longtext: 4BE4-pdb

content
HEADER    HYDROLASE                               05-MAR-13   4BE4
TITLE     CLOSED CONFORMATION OF O. PICEAE STEROL ESTERASE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: STEROL ESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: RESIDUES 13-549;
COMPND   5 EC: 3.1.1.13;
COMPND   6 ENGINEERED: YES;
COMPND   7 OTHER_DETAILS: N-ACETYLGLUCOSAMINE GLYCOSYLATIONS IN POSITIONS 362
COMPND   8  AND 380
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: OPHIOSTOMA PICEAE;
SOURCE   3 ORGANISM_TAXID: 61273;
SOURCE   4 EXPRESSION_SYSTEM: KOMAGATAELLA PASTORIS;
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 4922;
SOURCE   6 EXPRESSION_SYSTEM_STRAIN: KM71;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   8 EXPRESSION_SYSTEM_VECTOR: PPIC9
KEYWDS    HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.GUTIERREZ-FERNANDEZ,M.E.VAQUERO,A.PRIETO,J.BARRIUSO,M.J.GONZALEZ,
AUTHOR   2 J.A.HERMOSO
REVDAT   1   19-MAR-14 4BE4    0
JRNL        AUTH   J.GUTIERREZ-FERNANDEZ,M.E.VAQUERO,A.PRIETO,J.BARRIUSO,
JRNL        AUTH 2 M.J.GONZALEZ,J.A.HERMOSO
JRNL        TITL   CRYSTAL STRUCTURE OF OPE
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VICENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.600
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.228
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.33
REMARK   3   COMPLETENESS FOR RANGE        (%) : 97.00
REMARK   3   NUMBER OF REFLECTIONS             : 26656
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.1669
REMARK   3   R VALUE            (WORKING SET) : 0.1640
REMARK   3   FREE R VALUE                     : 0.2207
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 1344
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.2351 -  5.5990    0.93     2669   131  0.1894 0.2293
REMARK   3     2  5.5990 -  4.4453    0.97     2567   134  0.1289 0.1731
REMARK   3     3  4.4453 -  3.8837    0.98     2587   133  0.1182 0.1732
REMARK   3     4  3.8837 -  3.5287    0.98     2567   125  0.1348 0.1653
REMARK   3     5  3.5287 -  3.2759    0.97     2511   143  0.1524 0.2071
REMARK   3     6  3.2759 -  3.0828    0.98     2503   149  0.1758 0.2361
REMARK   3     7  3.0828 -  2.9284    0.98     2522   127  0.1956 0.2939
REMARK   3     8  2.9284 -  2.8010    0.97     2483   140  0.2036 0.2806
REMARK   3     9  2.8010 -  2.6931    0.97     2462   133  0.2267 0.3036
REMARK   3    10  2.6931 -  2.6002    0.96     2441   129  0.2574 0.3793
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.10
REMARK   3   SHRINKAGE RADIUS   : 0.86
REMARK   3   K_SOL              : 0.350
REMARK   3   B_SOL              : 41.977
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.63
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 22.23
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 37.51
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 5.0205
REMARK   3    B22 (A**2) : 5.0205
REMARK   3    B33 (A**2) : -10.0411
REMARK   3    B12 (A**2) : 0.0000
REMARK   3    B13 (A**2) : 0.0000
REMARK   3    B23 (A**2) : 0.0000
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           4372
REMARK   3   ANGLE     :  1.078           5933
REMARK   3   CHIRALITY :  0.073            656
REMARK   3   PLANARITY :  0.004            754
REMARK   3   DIHEDRAL  : 12.580           1577
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 6
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 11:61)
REMARK   3    ORIGIN FOR THE GROUP (A): -26.8263  40.0393 -49.2222
REMARK   3    T TENSOR
REMARK   3      T11:   0.3312 T22:   0.3679
REMARK   3      T33:   0.2106 T12:  -0.1496
REMARK   3      T13:   0.0234 T23:  -0.0577
REMARK   3    L TENSOR
REMARK   3      L11:   1.9734 L22:   2.0339
REMARK   3      L33:   1.4837 L12:   0.8762
REMARK   3      L13:  -0.4033 L23:   0.2499
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1922 S12:   0.6164 S13:  -0.1805
REMARK   3      S21:  -0.3741 S22:   0.2054 S23:   0.2020
REMARK   3      S31:   0.5010 S32:  -0.9199 S33:   0.0312
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 62:98)
REMARK   3    ORIGIN FOR THE GROUP (A): -25.8611  56.5362 -32.2748
REMARK   3    T TENSOR
REMARK   3      T11:   0.2639 T22:   0.2531
REMARK   3      T33:   0.3477 T12:   0.0036
REMARK   3      T13:   0.0297 T23:  -0.0547
REMARK   3    L TENSOR
REMARK   3      L11:   2.1636 L22:   2.3698
REMARK   3      L33:   4.0617 L12:   1.9456
REMARK   3      L13:   2.5319 L23:   1.4788
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0864 S12:  -0.0371 S13:   0.3703
REMARK   3      S21:   0.1497 S22:  -0.1763 S23:   0.2100
REMARK   3      S31:  -0.1314 S32:  -0.5511 S33:   0.2642
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 99:187)
REMARK   3    ORIGIN FOR THE GROUP (A): -20.8351  43.5877 -39.1054
REMARK   3    T TENSOR
REMARK   3      T11:   0.2316 T22:   0.2219
REMARK   3      T33:   0.1299 T12:  -0.0872
REMARK   3      T13:   0.0213 T23:  -0.0321
REMARK   3    L TENSOR
REMARK   3      L11:   2.6168 L22:   0.9809
REMARK   3      L33:   1.8221 L12:   0.5109
REMARK   3      L13:  -0.7624 L23:  -0.2305
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1051 S12:   0.1325 S13:  -0.1618
REMARK   3      S21:  -0.1750 S22:  -0.0235 S23:   0.0287
REMARK   3      S31:   0.3395 S32:  -0.4833 S33:   0.0949
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 188:277)
REMARK   3    ORIGIN FOR THE GROUP (A): -20.2922  35.2603 -31.7964
REMARK   3    T TENSOR
REMARK   3      T11:   0.3585 T22:   0.2240
REMARK   3      T33:   0.1949 T12:  -0.1150
REMARK   3      T13:   0.0551 T23:  -0.0095
REMARK   3    L TENSOR
REMARK   3      L11:   1.4298 L22:   0.8772
REMARK   3      L33:   1.8111 L12:   0.3571
REMARK   3      L13:  -0.2958 L23:  -0.3383
REMARK   3    S TENSOR
REMARK   3      S11:   0.0345 S12:  -0.1717 S13:  -0.1669
REMARK   3      S21:   0.0901 S22:  -0.0750 S23:   0.1518
REMARK   3      S31:   0.5066 S32:  -0.2880 S33:   0.0824
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 278:357)
REMARK   3    ORIGIN FOR THE GROUP (A): -26.1678  40.2221 -22.6761
REMARK   3    T TENSOR
REMARK   3      T11:   0.3468 T22:   0.4144
REMARK   3      T33:   0.2285 T12:  -0.1538
REMARK   3      T13:   0.0966 T23:   0.0113
REMARK   3    L TENSOR
REMARK   3      L11:   1.3059 L22:   0.8763
REMARK   3      L33:   0.5971 L12:   0.2902
REMARK   3      L13:  -0.1429 L23:   0.0655
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0960 S12:  -0.0473 S13:  -0.0613
REMARK   3      S21:   0.0884 S22:  -0.0210 S23:   0.1410
REMARK   3      S31:   0.4013 S32:  -0.5509 S33:   0.0449
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: CHAIN A AND (RESSEQ 358:549)
REMARK   3    ORIGIN FOR THE GROUP (A):  -7.6111  48.9759 -18.7030
REMARK   3    T TENSOR
REMARK   3      T11:   0.2178 T22:   0.2087
REMARK   3      T33:   0.1483 T12:  -0.0461
REMARK   3      T13:   0.0440 T23:   0.0315
REMARK   3    L TENSOR
REMARK   3      L11:   0.7422 L22:   0.7714
REMARK   3      L33:   1.8715 L12:  -0.2818
REMARK   3      L13:   0.0252 L23:   0.1964
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0561 S12:  -0.2449 S13:  -0.0759
REMARK   3      S21:   0.1271 S22:  -0.0192 S23:   0.0390
REMARK   3      S31:   0.2271 S32:   0.0254 S33:   0.0655
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS:  THE FIRST 8 AMINO ACIDS (EAEAYVEF)
REMARK   3  CORRESPOND TO A PURIFICATION TAG. ONLY THE LAST TWO AA (EF)
REMARK   3   WERE VISIBLE IN THE ELECTRON DENSITY MAP.
REMARK   4
REMARK   4 4BE4 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 05-MAR-13.
REMARK 100 THE PDBE ID CODE IS EBI-56045.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID29
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97908
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL (PILATUS 6M)
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 26901
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.60
REMARK 200  RESOLUTION RANGE LOW       (A) : 46.23
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 1.9
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 98.5
REMARK 200  DATA REDUNDANCY                : 3.4
REMARK 200  R MERGE                    (I) : 0.15
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 6.90
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.74
REMARK 200  COMPLETENESS FOR SHELL     (%) : 98.2
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.4
REMARK 200  R MERGE FOR SHELL          (I) : 0.82
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.90
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 65.4
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.55
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 10% 1,4-DIOXANE, 0.1 M MES PH
REMARK 280  6.5, 1.6 M AMMONIUM SULFATE
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 63 2 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z
REMARK 290       3555   -X+Y,-X,Z
REMARK 290       4555   -X,-Y,Z+1/2
REMARK 290       5555   Y,-X+Y,Z+1/2
REMARK 290       6555   X-Y,X,Z+1/2
REMARK 290       7555   Y,X,-Z
REMARK 290       8555   X-Y,-Y,-Z
REMARK 290       9555   -X,-X+Y,-Z
REMARK 290      10555   -Y,-X,-Z+1/2
REMARK 290      11555   -X+Y,Y,-Z+1/2
REMARK 290      12555   X,X-Y,-Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      103.38700
REMARK 290   SMTRY1   5  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   5 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000      103.38700
REMARK 290   SMTRY1   6  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000      103.38700
REMARK 290   SMTRY1   7 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   7  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   9 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   9 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   9  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1  10  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  10 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000      103.38700
REMARK 290   SMTRY1  11 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  11  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000      103.38700
REMARK 290   SMTRY1  12  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  12  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000      103.38700
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A     5
REMARK 465     ALA A     6
REMARK 465     GLU A     7
REMARK 465     ALA A     8
REMARK 465     TYR A     9
REMARK 465     VAL A    10
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O4   SO4 A  1567     O    HOH A  2209              2.07
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A  91      -47.73    -18.89
REMARK 500    SER A 220     -120.91     54.99
REMARK 500    SER A 252       79.56     64.38
REMARK 500    SER A 254     -107.06   -151.38
REMARK 500    SER A 312     -117.99     56.50
REMARK 500    VAL A 324      -57.44   -128.26
REMARK 500    ASP A 351       63.42   -115.64
REMARK 500    TYR A 377      -76.67   -104.21
REMARK 500    PHE A 427      -70.70   -125.66
REMARK 500    TYR A 454        0.67    -69.27
REMARK 500    PHE A 460      -29.02     65.69
REMARK 500
REMARK 500 REMARK: NULL
REMARK 600
REMARK 600 HETEROGEN
REMARK 600 N-ACETYL-D-GLUCOSAMINE (NAG): COVALENTLY BOUND TO ASN362
REMARK 600  AND ASN380 BY N-GLYCOSIDIC LINKAGES
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE DIO A1552
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE DIO A1553
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE DIO A1554
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE DIO A1555
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE DIO A1556
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE DIO A1557
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MES A1558
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A1559
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A1560
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A1561
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A1562
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A1563
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A1564
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A1565
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A1566
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SO4 A1567
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800  NAG A1550  BOUND TO ASN A 362
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800  NAG A1551  BOUND TO ASN A 380
DBREF  4BE4 A   13   549  UNP    Q2TFW1   Q2TFW1_9PEZI    13    549
SEQADV 4BE4 GLU A    5  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 ALA A    6  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 GLU A    7  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 ALA A    8  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 TYR A    9  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 VAL A   10  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 GLU A   11  UNP  Q2TFW1              EXPRESSION TAG
SEQADV 4BE4 PHE A   12  UNP  Q2TFW1              EXPRESSION TAG
SEQRES   1 A  545  GLU ALA GLU ALA TYR VAL GLU PHE THR THR VAL ASN VAL
SEQRES   2 A  545  ASN TYR PRO GLU GLY GLU VAL VAL GLY VAL SER VAL LEU
SEQRES   3 A  545  GLY ILE GLU SER PHE ARG GLY VAL PRO PHE ALA GLN PRO
SEQRES   4 A  545  PRO VAL GLY ASN LEU ARG LEU LYS PRO PRO VAL ARG TYR
SEQRES   5 A  545  THR GLU ASN ILE GLY THR LYS ASP THR THR GLY ILE GLY
SEQRES   6 A  545  PRO SER CYS PRO GLN MET TYR LEU SER THR GLY ASN GLY
SEQRES   7 A  545  GLU LEU LEU PHE GLN LEU VAL GLY ASN LEU ILE ASN ILE
SEQRES   8 A  545  PRO LEU PHE GLN THR ALA THR LEU SER SER GLU ASP CYS
SEQRES   9 A  545  LEU THR LEU ASN ILE GLN ARG PRO ALA GLY THR THR SER
SEQRES  10 A  545  ASN SER SER LEU PRO VAL LEU PHE TRP ILE PHE GLY GLY
SEQRES  11 A  545  GLY PHE GLU LEU GLY THR ASN GLN TYR TYR ASP GLY ILE
SEQRES  12 A  545  ASP LEU LEU THR GLU GLY ILE SER LEU GLY GLU PRO PHE
SEQRES  13 A  545  ILE PHE VAL ALA ILE ASN TYR ARG VAL GLY GLY PHE GLY
SEQRES  14 A  545  PHE LEU GLY GLY LYS GLU ILE LYS ALA ASP GLY SER SER
SEQRES  15 A  545  ASN LEU GLY LEU LEU ASP GLN ARG ILE ALA LEU GLU TRP
SEQRES  16 A  545  VAL ALA ASP ASN ILE ALA SER PHE GLY GLY ASP PRO SER
SEQRES  17 A  545  LYS VAL THR ILE TRP GLY GLU SER ALA GLY SER ILE SER
SEQRES  18 A  545  VAL PHE ASP GLN MET ALA LEU TYR GLY GLY ASN ASN LYS
SEQRES  19 A  545  TYR LYS GLY LYS ALA LEU PHE ARG GLY GLY ILE MET ASN
SEQRES  20 A  545  SER GLY SER VAL VAL PRO ALA ALA PRO VAL ASP GLY VAL
SEQRES  21 A  545  LYS ALA GLN ALA ILE TYR ASP HIS VAL VAL SER GLU ALA
SEQRES  22 A  545  GLY CYS ALA GLY THR SER ASP THR LEU ALA CYS LEU ARG
SEQRES  23 A  545  THR VAL ASP TYR THR LYS PHE LEU THR ALA VAL ASN SER
SEQRES  24 A  545  VAL PRO GLY ILE VAL SER TYR SER SER ILE ALA LEU SER
SEQRES  25 A  545  TYR LEU PRO ARG PRO ASP GLY VAL VAL LEU ILE ASP SER
SEQRES  26 A  545  PRO GLU GLU ILE VAL LYS ASN LYS GLN TYR ALA ALA VAL
SEQRES  27 A  545  PRO MET ILE ILE GLY ASP GLN GLU ASP GLU GLY THR LEU
SEQRES  28 A  545  PHE ALA VAL LEU PRO ASN ASN ILE THR SER THR ALA LYS
SEQRES  29 A  545  ILE VAL GLN TYR PHE GLN ASP LEU TYR PHE TYR ASN ALA
SEQRES  30 A  545  THR LYS GLU GLN LEU THR ALA PHE VAL ASN THR TYR PRO
SEQRES  31 A  545  THR ASP ILE THR ALA GLY SER PRO PHE ASN THR GLY ILE
SEQRES  32 A  545  PHE ASN GLU LEU TYR PRO GLY PHE LYS ARG LEU ALA ALA
SEQRES  33 A  545  ILE LEU GLY ASP MET THR PHE THR LEU ALA ARG ARG ALA
SEQRES  34 A  545  PHE LEU GLN LEU CYS SER GLU VAL ASN PRO ASP VAL PRO
SEQRES  35 A  545  SER TRP SER TYR LEU ALA SER TYR ASP TYR GLY PHE PRO
SEQRES  36 A  545  PHE LEU GLY THR PHE HIS ALA THR ASP ILE LEU GLN VAL
SEQRES  37 A  545  PHE TYR GLY VAL LEU PRO ASN TYR ALA SER GLY SER ILE
SEQRES  38 A  545  GLN LYS TYR TYR ILE ASN PHE VAL THR THR GLY ASP PRO
SEQRES  39 A  545  ASN LYS GLY ALA ALA VAL ASP ILE GLN TRP PRO GLN TRP
SEQRES  40 A  545  SER ALA LYS LYS ASN ILE LEU GLN ILE TYR ALA THR LYS
SEQRES  41 A  545  ALA VAL ILE VAL ALA ASP ASN PHE ARG ALA LYS SER TYR
SEQRES  42 A  545  GLU TYR LEU TYR ASN ASN ILE GLY ILE PHE ARG ILE
HET    NAG  A1550      14
HET    NAG  A1551      14
HET    DIO  A1552       6
HET    DIO  A1553       6
HET    DIO  A1554       6
HET    DIO  A1555       6
HET    DIO  A1556       6
HET    DIO  A1557       6
HET    MES  A1558      12
HET    GOL  A1559       6
HET    GOL  A1560       6
HET    GOL  A1561       6
HET    GOL  A1562       6
HET    GOL  A1563       6
HET    GOL  A1564       6
HET    SO4  A1565       5
HET    SO4  A1566       5
HET    SO4  A1567       5
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     DIO 1,4-DIETHYLENE DIOXIDE
HETNAM     MES 2-(N-MORPHOLINO)-ETHANESULFONIC ACID
HETNAM     GOL GLYCEROL
HETNAM     SO4 SULFATE ION
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
FORMUL   2  NAG    2(C8 H15 N O6)
FORMUL   3  DIO    6(C4 H8 O2)
FORMUL   4  MES    C6 H13 N O4 S
FORMUL   5  GOL    6(C3 H8 O3)
FORMUL   6  SO4    3(O4 S 2-)
FORMUL   7  HOH   *242(H2 O)
HELIX    1   1 VAL A   45  ARG A   49  5                                   5
HELIX    2   2 GLY A   82  VAL A   89  1                                   8
HELIX    3   3 GLY A   90  ILE A   95  5                                   6
HELIX    4   4 ILE A   95  GLN A   99  5                                   5
HELIX    5   5 THR A  140  TYR A  144  5                                   5
HELIX    6   6 GLY A  146  LEU A  156  1                                  11
HELIX    7   7 GLY A  170  LEU A  175  1                                   6
HELIX    8   8 GLY A  177  ASP A  183  1                                   7
HELIX    9   9 ASN A  187  ILE A  204  1                                  18
HELIX   10  10 ALA A  205  PHE A  207  5                                   3
HELIX   11  11 SER A  220  ASN A  236  1                                  17
HELIX   12  12 GLY A  263  GLY A  278  1                                  16
HELIX   13  13 ASP A  284  ARG A  290  1                                   7
HELIX   14  14 ASP A  293  SER A  303  1                                  11
HELIX   15  15 SER A  329  ASN A  336  1                                   8
HELIX   16  16 GLY A  353  VAL A  358  1                                   6
HELIX   17  17 SER A  365  TYR A  377  1                                  13
HELIX   18  18 THR A  382  ASN A  391  1                                  10
HELIX   19  19 GLY A  414  PHE A  427  1                                  14
HELIX   20  20 PHE A  427  ASN A  442  1                                  16
HELIX   21  21 ASP A  468  PHE A  473  1                                   6
HELIX   22  22 TYR A  480  GLY A  496  1                                  17
HELIX   23  23 GLN A  510  LYS A  515  1                                   6
HELIX   24  24 ARG A  533  ASN A  543  1                                  11
HELIX   25  25 ILE A  544  ARG A  548  5                                   5
SHEET    1  AA 2 PHE A  12  TYR A  19  0
SHEET    2  AA 2 GLY A  22  VAL A  29 -1  O  GLY A  22   N  TYR A  19
SHEET    1  AB 2 THR A  62  ASP A  64  0
SHEET    2  AB 2 GLY A  22  VAL A  29  1  O  GLU A  23   N  LYS A  63
SHEET    1  AC12 ALA A 525  ALA A 529  0
SHEET    2  AC12 ASN A 516  ILE A 520 -1  O  ILE A 517   N  VAL A 528
SHEET    3  AC12 SER A 447  ALA A 452  1  O  SER A 449   N  LEU A 518
SHEET    4  AC12 MET A 344  GLN A 349  1  O  MET A 344   N  TRP A 448
SHEET    5  AC12 GLY A 247  ASN A 251  1  O  GLY A 248   N  ILE A 345
SHEET    6  AC12 GLY A 209  GLU A 219  1  O  ILE A 216   N  ILE A 249
SHEET    7  AC12 LEU A 125  ILE A 131  1  O  LEU A 125   N  ASP A 210
SHEET    8  AC12 ILE A 161  ILE A 165  1  O  ILE A 161   N  LEU A 128
SHEET    9  AC12 THR A 110  PRO A 116 -1  O  ASN A 112   N  ALA A 164
SHEET   10  AC12 ILE A  32  PRO A  39 -1  O  GLU A  33   N  ARG A 115
SHEET   11  AC12 GLY A  22  VAL A  29 -1  O  VAL A  27   N  SER A  34
SHEET   12  AC12 THR A  62  ASP A  64  1  O  LYS A  63   N  VAL A  25
SHEET    1  AD12 ALA A 525  ALA A 529  0
SHEET    2  AD12 ASN A 516  ILE A 520 -1  O  ILE A 517   N  VAL A 528
SHEET    3  AD12 SER A 447  ALA A 452  1  O  SER A 449   N  LEU A 518
SHEET    4  AD12 MET A 344  GLN A 349  1  O  MET A 344   N  TRP A 448
SHEET    5  AD12 GLY A 247  ASN A 251  1  O  GLY A 248   N  ILE A 345
SHEET    6  AD12 GLY A 209  GLU A 219  1  O  ILE A 216   N  ILE A 249
SHEET    7  AD12 LEU A 125  ILE A 131  1  O  LEU A 125   N  ASP A 210
SHEET    8  AD12 ILE A 161  ILE A 165  1  O  ILE A 161   N  LEU A 128
SHEET    9  AD12 THR A 110  PRO A 116 -1  O  ASN A 112   N  ALA A 164
SHEET   10  AD12 ILE A  32  PRO A  39 -1  O  GLU A  33   N  ARG A 115
SHEET   11  AD12 GLY A  22  VAL A  29 -1  O  VAL A  27   N  SER A  34
SHEET   12  AD12 PHE A  12  TYR A  19 -1  O  THR A  13   N  SER A  28
SHEET    1  AE 2 LYS A 238  TYR A 239  0
SHEET    2  AE 2 LYS A 242  ALA A 243 -1  O  LYS A 242   N  TYR A 239
SSBOND   1 CYS A   72    CYS A  108                          1555   1555  2.04
SSBOND   2 CYS A  279    CYS A  288                          1555   1555  2.05
LINK         ND2 ASN A 362                 C1  NAG A1550     1555   1555  1.55
LINK         ND2 ASN A 380                 C1  NAG A1551     1555   1555  1.56
CISPEP   1 SER A  401    PRO A  402          0         6.23
SITE     1 AC1  4 GLY A 118  THR A 120  SER A 123  GLY A 501
SITE     1 AC2  3 GLU A 198  TYR A 233  HOH A2033
SITE     1 AC3  4 THR A 120  SER A 121  HOH A2004  HOH A2005
SITE     1 AC4  3 ALA A 117  LEU A 156  GLY A 501
SITE     1 AC5  4 PHE A 378  THR A 426  PHE A 547  ILE A 549
SITE     1 AC6  5 SER A 220  ILE A 224  ILE A 307  ILE A 313
SITE     2 AC6  5 LEU A 315
SITE     1 AC7  7 LEU A  92  TYR A 454  ASP A 455  GLN A 471
SITE     2 AC7  7 ASN A 479  ALA A 522  THR A 523
SITE     1 AC8  1 ASN A  94
SITE     1 AC9  3 ASN A  81  GLY A 306  SER A 309
SITE     1 BC1  2 ARG A 533  ALA A 534
SITE     1 BC2  2 TYR A 239  LYS A 240
SITE     1 BC3  4 TYR A 480  GLY A 483  LYS A 487  HOH A2216
SITE     1 BC4  3 VAL A 504  ASP A 505  HOH A2216
SITE     1 BC5  3 SER A 365  THR A 366  SO4 A1566
SITE     1 BC6  3 SER A 365  LYS A 368  SO4 A1565
SITE     1 BC7  5 TYR A 521  ALA A 522  THR A 523  LYS A 524
SITE     2 BC7  5 HOH A2209
SITE     1 BC8  3 TYR A 310  ASN A 362  HOH A2240
SITE     1 BC9  5 ALA A 259  ASN A 380  GLU A 384  ARG A 548
SITE     2 BC9  5 HOH A2241
CRYST1  119.274  119.274  206.774  90.00  90.00 120.00 P 63 2 2     12
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.008384  0.004841  0.000000        0.00000
SCALE2      0.000000  0.009681  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004836        0.00000
TER    4141      ILE A 549
MASTER      452    0   18   25   30    0   22    6 4508    1  133   42
END