.0021 L23:   0.1123
REMARK   3    S TENSOR
REMARK   3      S11:   0.0063 S12:   0.0274 S13:  -0.0402
REMARK   3      S21:   0.0028 S22:  -0.0320 S23:   0.0105
REMARK   3      S31:   0.0940 S32:   0.0622 S33:  -0.0000
REMARK   3   TLS GROUP : 7
REMARK   3    SELECTION: CHAIN B AND (RESID 1 THROUGH 21 )
REMARK   3    ORIGIN FOR THE GROUP (A):  16.6979  66.8211  34.6417
REMARK   3    T TENSOR
REMARK   3      T11:   0.2238 T22:   0.1287
REMARK   3      T33:   0.1785 T12:  -0.0308
REMARK   3      T13:   0.0350 T23:  -0.0080
REMARK   3    L TENSOR
REMARK   3      L11:   0.4677 L22:   0.4099
REMARK   3      L33:   0.3501 L12:  -0.0448
REMARK   3      L13:   0.3421 L23:  -0.0952
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0653 S12:   0.0657 S13:  -0.0964
REMARK   3      S21:   0.0179 S22:  -0.0440 S23:   0.0285
REMARK   3      S31:   0.3181 S32:  -0.0987 S33:   0.0223
REMARK   3   TLS GROUP : 8
REMARK   3    SELECTION: CHAIN B AND (RESID 22 THROUGH 145 )
REMARK   3    ORIGIN FOR THE GROUP (A):  19.4694  72.5514  45.6658
REMARK   3    T TENSOR
REMARK   3      T11:   0.1691 T22:   0.1241
REMARK   3      T33:   0.1571 T12:   0.0080
REMARK   3      T13:   0.0297 T23:   0.0138
REMARK   3    L TENSOR
REMARK   3      L11:   0.5138 L22:   0.3772
REMARK   3      L33:   0.8673 L12:  -0.1566
REMARK   3      L13:  -0.2186 L23:   0.4403
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0679 S12:  -0.0171 S13:  -0.0674
REMARK   3      S21:   0.0401 S22:  -0.0203 S23:   0.0795
REMARK   3      S31:   0.1715 S32:  -0.0421 S33:  -0.0012
REMARK   3   TLS GROUP : 9
REMARK   3    SELECTION: CHAIN B AND (RESID 146 THROUGH 168 )
REMARK   3    ORIGIN FOR THE GROUP (A):  29.6977  75.2712  53.7751
REMARK   3    T TENSOR
REMARK   3      T11:   0.1804 T22:   0.1792
REMARK   3      T33:   0.1535 T12:   0.0436
REMARK   3      T13:   0.0139 T23:   0.0243
REMARK   3    L TENSOR
REMARK   3      L11:   0.0747 L22:   0.0884
REMARK   3      L33:   0.0961 L12:   0.0372
REMARK   3      L13:  -0.0166 L23:   0.0714
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0512 S12:  -0.1029 S13:  -0.0740
REMARK   3      S21:   0.1058 S22:   0.0348 S23:   0.0540
REMARK   3      S31:   0.1021 S32:   0.0719 S33:  -0.0000
REMARK   3   TLS GROUP : 10
REMARK   3    SELECTION: CHAIN B AND (RESID 169 THROUGH 188 )
REMARK   3    ORIGIN FOR THE GROUP (A):  19.4948  88.4764  57.4932
REMARK   3    T TENSOR
REMARK   3      T11:   0.1777 T22:   0.1690
REMARK   3      T33:   0.1731 T12:   0.0012
REMARK   3      T13:   0.0177 T23:  -0.0062
REMARK   3    L TENSOR
REMARK   3      L11:   0.0333 L22:   0.0677
REMARK   3      L33:   0.0424 L12:  -0.0397
REMARK   3      L13:   0.0005 L23:  -0.0302
REMARK   3    S TENSOR
REMARK   3      S11:   0.0453 S12:  -0.0578 S13:   0.0708
REMARK   3      S21:   0.0835 S22:  -0.0357 S23:   0.0483
REMARK   3      S31:  -0.0460 S32:   0.0154 S33:  -0.0000
REMARK   3   TLS GROUP : 11
REMARK   3    SELECTION: CHAIN B AND (RESID 189 THROUGH 213 )
REMARK   3    ORIGIN FOR THE GROUP (A):  27.2404  82.3406  63.8736
REMARK   3    T TENSOR
REMARK   3      T11:   0.2068 T22:   0.2413
REMARK   3      T33:   0.1486 T12:   0.0231
REMARK   3      T13:   0.0025 T23:   0.0146
REMARK   3    L TENSOR
REMARK   3      L11:   0.2820 L22:   0.2861
REMARK   3      L33:   0.0785 L12:   0.2515
REMARK   3      L13:   0.0287 L23:  -0.0211
REMARK   3    S TENSOR
REMARK   3      S11:   0.0520 S12:  -0.2715 S13:  -0.0029
REMARK   3      S21:   0.1771 S22:  -0.0500 S23:   0.0471
REMARK   3      S31:   0.0660 S32:   0.2094 S33:   0.0015
REMARK   3   TLS GROUP : 12
REMARK   3    SELECTION: CHAIN B AND (RESID 214 THROUGH 229 )
REMARK   3    ORIGIN FOR THE GROUP (A):  28.0555  93.2504  46.8353
REMARK   3    T TENSOR
REMARK   3      T11:   0.1900 T22:   0.1638
REMARK   3      T33:   0.1674 T12:  -0.0012
REMARK   3      T13:   0.0180 T23:   0.0044
REMARK   3    L TENSOR
REMARK   3      L11:   0.0975 L22:   0.1175
REMARK   3      L33:   0.0979 L12:   0.1072
REMARK   3      L13:  -0.0071 L23:  -0.0092
REMARK   3    S TENSOR
REMARK   3      S11:   0.0924 S12:  -0.0496 S13:   0.0438
REMARK   3      S21:  -0.0287 S22:  -0.0385 S23:   0.0156
REMARK   3      S31:  -0.1455 S32:   0.1241 S33:  -0.0000
REMARK   3   TLS GROUP : 13
REMARK   3    SELECTION: CHAIN B AND (RESID 230 THROUGH 255 )
REMARK   3    ORIGIN FOR THE GROUP (A):  36.1722  90.0560  51.0750
REMARK   3    T TENSOR
REMARK   3      T11:   0.1738 T22:   0.2356
REMARK   3      T33:   0.1617 T12:  -0.0244
REMARK   3      T13:  -0.0237 T23:   0.0050
REMARK   3    L TENSOR
REMARK   3      L11:   0.1447 L22:   0.1775
REMARK   3      L33:   0.1928 L12:  -0.0846
REMARK   3      L13:  -0.1643 L23:   0.0656
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0201 S12:  -0.0746 S13:   0.1277
REMARK   3      S21:   0.0448 S22:  -0.0282 S23:  -0.1094
REMARK   3      S31:  -0.1248 S32:   0.3079 S33:   0.0004
REMARK   3   TLS GROUP : 14
REMARK   3    SELECTION: CHAIN B AND (RESID 256 THROUGH 275 )
REMARK   3    ORIGIN FOR THE GROUP (A):  32.9212  79.9926  39.3469
REMARK   3    T TENSOR
REMARK   3      T11:   0.1493 T22:   0.1596
REMARK   3      T33:   0.1462 T12:   0.0216
REMARK   3      T13:   0.0067 T23:   0.0115
REMARK   3    L TENSOR
REMARK   3      L11:   0.0612 L22:   0.0410
REMARK   3      L33:   0.1441 L12:  -0.0336
REMARK   3      L13:  -0.0840 L23:   0.0194
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0703 S12:  -0.0278 S13:  -0.0428
REMARK   3      S21:   0.0187 S22:   0.0592 S23:   0.0493
REMARK   3      S31:   0.0050 S32:   0.1258 S33:  -0.0031
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4B6G COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 10-OCT-11.
REMARK 100 THE PDBE ID CODE IS EBI-49945.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 30-OCT-10
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : AUSTRALIAN SYNCHROTRON
REMARK 200  BEAMLINE                       : MX2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.95369
REMARK 200  MONOCHROMATOR                  : DOUBLE CRYSTAL
REMARK 200                                   MONOCHROMATOR (SI111)
REMARK 200  OPTICS                         : SILICON MIRRORS (ADAPTIVE
REMARK 200                                   AND U-BENT)
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD (QUANTUM 315R)
REMARK 200  DETECTOR MANUFACTURER          : ADSC
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 151514
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.30
REMARK 200  RESOLUTION RANGE LOW       (A) : 19.84
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.4
REMARK 200  DATA REDUNDANCY                : 5.2
REMARK 200  R MERGE                    (I) : 0.06
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 15.30
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.30
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.37
REMARK 200  COMPLETENESS FOR SHELL     (%) : 98.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.1
REMARK 200  R MERGE FOR SHELL          (I) : 0.68
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.30
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 3FCX
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 45
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.23
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 25 MM BIS-TRIS (PH 6.5), 200 MM
REMARK 280  LITHIUM SULFATE AND 25% PEG 3350.
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 65 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+2/3
REMARK 290       3555   -X+Y,-X,Z+1/3
REMARK 290       4555   -X,-Y,Z+1/2
REMARK 290       5555   Y,-X+Y,Z+1/6
REMARK 290       6555   X-Y,X,Z+5/6
REMARK 290       7555   Y,X,-Z+2/3
REMARK 290       8555   X-Y,-Y,-Z
REMARK 290       9555   -X,-X+Y,-Z+1/3
REMARK 290      10555   -Y,-X,-Z+1/6
REMARK 290      11555   -X+Y,Y,-Z+1/2
REMARK 290      12555   X,X-Y,-Z+5/6
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      113.24067
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       56.62033
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000       84.93050
REMARK 290   SMTRY1   5  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   5 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000       28.31017
REMARK 290   SMTRY1   6  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000      141.55083
REMARK 290   SMTRY1   7 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   7  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000      113.24067
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   9 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   9 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   9  0.000000  0.000000 -1.000000       56.62033
REMARK 290   SMTRY1  10  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  10 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000       28.31017
REMARK 290   SMTRY1  11 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  11  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000       84.93050
REMARK 290   SMTRY1  12  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  12  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000      141.55083
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU B    -7
REMARK 465     ASN B    -6
REMARK 465     LEU B    -5
REMARK 465     TYR B    -4
REMARK 465     PHE B    -3
REMARK 465     GLN B    -2
REMARK 465     GLY B    -1
REMARK 465     ALA B     0
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A  37       62.89   -163.10
REMARK 500    THR A  53       -4.31     74.15
REMARK 500    THR A  55     -162.52   -112.79
REMARK 500    ALA A 100       48.74   -148.14
REMARK 500    SER A 145     -117.03     64.61
REMARK 500    GLN A 159     -107.63     48.91
REMARK 500    ASP A 253     -155.96   -105.73
REMARK 500    ASN B  37       65.64   -160.47
REMARK 500    THR B  53       -6.14     76.69
REMARK 500    THR B  55     -160.68   -112.54
REMARK 500    ALA B 100       47.69   -149.42
REMARK 500    SER B 145     -114.69     66.58
REMARK 500    ASP B 253     -155.26   -107.28
REMARK 500
REMARK 500 REMARK: NULL
DBREF  4B6G A    3   275  UNP    Q9JZ43   Q9JZ43_NEIMB     3    275
DBREF  4B6G B    3   275  UNP    Q9JZ43   Q9JZ43_NEIMB     3    275
SEQADV 4B6G GLU A   -7  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G ASN A   -6  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G LEU A   -5  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G TYR A   -4  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G PHE A   -3  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLN A   -2  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLY A   -1  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G ALA A    0  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G MET A    1  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLU A    2  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLU B   -7  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G ASN B   -6  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G LEU B   -5  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G TYR B   -4  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G PHE B   -3  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLN B   -2  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLY B   -1  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G ALA B    0  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G MET B    1  UNP  Q9JZ43              EXPRESSION TAG
SEQADV 4B6G GLU B    2  UNP  Q9JZ43              EXPRESSION TAG
SEQRES   1 A  283  GLU ASN LEU TYR PHE GLN GLY ALA MET GLU LEU ILE GLU
SEQRES   2 A  283  GLN HIS GLN ILE PHE GLY GLY SER GLN GLN VAL TRP ALA
SEQRES   3 A  283  HIS HIS ALA GLN THR LEU GLN CYS GLU MET LYS PHE ALA
SEQRES   4 A  283  VAL TYR LEU PRO ASN ASN PRO GLU ASN ARG PRO LEU GLY
SEQRES   5 A  283  VAL ILE TYR TRP LEU SER GLY LEU THR CYS THR GLU GLN
SEQRES   6 A  283  ASN PHE ILE THR LYS SER GLY PHE GLN ARG TYR ALA ALA
SEQRES   7 A  283  GLU HIS GLN VAL ILE VAL VAL ALA PRO ASP THR SER PRO
SEQRES   8 A  283  ARG GLY GLU GLN VAL PRO ASN ASP ASP ALA TYR ASP LEU
SEQRES   9 A  283  GLY GLN SER ALA GLY PHE TYR LEU ASN ALA THR GLU GLN
SEQRES  10 A  283  PRO TRP ALA ALA ASN TYR GLN MET TYR ASP TYR ILE LEU
SEQRES  11 A  283  ASN GLU LEU PRO ARG LEU ILE GLU LYS HIS PHE PRO THR
SEQRES  12 A  283  ASN GLY LYS ARG SER ILE MET GLY HIS SER MET GLY GLY
SEQRES  13 A  283  HIS GLY ALA LEU VAL LEU ALA LEU ARG ASN GLN GLU ARG
SEQRES  14 A  283  TYR GLN SER VAL SER ALA PHE SER PRO ILE LEU SER PRO
SEQRES  15 A  283  SER LEU VAL PRO TRP GLY GLU LYS ALA PHE THR ALA TYR
SEQRES  16 A  283  LEU GLY LYS ASP ARG GLU LYS TRP GLN GLN TYR ASP ALA
SEQRES  17 A  283  ASN SER LEU ILE GLN GLN GLY TYR LYS VAL GLN GLY MET
SEQRES  18 A  283  ARG ILE ASP GLN GLY LEU GLU ASP GLU PHE LEU PRO THR
SEQRES  19 A  283  GLN LEU ARG THR GLU ASP PHE ILE GLU THR CYS ARG ALA
SEQRES  20 A  283  ALA ASN GLN PRO VAL ASP VAL ARG PHE HIS LYS GLY TYR
SEQRES  21 A  283  ASP HIS SER TYR TYR PHE ILE ALA SER PHE ILE GLY GLU
SEQRES  22 A  283  HIS ILE ALA TYR HIS ALA ALA PHE LEU LYS
SEQRES   1 B  283  GLU ASN LEU TYR PHE GLN GLY ALA MET GLU LEU ILE GLU
SEQRES   2 B  283  GLN HIS GLN ILE PHE GLY GLY SER GLN GLN VAL TRP ALA
SEQRES   3 B  283  HIS HIS ALA GLN THR LEU GLN CYS GLU MET LYS PHE ALA
SEQRES   4 B  283  VAL TYR LEU PRO ASN ASN PRO GLU ASN ARG PRO LEU GLY
SEQRES   5 B  283  VAL ILE TYR TRP LEU SER GLY LEU THR CYS THR GLU GLN
SEQRES   6 B  283  ASN PHE ILE THR LYS SER GLY PHE GLN ARG TYR ALA ALA
SEQRES   7 B  283  GLU HIS GLN VAL ILE VAL VAL ALA PRO ASP THR SER PRO
SEQRES   8 B  283  ARG GLY GLU GLN VAL PRO ASN ASP ASP ALA TYR ASP LEU
SEQRES   9 B  283  GLY GLN SER ALA GLY PHE TYR LEU ASN ALA THR GLU GLN
SEQRES  10 B  283  PRO TRP ALA ALA ASN TYR GLN MET TYR ASP TYR ILE LEU
SEQRES  11 B  283  ASN GLU LEU PRO ARG LEU ILE GLU LYS HIS PHE PRO THR
SEQRES  12 B  283  ASN GLY LYS ARG SER ILE MET GLY HIS SER MET GLY GLY
SEQRES  13 B  283  HIS GLY ALA LEU VAL LEU ALA LEU ARG ASN GLN GLU ARG
SEQRES  14 B  283  TYR GLN SER VAL SER ALA PHE SER PRO ILE LEU SER PRO
SEQRES  15 B  283  SER LEU VAL PRO TRP GLY GLU LYS ALA PHE THR ALA TYR
SEQRES  16 B  283  LEU GLY LYS ASP ARG GLU LYS TRP GLN GLN TYR ASP ALA
SEQRES  17 B  283  ASN SER LEU ILE GLN GLN GLY TYR LYS VAL GLN GLY MET
SEQRES  18 B  283  ARG ILE ASP GLN GLY LEU GLU ASP GLU PHE LEU PRO THR
SEQRES  19 B  283  GLN LEU ARG THR GLU ASP PHE ILE GLU THR CYS ARG ALA
SEQRES  20 B  283  ALA ASN GLN PRO VAL ASP VAL ARG PHE HIS LYS GLY TYR
SEQRES  21 B  283  ASP HIS SER TYR TYR PHE ILE ALA SER PHE ILE GLY GLU
SEQRES  22 B  283  HIS ILE ALA TYR HIS ALA ALA PHE LEU LYS
FORMUL   2  HOH   *726(H2 O)
HELIX    1   1 GLU A   56  SER A   63  1                                   8
HELIX    2   2 PHE A   65  GLN A   73  1                                   9
HELIX    3   3 PRO A  110  ASN A  114  5                                   5
HELIX    4   4 GLN A  116  ASN A  123  1                                   8
HELIX    5   5 ASN A  123  PHE A  133  1                                  11
HELIX    6   6 SER A  145  GLN A  159  1                                  15
HELIX    7   7 GLU A  160  TYR A  162  5                                   3
HELIX    8   8 SER A  173  LEU A  176  5                                   4
HELIX    9   9 VAL A  177  GLY A  189  1                                  13
HELIX   10  10 ASP A  191  TYR A  198  5                                   8
HELIX   11  11 ASP A  199  GLY A  207  1                                   9
HELIX   12  12 PHE A  223  LEU A  228  1                                   6
HELIX   13  13 ARG A  229  ASN A  241  1                                  13
HELIX   14  14 SER A  255  ALA A  272  1                                  18
HELIX   15  15 GLU B   56  SER B   63  1                                   8
HELIX   16  16 PHE B   65  GLN B   73  1                                   9
HELIX   17  17 PRO B  110  ASN B  114  5                                   5
HELIX   18  18 GLN B  116  ASN B  123  1                                   8
HELIX   19  19 ASN B  123  PHE B  133  1                                  11
HELIX   20  20 SER B  145  ASN B  158  1                                  14
HELIX   21  21 GLN B  159  TYR B  162  5                                   4
HELIX   22  22 SER B  173  LEU B  176  5                                   4
HELIX   23  23 VAL B  177  GLY B  189  1                                  13
HELIX   24  24 ASP B  191  TYR B  198  5                                   8
HELIX   25  25 ASP B  199  GLN B  206  1                                   8
HELIX   26  26 PHE B  223  LEU B  228  1                                   6
HELIX   27  27 ARG B  229  ASN B  241  1                                  13
HELIX   28  28 SER B  255  LEU B  274  1                                  20
SHEET    1  AA10 PHE A  -3  GLN A  -2  0
SHEET    2  AA10 MET A   1  ILE A   9 -1  O  MET A   1   N  GLN A  -2
SHEET    3  AA10 GLY A  12  ALA A  21 -1  O  GLY A  12   N  ILE A   9
SHEET    4  AA10 CYS A  26  LEU A  34 -1  O  CYS A  26   N  ALA A  21
SHEET    5  AA10 ILE A  75  PRO A  79 -1  O  VAL A  76   N  TYR A  33
SHEET    6  AA10 LEU A  43  LEU A  49  1  O  GLY A  44   N  ILE A  75
SHEET    7  AA10 THR A 135  HIS A 144  1  N  ASN A 136   O  LEU A  43
SHEET    8  AA10 VAL A 165  PHE A 168  1  O  SER A 166   N  GLY A 143
SHEET    9  AA10 ARG A 214  GLY A 218  1  O  ARG A 214   N  ALA A 167
SHEET   10  AA10 ASP A 245  HIS A 249  1  O  ASP A 245   N  ILE A 215
SHEET    1  BA 9 GLU B   2  ILE B   9  0
SHEET    2  BA 9 GLY B  12  ALA B  21 -1  O  GLY B  12   N  ILE B   9
SHEET    3  BA 9 CYS B  26  LEU B  34 -1  O  CYS B  26   N  ALA B  21
SHEET    4  BA 9 ILE B  75  PRO B  79 -1  O  VAL B  76   N  TYR B  33
SHEET    5  BA 9 LEU B  43  LEU B  49  1  O  GLY B  44   N  ILE B  75
SHEET    6  BA 9 THR B 135  HIS B 144  1  N  ASN B 136   O  LEU B  43
SHEET    7  BA 9 VAL B 165  PHE B 168  1  O  SER B 166   N  GLY B 143
SHEET    8  BA 9 ARG B 214  GLY B 218  1  O  ARG B 214   N  ALA B 167
SHEET    9  BA 9 ASP B 245  HIS B 249  1  O  ASP B 245   N  ILE B 215
CISPEP   1 GLN A  109    PRO A  110          0         6.77
CISPEP   2 GLN B  109    PRO B  110          0         3.44
CRYST1  111.268  111.268  169.861  90.00  90.00 120.00 P 65 2 2     24
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008987  0.005189  0.000000        0.00000
SCALE2      0.000000  0.010378  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005887        0.00000
TER    4494      LYS A 275
TER    8844      LYS B 275
MASTER      512    0    0   28   19    0    0    6 9568    2    0   44
END