longtext: 4BYM-pdb

content
HEADER    HYDROLASE                               20-JUL-13   4BYM
TITLE     STRUCTURE OF PHAZ7 PHB DEPOLYMERASE Y105E MUTANT
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PHB DEPOLYMERASE PHAZ7;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: PHB DEPOLYMERASE;
COMPND   5 EC: 3.1.1.75;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PAUCIMONAS LEMOIGNEI;
SOURCE   3 ORGANISM_TAXID: 29443;
SOURCE   4 EXPRESSION_SYSTEM: BACILLUS SUBTILIS;
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 1423;
SOURCE   6 EXPRESSION_SYSTEM_STRAIN: WB800
KEYWDS    HYDROLASE, CATALYTIC TRIAD, SUBSTRATE-BINDING CAVITY, BINDING
KEYWDS   2 CAVITY, 3HB TRIMER BINDING
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.HERMAWAN,B.SUBEDI,A.C.PAPAGEORGIOU,D.JENDROSSEK
REVDAT   1   18-SEP-13 4BYM    0
JRNL        AUTH   D.JENDROSSEK,S.HERMAWAN,B.SUBEDI,A.C.PAPAGEORGIOU
JRNL        TITL   BIOCHEMICAL ANALYSIS AND STRUCTURE DETERMINATION OF
JRNL        TITL 2 PAUCIMONAS LEMOIGNEI POLY(3-HYDROXYBUTYRATE) (PHB)
JRNL        TITL 3 DEPOLYMERASE PHAZ7 MUTEINS REVEAL THE PHB BINDING SITE AND
JRNL        TITL 4 DETAILS OF SUBSTRATE-ENZYME INTERACTIONS.
JRNL        REF    MOL.MICROBIOL.                             2013
JRNL        REFN                   ESSN 1365-2958
JRNL        PMID   24007310
JRNL        DOI    10.1111/MMI.12391
REMARK   2
REMARK   2 RESOLUTION.    1.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VICENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : MLHL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.598
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.843
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.99
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.13
REMARK   3   NUMBER OF REFLECTIONS             : 85301
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.1497
REMARK   3   R VALUE            (WORKING SET) : 0.1484
REMARK   3   FREE R VALUE                     : 0.1743
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 4266
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 19.8448 -  4.9409    0.95     2663   141  0.1370 0.1465
REMARK   3     2  4.9409 -  3.9317    0.96     2661   140  0.1158 0.1426
REMARK   3     3  3.9317 -  3.4377    0.97     2655   140  0.1241 0.1385
REMARK   3     4  3.4377 -  3.1247    0.98     2709   142  0.1370 0.1412
REMARK   3     5  3.1247 -  2.9015    0.98     2692   142  0.1430 0.1575
REMARK   3     6  2.9015 -  2.7308    0.98     2713   143  0.1416 0.1626
REMARK   3     7  2.7308 -  2.5944    0.99     2689   141  0.1428 0.1691
REMARK   3     8  2.5944 -  2.4817    0.98     2730   144  0.1420 0.1520
REMARK   3     9  2.4817 -  2.3863    0.98     2729   143  0.1442 0.1700
REMARK   3    10  2.3863 -  2.3041    0.99     2702   143  0.1462 0.1760
REMARK   3    11  2.3041 -  2.2321    0.98     2728   143  0.1449 0.1868
REMARK   3    12  2.2321 -  2.1684    0.99     2747   145  0.1437 0.1726
REMARK   3    13  2.1684 -  2.1114    0.99     2687   141  0.1438 0.1985
REMARK   3    14  2.1114 -  2.0599    0.99     2747   145  0.1481 0.1850
REMARK   3    15  2.0599 -  2.0132    0.99     2710   142  0.1479 0.2015
REMARK   3    16  2.0132 -  1.9703    1.00     2755   145  0.1494 0.1535
REMARK   3    17  1.9703 -  1.9310    0.99     2722   144  0.1554 0.1912
REMARK   3    18  1.9310 -  1.8946    1.00     2749   144  0.1605 0.2048
REMARK   3    19  1.8946 -  1.8607    0.99     2666   141  0.1665 0.2145
REMARK   3    20  1.8607 -  1.8292    0.99     2823   148  0.1641 0.2091
REMARK   3    21  1.8292 -  1.7997    1.00     2668   141  0.1683 0.1954
REMARK   3    22  1.7997 -  1.7721    0.99     2762   145  0.1675 0.1813
REMARK   3    23  1.7721 -  1.7460    0.99     2768   146  0.1702 0.1884
REMARK   3    24  1.7460 -  1.7214    1.00     2661   140  0.1727 0.2304
REMARK   3    25  1.7214 -  1.6982    0.99     2786   147  0.1824 0.1976
REMARK   3    26  1.6982 -  1.6762    1.00     2738   144  0.1822 0.2235
REMARK   3    27  1.6762 -  1.6552    1.00     2694   141  0.1927 0.2322
REMARK   3    28  1.6552 -  1.6353    0.99     2813   148  0.2054 0.2158
REMARK   3    29  1.6353 -  1.6163    1.00     2707   143  0.2215 0.2815
REMARK   3    30  1.6163 -  1.5981    0.80     2161   114  0.2452 0.2893
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.16
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 16.24
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 19
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 13.8
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.008           5335
REMARK   3   ANGLE     :  1.037           7261
REMARK   3   CHIRALITY :  0.070            768
REMARK   3   PLANARITY :  0.005            943
REMARK   3   DIHEDRAL  : 12.199           1843
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4BYM COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 22-JUL-13.
REMARK 100 THE PDBE ID CODE IS EBI-57355.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 19-NOV-10
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : EMBL/DESY, HAMBURG
REMARK 200  BEAMLINE                       : X13
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.8123
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 85362
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.60
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.00
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.8
REMARK 200  DATA REDUNDANCY                : 3.1
REMARK 200  R MERGE                    (I) : 0.05
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 16.30
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.69
REMARK 200  COMPLETENESS FOR SHELL     (%) : 95.5
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.9
REMARK 200  R MERGE FOR SHELL          (I) : 0.37
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.90
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: PDB ENTRY 4BRS
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 43
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.3
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: HANGING DROP VAPOUR DIFFUSION.
REMARK 280  PROTEIN CONCENTRATION 10 MG/ML. 0.1 M NAOAC (PH 5.0), 0.2
REMARK 280  M LICL, 18% W/V PEG6000
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000      100.17000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     TYR A 293    CG   CD1  CD2  CE1  CE2  CZ   OH
REMARK 470     ASN A 294    CG   OD1  ND2
REMARK 470     ASN B 294    CG   OD1  ND2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OG   SER A   136     O    HOH A  2158              2.16
REMARK 500   OH   TYR B    66     O    HOH B  2216              2.20
REMARK 500   OG   SER B   136     O    HOH B  2177              2.02
REMARK 500   O    TYR B   293     O    HOH B  2577              2.00
REMARK 500   OE1A GLU B   333     O    HOH A  2129              2.16
REMARK 500   OE2B GLU B   333     O    HOH A  2129              1.94
REMARK 500   O    HOH A  2046     O    HOH A  2100              1.88
REMARK 500   O    HOH A  2066     O    HOH A  2540              2.15
REMARK 500   O    HOH A  2112     O    HOH A  2113              2.14
REMARK 500   O    HOH A  2120     O    HOH A  2121              2.05
REMARK 500   O    HOH A  2121     O    HOH A  2244              2.15
REMARK 500   O    HOH A  2141     O    HOH A  2265              2.05
REMARK 500   O    HOH A  2158     O    HOH A  2339              1.97
REMARK 500   O    HOH A  2159     O    HOH A  2160              2.10
REMARK 500   O    HOH A  2179     O    HOH A  2180              2.10
REMARK 500   O    HOH A  2180     O    HOH A  2346              2.03
REMARK 500   O    HOH A  2298     O    HOH A  2535              2.06
REMARK 500   O    HOH A  2321     O    HOH A  2322              2.17
REMARK 500   O    HOH A  2321     O    HOH A  2572              2.19
REMARK 500   O    HOH A  2325     O    HOH A  2326              2.13
REMARK 500   O    HOH A  2357     O    HOH A  2540              2.13
REMARK 500   O    HOH A  2364     O    HOH A  2365              2.12
REMARK 500   O    HOH A  2448     O    HOH A  2449              1.99
REMARK 500   O    HOH A  2539     O    HOH A  2540              2.19
REMARK 500   O    HOH A  2565     O    HOH A  2567              2.08
REMARK 500   O    HOH A  2575     O    HOH A  2578              2.12
REMARK 500   O    HOH B  2011     O    HOH B  2074              2.03
REMARK 500   O    HOH B  2017     O    HOH B  2025              2.14
REMARK 500   O    HOH B  2020     O    HOH B  2588              2.00
REMARK 500   O    HOH B  2037     O    HOH B  2059              2.11
REMARK 500   O    HOH B  2038     O    HOH B  2087              2.16
REMARK 500   O    HOH B  2082     O    HOH B  2147              2.10
REMARK 500   O    HOH B  2083     O    HOH B  2163              2.19
REMARK 500   O    HOH B  2142     O    HOH B  2282              2.14
REMARK 500   O    HOH B  2142     O    HOH B  2158              2.17
REMARK 500   O    HOH B  2153     O    HOH B  2154              1.97
REMARK 500   O    HOH B  2166     O    HOH B  2373              2.14
REMARK 500   O    HOH B  2177     O    HOH B  2380              2.15
REMARK 500   O    HOH B  2180     O    HOH B  2361              2.11
REMARK 500   O    HOH B  2194     O    HOH B  2388              1.98
REMARK 500   O    HOH B  2203     O    HOH B  2402              1.96
REMARK 500   O    HOH B  2251     O    HOH A  2315              2.16
REMARK 500   O    HOH B  2275     O    HOH B  2282              2.14
REMARK 500   O    HOH B  2293     O    HOH B  2627              1.85
REMARK 500   O    HOH B  2293     O    HOH B  2628              2.09
REMARK 500   O    HOH B  2299     O    HOH B  2537              2.08
REMARK 500   O    HOH B  2299     O    HOH B  2534              2.19
REMARK 500   O    HOH B  2337     O    HOH B  2341              2.07
REMARK 500   O    HOH B  2389     O    HOH B  2390              2.03
REMARK 500   O    HOH B  2460     O    HOH B  2467              1.97
REMARK 500
REMARK 500 THIS ENTRY HAS      53 CLOSE CONTACTS
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O    HOH A  2117     O    HOH B  2362     1656     2.08
REMARK 500   O    HOH B  2045     O    HOH B  2200     1655     2.04
REMARK 500   O    HOH B  2190     O    HOH B  2630     1455     2.20
REMARK 500   O    HOH B  2214     O    HOH B  2339     1656     2.18
REMARK 500   O    HOH B  2529     O    HOH B  2584     1455     2.03
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A 136     -129.83     54.75
REMARK 500    ARG A 156      -61.25    -95.75
REMARK 500    ALA A 177       45.89   -142.96
REMARK 500    SER A 186     -157.93    -81.56
REMARK 500    TYR A 204     -115.40     54.13
REMARK 500    ASN A 210       83.25   -155.53
REMARK 500    ALA B  23       84.77   -151.99
REMARK 500    SER B 136     -133.00     58.21
REMARK 500    ARG B 156      -61.84    -92.01
REMARK 500    ALA B 177       40.07   -142.02
REMARK 500    ALA B 181       65.04   -150.25
REMARK 500    SER B 186     -155.83    -83.42
REMARK 500    TYR B 204     -119.27     53.31
REMARK 500    ASN B 210       81.65   -156.74
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A2142        DISTANCE =  5.18 ANGSTROMS
REMARK 525    HOH A2575        DISTANCE =  6.92 ANGSTROMS
REMARK 525    HOH A2576        DISTANCE =  5.94 ANGSTROMS
REMARK 525    HOH A2578        DISTANCE =  7.96 ANGSTROMS
REMARK 525    HOH B2062        DISTANCE =  6.78 ANGSTROMS
REMARK 525    HOH B2152        DISTANCE =  6.20 ANGSTROMS
REMARK 525    HOH B2265        DISTANCE =  5.98 ANGSTROMS
REMARK 525    HOH B2345        DISTANCE =  6.83 ANGSTROMS
REMARK 525    HOH B2369        DISTANCE =  6.89 ANGSTROMS
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA A1343  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 LYS A 122   NZ
REMARK 620 2 HOH A2311   O    62.3
REMARK 620 3 HOH A2574   O   113.2 107.2
REMARK 620 4 HOH A2329   O   124.7 111.0 120.3
REMARK 620 5 SER A 128   O    89.7 143.8 104.7  65.1
REMARK 620 N                    1     2     3     4
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA B1344  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLY B 167   O
REMARK 620 2 HOH B2416   O    72.9
REMARK 620 3 HOH B2418   O   155.2 126.7
REMARK 620 N                    1     2
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL B1343
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  NA B1344
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  NA A1343
DBREF  4BYM A    1   342  UNP    Q939Q9   Q939Q9_PSELE    39    380
DBREF  4BYM B    1   342  UNP    Q939Q9   Q939Q9_PSELE    39    380
SEQADV 4BYM GLU A  105  UNP  Q939Q9    TYR   143 ENGINEERED MUTATION
SEQADV 4BYM GLU B  105  UNP  Q939Q9    TYR   143 ENGINEERED MUTATION
SEQRES   1 A  342  LEU THR CYS GLY THR ASN SER GLY PHE VAL CYS LYS GLY
SEQRES   2 A  342  THR GLN THR GLN TYR ALA GLY GLY PHE ALA PRO GLY VAL
SEQRES   3 A  342  GLY TYR GLY GLY PHE GLY GLY GLY SER CYS THR ALA THR
SEQRES   4 A  342  LYS THR PRO VAL ILE PHE ILE HIS GLY ASN GLY ASP ASN
SEQRES   5 A  342  ALA ILE SER PHE ASP MET PRO PRO GLY ASN VAL SER GLY
SEQRES   6 A  342  TYR GLY THR PRO ALA ARG SER VAL TYR ALA GLU LEU LYS
SEQRES   7 A  342  ALA ARG GLY TYR ASN ASP CYS GLU ILE PHE GLY VAL THR
SEQRES   8 A  342  TYR LEU SER SER SER GLU GLN GLY SER ALA GLN TYR ASN
SEQRES   9 A  342  GLU HIS SER SER THR LYS TYR ALA ILE ILE LYS THR PHE
SEQRES  10 A  342  ILE ASP LYS VAL LYS ALA TYR THR GLY LYS SER GLN VAL
SEQRES  11 A  342  ASP ILE VAL ALA HIS SER MET GLY VAL SER MET SER LEU
SEQRES  12 A  342  ALA THR LEU GLN TYR TYR ASN ASN TRP THR SER VAL ARG
SEQRES  13 A  342  LYS PHE ILE ASN LEU ALA GLY GLY ILE ARG GLY LEU TYR
SEQRES  14 A  342  SER CYS TYR TYR THR GLY TYR ALA ASN ALA ALA ALA PRO
SEQRES  15 A  342  THR CYS GLY SER GLN ASN TYR TYR ASN SER TYR THR PHE
SEQRES  16 A  342  GLY PHE PHE PRO GLU GLY TRP TYR TYR GLY VAL TRP VAL
SEQRES  17 A  342  SER ASN PRO TRP THR GLY SER GLY SER THR ASN SER MET
SEQRES  18 A  342  ARG ASP MET PRO ALA LYS ARG THR ALA VAL SER PHE TYR
SEQRES  19 A  342  THR LEU SER ALA GLY PHE LYS ASP GLN VAL GLY CYS ALA
SEQRES  20 A  342  THR ALA SER PHE TRP ALA GLY CYS ASP SER ALA ALA LYS
SEQRES  21 A  342  PHE ALA SER THR THR SER ASN VAL LYS ALA GLN ILE ASN
SEQRES  22 A  342  VAL GLY ALA GLY SER ASN ALA THR GLN ALA ASP TYR ASP
SEQRES  23 A  342  TRP ALA ASP GLY MET PRO TYR ASN ALA GLY GLY GLY ASP
SEQRES  24 A  342  THR THR ASN GLY VAL GLY HIS PHE ARG THR LYS THR ASN
SEQRES  25 A  342  THR GLY ALA ILE ILE GLN ARG MET LEU LEU THR THR CYS
SEQRES  26 A  342  THR GLY LEU ASP CYS ALA ALA GLU TYR THR THR GLY PRO
SEQRES  27 A  342  LYS ALA ALA TYR
SEQRES   1 B  342  LEU THR CYS GLY THR ASN SER GLY PHE VAL CYS LYS GLY
SEQRES   2 B  342  THR GLN THR GLN TYR ALA GLY GLY PHE ALA PRO GLY VAL
SEQRES   3 B  342  GLY TYR GLY GLY PHE GLY GLY GLY SER CYS THR ALA THR
SEQRES   4 B  342  LYS THR PRO VAL ILE PHE ILE HIS GLY ASN GLY ASP ASN
SEQRES   5 B  342  ALA ILE SER PHE ASP MET PRO PRO GLY ASN VAL SER GLY
SEQRES   6 B  342  TYR GLY THR PRO ALA ARG SER VAL TYR ALA GLU LEU LYS
SEQRES   7 B  342  ALA ARG GLY TYR ASN ASP CYS GLU ILE PHE GLY VAL THR
SEQRES   8 B  342  TYR LEU SER SER SER GLU GLN GLY SER ALA GLN TYR ASN
SEQRES   9 B  342  GLU HIS SER SER THR LYS TYR ALA ILE ILE LYS THR PHE
SEQRES  10 B  342  ILE ASP LYS VAL LYS ALA TYR THR GLY LYS SER GLN VAL
SEQRES  11 B  342  ASP ILE VAL ALA HIS SER MET GLY VAL SER MET SER LEU
SEQRES  12 B  342  ALA THR LEU GLN TYR TYR ASN ASN TRP THR SER VAL ARG
SEQRES  13 B  342  LYS PHE ILE ASN LEU ALA GLY GLY ILE ARG GLY LEU TYR
SEQRES  14 B  342  SER CYS TYR TYR THR GLY TYR ALA ASN ALA ALA ALA PRO
SEQRES  15 B  342  THR CYS GLY SER GLN ASN TYR TYR ASN SER TYR THR PHE
SEQRES  16 B  342  GLY PHE PHE PRO GLU GLY TRP TYR TYR GLY VAL TRP VAL
SEQRES  17 B  342  SER ASN PRO TRP THR GLY SER GLY SER THR ASN SER MET
SEQRES  18 B  342  ARG ASP MET PRO ALA LYS ARG THR ALA VAL SER PHE TYR
SEQRES  19 B  342  THR LEU SER ALA GLY PHE LYS ASP GLN VAL GLY CYS ALA
SEQRES  20 B  342  THR ALA SER PHE TRP ALA GLY CYS ASP SER ALA ALA LYS
SEQRES  21 B  342  PHE ALA SER THR THR SER ASN VAL LYS ALA GLN ILE ASN
SEQRES  22 B  342  VAL GLY ALA GLY SER ASN ALA THR GLN ALA ASP TYR ASP
SEQRES  23 B  342  TRP ALA ASP GLY MET PRO TYR ASN ALA GLY GLY GLY ASP
SEQRES  24 B  342  THR THR ASN GLY VAL GLY HIS PHE ARG THR LYS THR ASN
SEQRES  25 B  342  THR GLY ALA ILE ILE GLN ARG MET LEU LEU THR THR CYS
SEQRES  26 B  342  THR GLY LEU ASP CYS ALA ALA GLU TYR THR THR GLY PRO
SEQRES  27 B  342  LYS ALA ALA TYR
HET     CL  B1343       1
HET     NA  B1344       1
HET     NA  A1343       1
HETNAM      CL CHLORIDE ION
HETNAM      NA SODIUM ION
FORMUL   3   CL    CL 1-
FORMUL   4   NA    2(NA 1+)
FORMUL   5  HOH   *1210(H2 O)
HELIX    1   1 ASN A   52  MET A   58  5                                   7
HELIX    2   2 SER A   72  ARG A   80  1                                   9
HELIX    3   3 SER A   94  GLY A   99  1                                   6
HELIX    4   4 SER A  100  ASN A  104  5                                   5
HELIX    5   5 SER A  107  GLY A  126  1                                  20
HELIX    6   6 SER A  136  ASN A  150  1                                  15
HELIX    7   7 ASN A  151  THR A  153  5                                   3
HELIX    8   8 LEU A  168  TYR A  172  5                                   5
HELIX    9   9 ALA A  181  GLY A  185  5                                   5
HELIX   10  10 SER A  220  ARG A  222  5                                   3
HELIX   11  11 ASP A  223  ARG A  228  1                                   6
HELIX   12  12 ALA A  238  ASP A  242  5                                   5
HELIX   13  13 GLY A  254  ALA A  258  5                                   5
HELIX   14  14 ASP A  286  GLY A  290  5                                   5
HELIX   15  15 PHE A  307  ASN A  312  1                                   6
HELIX   16  16 THR A  313  THR A  323  1                                  11
HELIX   17  17 THR A  326  ALA A  332  5                                   7
HELIX   18  18 ASN B   52  MET B   58  5                                   7
HELIX   19  19 SER B   72  ARG B   80  1                                   9
HELIX   20  20 SER B   94  GLY B   99  1                                   6
HELIX   21  21 SER B  100  ASN B  104  5                                   5
HELIX   22  22 SER B  107  GLY B  126  1                                  20
HELIX   23  23 SER B  136  ASN B  150  1                                  15
HELIX   24  24 ASN B  151  THR B  153  5                                   3
HELIX   25  25 LEU B  168  TYR B  172  5                                   5
HELIX   26  26 ALA B  181  GLY B  185  5                                   5
HELIX   27  27 SER B  220  ARG B  228  5                                   9
HELIX   28  28 ALA B  238  ASP B  242  5                                   5
HELIX   29  29 GLY B  254  ALA B  258  5                                   5
HELIX   30  30 ASP B  286  GLY B  290  5                                   5
HELIX   31  31 PHE B  307  ASN B  312  1                                   6
HELIX   32  32 THR B  313  THR B  323  1                                  11
HELIX   33  33 THR B  326  ALA B  332  5                                   7
SHEET    1  AA 9 THR A  16  TYR A  18  0
SHEET    2  AA 9 GLY A  30  GLY A  32  1  O  GLY A  30   N  GLN A  17
SHEET    3  AA 9 ILE A  87  VAL A  90 -1  O  GLY A  89   N  PHE A  31
SHEET    4  AA 9 VAL A  43  ILE A  46  1  O  VAL A  43   N  PHE A  88
SHEET    5  AA 9 VAL A 130  HIS A 135  1  O  ASP A 131   N  ILE A  44
SHEET    6  AA 9 VAL A 155  LEU A 161  1  N  ARG A 156   O  VAL A 130
SHEET    7  AA 9 SER A 232  SER A 237  1  O  SER A 232   N  PHE A 158
SHEET    8  AA 9 VAL A 268  ASN A 273  1  N  LYS A 269   O  PHE A 233
SHEET    9  AA 9 ALA A 340  ALA A 341 -1  O  ALA A 340   N  ASN A 273
SHEET    1  AB 2 GLY A 201  TYR A 203  0
SHEET    2  AB 2 VAL A 206  VAL A 208 -1  O  VAL A 206   N  TYR A 203
SHEET    1  BA 9 THR B  16  TYR B  18  0
SHEET    2  BA 9 GLY B  30  GLY B  32  1  O  GLY B  30   N  GLN B  17
SHEET    3  BA 9 ILE B  87  VAL B  90 -1  O  GLY B  89   N  PHE B  31
SHEET    4  BA 9 VAL B  43  ILE B  46  1  O  VAL B  43   N  PHE B  88
SHEET    5  BA 9 VAL B 130  HIS B 135  1  O  ASP B 131   N  ILE B  44
SHEET    6  BA 9 VAL B 155  LEU B 161  1  N  ARG B 156   O  VAL B 130
SHEET    7  BA 9 SER B 232  SER B 237  1  O  SER B 232   N  PHE B 158
SHEET    8  BA 9 VAL B 268  ASN B 273  1  N  LYS B 269   O  PHE B 233
SHEET    9  BA 9 ALA B 340  ALA B 341 -1  O  ALA B 340   N  ASN B 273
SHEET    1  BB 2 GLY B 201  TYR B 203  0
SHEET    2  BB 2 VAL B 206  VAL B 208 -1  O  VAL B 206   N  TYR B 203
SSBOND   1 CYS A    3    CYS A   11                          1555   1555  2.03
SSBOND   2 CYS A   36    CYS A   85                          1555   1555  2.04
SSBOND   3 CYS A  171    CYS A  184                          1555   1555  2.04
SSBOND   4 CYS A  246    CYS A  255                          1555   1555  2.05
SSBOND   5 CYS A  325    CYS A  330                          1555   1555  2.04
SSBOND   6 CYS B    3    CYS B   11                          1555   1555  2.03
SSBOND   7 CYS B   36    CYS B   85                          1555   1555  2.03
SSBOND   8 CYS B  171    CYS B  184                          1555   1555  2.04
SSBOND   9 CYS B  246    CYS B  255                          1555   1555  2.05
SSBOND  10 CYS B  325    CYS B  330                          1555   1555  2.03
LINK        NA    NA A1343                 NZ  LYS A 122     1555   1555  2.73
LINK        NA    NA A1343                 O   HOH A2311     1555   1555  2.90
LINK        NA    NA A1343                 O   HOH A2574     1555   1555  2.54
LINK        NA    NA A1343                 O   HOH A2329     1555   1555  2.01
LINK        NA    NA A1343                 O   SER A 128     1555   1555  3.03
LINK        NA    NA B1344                 O   HOH B2416     1555   1555  2.95
LINK        NA    NA B1344                 O   HOH B2418     1555   1555  2.71
LINK        NA    NA B1344                 O   GLY B 167     1555   1555  3.09
SITE     1 AC1  3 PHE B 240  ASN B 273  HOH B2517
SITE     1 AC2  6 ARG B 166  GLY B 167  TYR B 169  ALA B 258
SITE     2 AC2  6 HOH B2416  HOH B2418
SITE     1 AC3  5 LYS A 122  SER A 128  HOH A2311  HOH A2329
SITE     2 AC3  5 HOH A2574
CRYST1   41.560  200.340   44.520  90.00 114.57  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.024062  0.000000  0.011001        0.00000
SCALE2      0.000000  0.004992  0.000000        0.00000
SCALE3      0.000000  0.000000  0.024698        0.00000
TER    2568      TYR A 342
TER    5164      TYR B 342
MASTER      398    0    3   33   22    0    5    6 6375    2   31   54
END