longtext: 4C4Z-pdb

content
HEADER    HYDROLASE                               09-SEP-13   4C4Z
TITLE     CRYSTAL STRUCTURE OF HUMAN BIFUNCTIONAL EPOXIDE HYDROXYLASE 2
TITLE    2 COMPLEXED WITH A8
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: BIFUNCTIONAL EPOXIDE HYDROLASE 2;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: EPOXIDE HYDROXYLASE DOMAIN RESIDUES 230-555;
COMPND   5 SYNONYM: BIFUNCTIONAL EPOXIDE HYDROXYLASE 2, CYTOSOLIC EPOXIDE
COMPND   6  HYDROLASE 2, CEH, EPOXIDE HYDRATASE, SOLUBLE EPOXIDE HYDROLASE,
COMPND   7  SEH, LIPID-PHOSPHATE PHOSPHATASE;
COMPND   8 EC: 3.3.2.10, 3.1.3.76;
COMPND   9 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR: PET16B
KEYWDS    HYDROLASE, DRUG DESIGN, MULTIPLE BINDING MODES
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.PILGER,A.MAZUR,P.MONECKE,H.SCHREUDER,B.ELSHORST,T.LANGER,
AUTHOR   2 A.SCHIFFER,I.KRIMM,M.WEGSTROTH,D.LEE,G.HESSLER,K.-U.WENDT,S.BECKER,
AUTHOR   3 C.GRIESINGER
REVDAT   1   01-OCT-14 4C4Z    0
JRNL        AUTH   J.PILGER,A.MAZUR,P.MONECKE,H.SCHREUDER,B.ELSHORST,T.LANGER,
JRNL        AUTH 2 A.SCHIFFER,I.KRIMM,M.WEGSTROTH,D.LEE,G.HESSLER,K.-U.WENDT,
JRNL        AUTH 3 S.BECKER,C.GRIESINGER
JRNL        TITL   A COMBINATION OF SPIN DIFFUSION METHODS FOR THE
JRNL        TITL 2 DETERMINATION OF PROTEIN-LIGAND COMPLEX STRUCTURAL
JRNL        TITL 3 ENSEMBLES
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.06 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : BUSTER 2.11.2
REMARK   3   AUTHORS     : BRICOGNE,BLANC,BRANDL,FLENSBURG,KELLER,
REMARK   3               : PACIOREK,ROVERSI,SHARFF,SMART,VONRHEIN,WOMACK;
REMARK   3               : MATTHEWS,TEN EYCK,TRONRUD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.06
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 28.67
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.0
REMARK   3   COMPLETENESS FOR RANGE        (%) : 89.94
REMARK   3   NUMBER OF REFLECTIONS             : 35547
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD           : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION   : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET)  : 0.2223
REMARK   3   R VALUE            (WORKING SET)  : 0.2200
REMARK   3   FREE R VALUE                      : 0.2530
REMARK   3   FREE R VALUE TEST SET SIZE   (%)  : 7.17
REMARK   3   FREE R VALUE TEST SET COUNT       : 2549
REMARK   3   ESTIMATED ERROR OF FREE R VALUE   : NULL
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED               : 18
REMARK   3   BIN RESOLUTION RANGE HIGH   (ANGSTROMS) : 2.06
REMARK   3   BIN RESOLUTION RANGE LOW    (ANGSTROMS) : 2.12
REMARK   3   BIN COMPLETENESS (WORKING+TEST)     (%) : 89.94
REMARK   3   REFLECTIONS IN BIN (WORKING + TEST SET) : 2638
REMARK   3   BIN R VALUE        (WORKING + TEST SET) : 0.2460
REMARK   3   REFLECTIONS IN BIN        (WORKING SET) : 2442
REMARK   3   BIN R VALUE               (WORKING SET) : 0.2422
REMARK   3   BIN FREE R VALUE                        : 0.2938
REMARK   3   BIN FREE R VALUE TEST SET SIZE      (%) : 7.43
REMARK   3   BIN FREE R VALUE TEST SET COUNT         : 196
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE     : NULL
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 5124
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 32
REMARK   3   SOLVENT ATOMS            : 602
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 25.10
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 27.24
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.9085
REMARK   3    B22 (A**2) : 7.8062
REMARK   3    B33 (A**2) : -8.7146
REMARK   3    B12 (A**2) : 0.0000
REMARK   3    B13 (A**2) : -2.9787
REMARK   3    B23 (A**2) : 0.0000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT                    (A) : 0.323
REMARK   3   DPI (BLOW EQ-10) BASED ON R VALUE        (A) : 0.349
REMARK   3   DPI (BLOW EQ-9) BASED ON FREE R VALUE    (A) : 0.221
REMARK   3   DPI (CRUICKSHANK) BASED ON R VALUE       (A) : 0.317
REMARK   3   DPI (CRUICKSHANK) BASED ON FREE R VALUE  (A) : 0.216
REMARK   3
REMARK   3   REFERENCES: BLOW, D. (2002) ACTA CRYST D58, 792-797
REMARK   3               CRUICKSHANK, D.W.J. (1999) ACTA CRYST D55, 583-601
REMARK   3
REMARK   3  CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.8927
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.8672
REMARK   3
REMARK   3   NUMBER OF GEOMETRIC FUNCTION TERMS DEFINED : 15
REMARK   3   TERM                          COUNT    WEIGHT   FUNCTION.
REMARK   3    BOND LENGTHS              :   5310 ;   2.00 ;  HARMONIC
REMARK   3    BOND ANGLES               :   7200 ;   2.00 ;  HARMONIC
REMARK   3    TORSION ANGLES            :   1808 ;   2.00 ;  SINUSOIDAL
REMARK   3    TRIGONAL CARBON PLANES    :    124 ;   2.00 ;  HARMONIC
REMARK   3    GENERAL PLANES            :    760 ;   5.00 ;  HARMONIC
REMARK   3    ISOTROPIC THERMAL FACTORS :   5310 ;  20.00 ;  HARMONIC
REMARK   3    BAD NON-BONDED CONTACTS   :   NULL ;   NULL ;  NULL
REMARK   3    IMPROPER TORSIONS         :   NULL ;   NULL ;  NULL
REMARK   3    PSEUDOROTATION ANGLES     :   NULL ;   NULL ;  NULL
REMARK   3    CHIRAL IMPROPER TORSION   :    640 ;   5.00 ;  SEMIHARMONIC
REMARK   3    SUM OF OCCUPANCIES        :   NULL ;   NULL ;  NULL
REMARK   3    UTILITY DISTANCES         :   NULL ;   NULL ;  NULL
REMARK   3    UTILITY ANGLES            :   NULL ;   NULL ;  NULL
REMARK   3    UTILITY TORSION           :   NULL ;   NULL ;  NULL
REMARK   3    IDEAL-DIST CONTACT TERM   :   6643 ;   4.00 ;  SEMIHARMONIC
REMARK   3
REMARK   3   RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3    BOND LENGTHS                       (A) : 0.007
REMARK   3    BOND ANGLES                  (DEGREES) : 0.92
REMARK   3    PEPTIDE OMEGA TORSION ANGLES (DEGREES) : 2.03
REMARK   3    OTHER TORSION ANGLES         (DEGREES) : 18.78
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: IDEAL-DIST CONTACT TERM CONTACT SETUP.
REMARK   3    ALL ATOMS HAVE CCP4 ATOM TYPE FROM LIBRARY.
REMARK   4
REMARK   4 4C4Z COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 09-SEP-13.
REMARK 100 THE PDBE ID CODE IS EBI-58283.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 21-DEC-12
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.3
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : ROTATING ANODE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU MICROMAX-007 HF
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : OSMIC MAXFLUX
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE (MAR345)
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 36430
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.05
REMARK 200  RESOLUTION RANGE LOW       (A) : 79.85
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 90.7
REMARK 200  DATA REDUNDANCY                : 2.7
REMARK 200  R MERGE                    (I) : 0.03
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 21.70
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.05
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.16
REMARK 200  COMPLETENESS FOR SHELL     (%) : 82.6
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.5
REMARK 200  R MERGE FOR SHELL          (I) : 0.16
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 8.30
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: OTHER
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NONE
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 44.6
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.22
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       39.92500
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ASN A   548
REMARK 465     PRO A   549
REMARK 465     PRO A   550
REMARK 465     VAL A   551
REMARK 465     VAL A   552
REMARK 465     SER A   553
REMARK 465     LYS A   554
REMARK 465     MET A   555
REMARK 465     ASN B   548
REMARK 465     PRO B   549
REMARK 465     PRO B   550
REMARK 465     VAL B   551
REMARK 465     VAL B   552
REMARK 465     SER B   553
REMARK 465     LYS B   554
REMARK 465     MET B   555
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A 247      -18.16    108.60
REMARK 500    PRO A 268       75.26   -101.77
REMARK 500    GLU A 269     -157.30    -98.26
REMARK 500    ASP A 335     -116.14     58.37
REMARK 500    ASN A 359      -40.92     72.11
REMARK 500    LEU A 480      107.68     16.49
REMARK 500    VAL A 498      -60.18    -97.20
REMARK 500    HIS A 513       48.89   -105.89
REMARK 500    PRO B 268       77.87   -100.72
REMARK 500    GLU B 269     -157.75   -100.99
REMARK 500    ASP B 335     -115.69     55.67
REMARK 500    ASN B 359      -44.30     68.19
REMARK 500    PHE B 381       39.51    -96.82
REMARK 500    PHE B 429       40.46    -90.71
REMARK 500    LEU B 480      107.90     -7.19
REMARK 500    VAL B 498      -60.14    -93.76
REMARK 500    HIS B 513       51.97    -93.53
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    GLU A 269        24.8      L          L   OUTSIDE RANGE
REMARK 500    VAL A 341        24.8      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE W9L A1548
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE W9L B1548
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4C4X   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF HUMAN BIFUNCTIONAL EPOXIDE
REMARK 900  HYDROXYLASE 2 COMPLEXED WITH C9
REMARK 900 RELATED ID: 4C4Y   RELATED DB: PDB
REMARK 900  CRYSTAL STRUCTURE OF HUMAN BIFUNCTIONAL EPOXIDE
REMARK 900  HYDROXYLASE 2 COMPLEXED WITH A4
DBREF  4C4Z A  230   555  UNP    P34913   HYES_HUMAN     230    555
DBREF  4C4Z B  230   555  UNP    P34913   HYES_HUMAN     230    555
SEQRES   1 A  326  THR SER CYS ASN PRO SER ASP MET SER HIS GLY TYR VAL
SEQRES   2 A  326  THR VAL LYS PRO ARG VAL ARG LEU HIS PHE VAL GLU LEU
SEQRES   3 A  326  GLY SER GLY PRO ALA VAL CYS LEU CYS HIS GLY PHE PRO
SEQRES   4 A  326  GLU SER TRP TYR SER TRP ARG TYR GLN ILE PRO ALA LEU
SEQRES   5 A  326  ALA GLN ALA GLY TYR ARG VAL LEU ALA MET ASP MET LYS
SEQRES   6 A  326  GLY TYR GLY GLU SER SER ALA PRO PRO GLU ILE GLU GLU
SEQRES   7 A  326  TYR CYS MET GLU VAL LEU CYS LYS GLU MET VAL THR PHE
SEQRES   8 A  326  LEU ASP LYS LEU GLY LEU SER GLN ALA VAL PHE ILE GLY
SEQRES   9 A  326  HIS ASP TRP GLY GLY MET LEU VAL TRP TYR MET ALA LEU
SEQRES  10 A  326  PHE TYR PRO GLU ARG VAL ARG ALA VAL ALA SER LEU ASN
SEQRES  11 A  326  THR PRO PHE ILE PRO ALA ASN PRO ASN MET SER PRO LEU
SEQRES  12 A  326  GLU SER ILE LYS ALA ASN PRO VAL PHE ASP TYR GLN LEU
SEQRES  13 A  326  TYR PHE GLN GLU PRO GLY VAL ALA GLU ALA GLU LEU GLU
SEQRES  14 A  326  GLN ASN LEU SER ARG THR PHE LYS SER LEU PHE ARG ALA
SEQRES  15 A  326  SER ASP GLU SER VAL LEU SER MET HIS LYS VAL CYS GLU
SEQRES  16 A  326  ALA GLY GLY LEU PHE VAL ASN SER PRO GLU GLU PRO SER
SEQRES  17 A  326  LEU SER ARG MET VAL THR GLU GLU GLU ILE GLN PHE TYR
SEQRES  18 A  326  VAL GLN GLN PHE LYS LYS SER GLY PHE ARG GLY PRO LEU
SEQRES  19 A  326  ASN TRP TYR ARG ASN MET GLU ARG ASN TRP LYS TRP ALA
SEQRES  20 A  326  CYS LYS SER LEU GLY ARG LYS ILE LEU ILE PRO ALA LEU
SEQRES  21 A  326  MET VAL THR ALA GLU LYS ASP PHE VAL LEU VAL PRO GLN
SEQRES  22 A  326  MET SER GLN HIS MET GLU ASP TRP ILE PRO HIS LEU LYS
SEQRES  23 A  326  ARG GLY HIS ILE GLU ASP CYS GLY HIS TRP THR GLN MET
SEQRES  24 A  326  ASP LYS PRO THR GLU VAL ASN GLN ILE LEU ILE LYS TRP
SEQRES  25 A  326  LEU ASP SER ASP ALA ARG ASN PRO PRO VAL VAL SER LYS
SEQRES  26 A  326  MET
SEQRES   1 B  326  THR SER CYS ASN PRO SER ASP MET SER HIS GLY TYR VAL
SEQRES   2 B  326  THR VAL LYS PRO ARG VAL ARG LEU HIS PHE VAL GLU LEU
SEQRES   3 B  326  GLY SER GLY PRO ALA VAL CYS LEU CYS HIS GLY PHE PRO
SEQRES   4 B  326  GLU SER TRP TYR SER TRP ARG TYR GLN ILE PRO ALA LEU
SEQRES   5 B  326  ALA GLN ALA GLY TYR ARG VAL LEU ALA MET ASP MET LYS
SEQRES   6 B  326  GLY TYR GLY GLU SER SER ALA PRO PRO GLU ILE GLU GLU
SEQRES   7 B  326  TYR CYS MET GLU VAL LEU CYS LYS GLU MET VAL THR PHE
SEQRES   8 B  326  LEU ASP LYS LEU GLY LEU SER GLN ALA VAL PHE ILE GLY
SEQRES   9 B  326  HIS ASP TRP GLY GLY MET LEU VAL TRP TYR MET ALA LEU
SEQRES  10 B  326  PHE TYR PRO GLU ARG VAL ARG ALA VAL ALA SER LEU ASN
SEQRES  11 B  326  THR PRO PHE ILE PRO ALA ASN PRO ASN MET SER PRO LEU
SEQRES  12 B  326  GLU SER ILE LYS ALA ASN PRO VAL PHE ASP TYR GLN LEU
SEQRES  13 B  326  TYR PHE GLN GLU PRO GLY VAL ALA GLU ALA GLU LEU GLU
SEQRES  14 B  326  GLN ASN LEU SER ARG THR PHE LYS SER LEU PHE ARG ALA
SEQRES  15 B  326  SER ASP GLU SER VAL LEU SER MET HIS LYS VAL CYS GLU
SEQRES  16 B  326  ALA GLY GLY LEU PHE VAL ASN SER PRO GLU GLU PRO SER
SEQRES  17 B  326  LEU SER ARG MET VAL THR GLU GLU GLU ILE GLN PHE TYR
SEQRES  18 B  326  VAL GLN GLN PHE LYS LYS SER GLY PHE ARG GLY PRO LEU
SEQRES  19 B  326  ASN TRP TYR ARG ASN MET GLU ARG ASN TRP LYS TRP ALA
SEQRES  20 B  326  CYS LYS SER LEU GLY ARG LYS ILE LEU ILE PRO ALA LEU
SEQRES  21 B  326  MET VAL THR ALA GLU LYS ASP PHE VAL LEU VAL PRO GLN
SEQRES  22 B  326  MET SER GLN HIS MET GLU ASP TRP ILE PRO HIS LEU LYS
SEQRES  23 B  326  ARG GLY HIS ILE GLU ASP CYS GLY HIS TRP THR GLN MET
SEQRES  24 B  326  ASP LYS PRO THR GLU VAL ASN GLN ILE LEU ILE LYS TRP
SEQRES  25 B  326  LEU ASP SER ASP ALA ARG ASN PRO PRO VAL VAL SER LYS
SEQRES  26 B  326  MET
HET    W9L  A1548      16
HET    W9L  B1548      16
HETNAM     W9L 1-ETHYL-3-NAPHTHALEN-1-YLUREA
FORMUL   3  W9L    2(C13 H14 N2 O)
FORMUL   4  HOH   *602(H2 O)
HELIX    1   1 ASN A  233  MET A  237  5                                   5
HELIX    2   2 SER A  270  ARG A  275  5                                   6
HELIX    3   3 TYR A  276  ALA A  284  1                                   9
HELIX    4   4 ILE A  305  TYR A  308  5                                   4
HELIX    5   5 CYS A  309  LEU A  324  1                                  16
HELIX    6   6 ASP A  335  TYR A  348  1                                  14
HELIX    7   7 SER A  370  ASN A  378  1                                   9
HELIX    8   8 PRO A  379  PHE A  381  5                                   3
HELIX    9   9 ASP A  382  PHE A  387  1                                   6
HELIX   10  10 GLY A  391  ASN A  400  1                                  10
HELIX   11  11 ASN A  400  PHE A  409  1                                  10
HELIX   12  12 LYS A  421  GLY A  426  1                                   6
HELIX   13  13 THR A  443  LYS A  455  1                                  13
HELIX   14  14 PHE A  459  TRP A  465  1                                   7
HELIX   15  15 ASN A  468  CYS A  477  1                                  10
HELIX   16  16 VAL A  500  GLN A  505  5                                   6
HELIX   17  17 HIS A  506  TRP A  510  5                                   5
HELIX   18  18 TRP A  525  LYS A  530  1                                   6
HELIX   19  19 LYS A  530  ASP A  545  1                                  16
HELIX   20  20 ASN B  233  MET B  237  5                                   5
HELIX   21  21 SER B  270  ARG B  275  5                                   6
HELIX   22  22 TYR B  276  GLY B  285  1                                  10
HELIX   23  23 ILE B  305  TYR B  308  5                                   4
HELIX   24  24 CYS B  309  LEU B  324  1                                  16
HELIX   25  25 ASP B  335  TYR B  348  1                                  14
HELIX   26  26 SER B  370  ASN B  378  1                                   9
HELIX   27  27 PHE B  381  PHE B  387  1                                   7
HELIX   28  28 GLY B  391  ASN B  400  1                                  10
HELIX   29  29 ASN B  400  PHE B  409  1                                  10
HELIX   30  30 THR B  443  PHE B  454  1                                  12
HELIX   31  31 PHE B  459  ASN B  464  1                                   6
HELIX   32  32 ASN B  468  CYS B  477  1                                  10
HELIX   33  33 VAL B  500  GLN B  505  5                                   6
HELIX   34  34 HIS B  506  TRP B  510  5                                   5
HELIX   35  35 TRP B  525  LYS B  530  1                                   6
HELIX   36  36 LYS B  530  ALA B  546  1                                  17
SHEET    1  AA16 LEU A 514  ILE A 519  0
SHEET    2  AA16 ALA A 488  ALA A 493  1  O  ALA A 488   N  LYS A 515
SHEET    3  AA16 VAL A 352  LEU A 358  1  O  VAL A 355   N  LEU A 489
SHEET    4  AA16 ALA A 329  HIS A 334  1  O  ALA A 329   N  ARG A 353
SHEET    5  AA16 ALA A 260  CYS A 264  1  O  ALA A 260   N  VAL A 330
SHEET    6  AA16 ARG A 287  ASP A 292  1  O  ARG A 287   N  VAL A 261
SHEET    7  AA16 VAL A 248  LEU A 255 -1  O  HIS A 251   N  ASP A 292
SHEET    8  AA16 SER A 238  LYS A 245 -1  O  SER A 238   N  GLU A 254
SHEET    9  AA16 SER B 238  LYS B 245 -1  O  HIS B 239   N  TYR A 241
SHEET   10  AA16 VAL B 248  LEU B 255 -1  O  VAL B 248   N  LYS B 245
SHEET   11  AA16 ARG B 287  MET B 291 -1  O  VAL B 288   N  LEU B 255
SHEET   12  AA16 ALA B 260  CYS B 264  1  O  VAL B 261   N  LEU B 289
SHEET   13  AA16 ALA B 329  HIS B 334  1  O  VAL B 330   N  CYS B 262
SHEET   14  AA16 VAL B 352  LEU B 358  1  N  ARG B 353   O  ALA B 329
SHEET   15  AA16 ALA B 488  ALA B 493  1  O  LEU B 489   N  SER B 357
SHEET   16  AA16 LEU B 514  ILE B 519  1  O  LYS B 515   N  MET B 490
CISPEP   1 PHE A  267    PRO A  268          0        -5.15
CISPEP   2 PHE B  267    PRO B  268          0        -5.35
SITE     1 AC1  8 PHE A 267  ASP A 335  TYR A 383  LEU A 408
SITE     2 AC1  8 MET A 419  TYR A 466  VAL A 498  HIS A 524
SITE     1 AC2  7 PHE B 267  ASP B 335  TYR B 383  MET B 419
SITE     2 AC2  7 TYR B 466  VAL B 498  HIS B 524
CRYST1   46.490   79.850   87.220  90.00  89.38  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.021510  0.000000 -0.000233        0.00000
SCALE2      0.000000  0.012523  0.000000        0.00000
SCALE3      0.000000  0.000000  0.011466        0.00000
TER    2563      ARG A 547
TER    5126      ARG B 547
MASTER      307    0    2   36   16    0    4    6 5758    2   32   52
END