longtext: 4G4J-pdb

content
HEADER    HYDROLASE                               16-JUL-12   4G4J
TITLE     CRYSTAL STRUCTURE OF GLUCURONOYL ESTERASE S213A MUTANT FROM
TITLE    2 SPOROTRICHUM THERMOPHILE IN COMPLEX WITH METHYL 4-O-METHYL-BETA-D-
TITLE    3 GLUCOPYRANURONATE DETERMINED AT 2.35 A RESOLUTION
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: 4-O-METHYL-GLUCURONOYL METHYLESTERASE;
COMPND   3 CHAIN: A;
COMPND   4 EC: 3.1.1.-;
COMPND   5 ENGINEERED: YES;
COMPND   6 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: MYCELIOPHTHORA THERMOPHILA;
SOURCE   3 ORGANISM_TAXID: 573729;
SOURCE   4 STRAIN: ATCC 42464;
SOURCE   5 GENE: MYCTH_55568;
SOURCE   6 EXPRESSION_SYSTEM: KOMAGATAELLA PASTORIS;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 4922;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: X-33;
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PPICZALPHAC
KEYWDS    ALPHA/BETA HYDROLASE, 3-LAYER ALPHA/BETA/ALPHA SANDWICH, ROSSMANN
KEYWDS   2 FOLD, CARBOHYDRATE BINDING, GLUCURONOYL ESTERASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.D.CHARAVGI,M.DIMAROGONA,E.TOPAKAS,P.CHRISTAKOPOULOS,E.D.CHRYSINA
REVDAT   1   02-JAN-13 4G4J    0
JRNL        AUTH   M.-D.CHARAVGI,M.DIMAROGONA,E.TOPAKAS,P.CHRISTAKOPOULOS,
JRNL        AUTH 2 E.D.CHRYSINA
JRNL        TITL   THE STRUCTURE OF A NOVEL GLUCURONOYL ESTERASE FROM
JRNL        TITL 2 MYCELIOPHTHORA THERMOPHILA GIVES NEW INSIGHTS INTO ITS ROLE
JRNL        TITL 3 AS A POTENTIAL BIOCATALYST.
JRNL        REF    ACTA CRYSTALLOGR.,SECT.D      V.  69    63 2013
JRNL        REFN                   ISSN 0907-4449
JRNL        DOI    10.1107/S0907444912042400
REMARK   2
REMARK   2 RESOLUTION.    2.35 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.6.0117
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.35
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 13.44
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 92.0
REMARK   3   NUMBER OF REFLECTIONS             : 14330
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.189
REMARK   3   R VALUE            (WORKING SET) : 0.186
REMARK   3   FREE R VALUE                     : 0.245
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 760
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.35
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.41
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 902
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 89.20
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2300
REMARK   3   BIN FREE R VALUE SET COUNT          : 48
REMARK   3   BIN FREE R VALUE                    : 0.2980
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2736
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 107
REMARK   3   SOLVENT ATOMS            : 212
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 14.80
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 9.22
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.01000
REMARK   3    B22 (A**2) : -0.00000
REMARK   3    B33 (A**2) : -0.01000
REMARK   3    B12 (A**2) : -0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.549
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.278
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.195
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 8.156
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.910
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.845
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  2889 ; 0.006 ; 0.020
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  3893 ; 1.006 ; 1.955
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   368 ; 5.089 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   121 ;30.720 ;24.628
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   410 ;12.863 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    14 ;14.599 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   419 ; 0.067 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  2196 ; 0.003 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN USED IF PRESENT IN
REMARK   3  THE INPUT
REMARK   4
REMARK   4 4G4J COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 18-JUL-12.
REMARK 100 THE RCSB ID CODE IS RCSB073728.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 16-NOV-11
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 4.6
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : SEALED TUBE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : OTHER
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5419
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : MUTLTI-LAYER XRAY OPTIC
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : AGILENT ATLAS CCD
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : CRYSALISPRO
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 15099
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.350
REMARK 200  RESOLUTION RANGE LOW       (A) : 13.480
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 92.8
REMARK 200  DATA REDUNDANCY                : 3.400
REMARK 200  R MERGE                    (I) : 0.11000
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 7.7000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.35
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.48
REMARK 200  COMPLETENESS FOR SHELL     (%) : 90.4
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.15000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 5.300
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 4G4I
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 45.60
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.26
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 25% PEG 550 MME, 0.1M SODIUM ACETATE,
REMARK 280  PH 4.6, VAPOR DIFFUSION, SITTING DROP, TEMPERATURE 289K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 2 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y,-Z
REMARK 290       3555   X+1/2,-Y,-Z+1/2
REMARK 290       4555   -X+1/2,-Y,Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       26.04800
REMARK 290   SMTRY2   3  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       51.97500
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       26.04800
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000       51.97500
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     SER A   -13
REMARK 465     MET A   -12
REMARK 465     ALA A   -11
REMARK 465     PRO A   -10
REMARK 465     MET A    -9
REMARK 465     ASN A    -8
REMARK 465     HIS A    -7
REMARK 465     ILE A    -6
REMARK 465     PHE A    -5
REMARK 465     GLU A    -4
REMARK 465     ARG A    -3
REMARK 465     GLN A    -2
REMARK 465     ASP A    -1
REMARK 465     THR A     0
REMARK 465     MET A     1
REMARK 465     VAL A     2
REMARK 465     HIS A     3
REMARK 465     LEU A     4
REMARK 465     THR A     5
REMARK 465     SER A     6
REMARK 465     ALA A     7
REMARK 465     LEU A     8
REMARK 465     LEU A     9
REMARK 465     VAL A    10
REMARK 465     ALA A    11
REMARK 465     GLY A    12
REMARK 465     ALA A    13
REMARK 465     ALA A    14
REMARK 465     PHE A    15
REMARK 465     ALA A    16
REMARK 465     ALA A    17
REMARK 465     ALA A    18
REMARK 465     ALA A    19
REMARK 465     PRO A    20
REMARK 465     MET A    21
REMARK 465     ASN A    22
REMARK 465     HIS A    23
REMARK 465     ILE A    24
REMARK 465     PHE A    25
REMARK 465     GLU A    26
REMARK 465     ARG A    27
REMARK 465     GLN A    28
REMARK 465     LEU A   398
REMARK 465     GLU A   399
REMARK 465     GLN A   400
REMARK 465     LYS A   401
REMARK 465     LEU A   402
REMARK 465     ILE A   403
REMARK 465     SER A   404
REMARK 465     GLU A   405
REMARK 465     GLU A   406
REMARK 465     ASP A   407
REMARK 465     LEU A   408
REMARK 465     ASN A   409
REMARK 465     SER A   410
REMARK 465     ALA A   411
REMARK 465     VAL A   412
REMARK 465     ASP A   413
REMARK 465     HIS A   414
REMARK 465     HIS A   415
REMARK 465     HIS A   416
REMARK 465     HIS A   417
REMARK 465     HIS A   418
REMARK 465     HIS A   419
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    GLU A  78      -54.22   -148.52
REMARK 500    ALA A 213     -114.67     57.36
REMARK 500    PRO A 269       42.19    -81.58
REMARK 500    SER A 281       63.76   -154.01
REMARK 500    PRO A 286       39.77    -76.27
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MCU A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 502
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 503
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 504
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 505
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 506
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 507
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 508
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 509
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 510
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 511
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 512
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 513
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 514
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 515
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 516
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 517
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 518
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 519
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 520
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 521
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 522
REMARK 800
REMARK 800 SITE_IDENTIFIER: CC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE GOL A 523
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4G4G   RELATED DB: PDB
REMARK 900 CRYSTAL STRUCTURE OF GLUCURONOYL ESTERASE FROM SPOROTRICHUM
REMARK 900 THERMOPHILE DETERMINED AT 1.55 A RESOLUTION
REMARK 900 RELATED ID: 4G4I   RELATED DB: PDB
REMARK 900 CRYSTAL STRUCTURE OF GLUCURONOYL ESTERASE S213A MUTANT FROM
REMARK 900 SPOROTRICHUM THERMOPHILE DETERMINED AT 1.9 A RESOLUTION
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 THIS SEGMENT CORRESPONDS TO A C-MYC EPITOPE
DBREF  4G4J A    1   397  UNP    G2QJR6   G2QJR6_THIHA     1    397
SEQADV 4G4J SER A  -13  UNP  G2QJR6              INSERTION
SEQADV 4G4J MET A  -12  UNP  G2QJR6              INSERTION
SEQADV 4G4J ALA A  -11  UNP  G2QJR6              INSERTION
SEQADV 4G4J PRO A  -10  UNP  G2QJR6              INSERTION
SEQADV 4G4J MET A   -9  UNP  G2QJR6              INSERTION
SEQADV 4G4J ASN A   -8  UNP  G2QJR6              INSERTION
SEQADV 4G4J HIS A   -7  UNP  G2QJR6              INSERTION
SEQADV 4G4J ILE A   -6  UNP  G2QJR6              INSERTION
SEQADV 4G4J PHE A   -5  UNP  G2QJR6              INSERTION
SEQADV 4G4J GLU A   -4  UNP  G2QJR6              INSERTION
SEQADV 4G4J ARG A   -3  UNP  G2QJR6              INSERTION
SEQADV 4G4J GLN A   -2  UNP  G2QJR6              INSERTION
SEQADV 4G4J ASP A   -1  UNP  G2QJR6              INSERTION
SEQADV 4G4J THR A    0  UNP  G2QJR6              INSERTION
SEQADV 4G4J ALA A  213  UNP  G2QJR6    SER   213 ENGINEERED MUTATION
SEQADV 4G4J LEU A  398  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J GLU A  399  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J GLN A  400  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J LYS A  401  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J LEU A  402  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J ILE A  403  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J SER A  404  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J GLU A  405  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J GLU A  406  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J ASP A  407  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J LEU A  408  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J ASN A  409  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J SER A  410  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J ALA A  411  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J VAL A  412  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J ASP A  413  UNP  G2QJR6              SEE REMARK 999
SEQADV 4G4J HIS A  414  UNP  G2QJR6              EXPRESSION TAG
SEQADV 4G4J HIS A  415  UNP  G2QJR6              EXPRESSION TAG
SEQADV 4G4J HIS A  416  UNP  G2QJR6              EXPRESSION TAG
SEQADV 4G4J HIS A  417  UNP  G2QJR6              EXPRESSION TAG
SEQADV 4G4J HIS A  418  UNP  G2QJR6              EXPRESSION TAG
SEQADV 4G4J HIS A  419  UNP  G2QJR6              EXPRESSION TAG
SEQRES   1 A  433  SER MET ALA PRO MET ASN HIS ILE PHE GLU ARG GLN ASP
SEQRES   2 A  433  THR MET VAL HIS LEU THR SER ALA LEU LEU VAL ALA GLY
SEQRES   3 A  433  ALA ALA PHE ALA ALA ALA ALA PRO MET ASN HIS ILE PHE
SEQRES   4 A  433  GLU ARG GLN ASP THR CYS SER VAL SER ASP ASN TYR PRO
SEQRES   5 A  433  THR VAL ASN SER ALA LYS LEU PRO ASP PRO PHE THR THR
SEQRES   6 A  433  ALA SER GLY GLU LYS VAL THR THR LYS ASP GLN PHE GLU
SEQRES   7 A  433  CYS ARG ARG ALA GLU ILE ASN LYS ILE LEU GLN GLN TYR
SEQRES   8 A  433  GLU LEU GLY GLU TYR PRO GLY PRO PRO ASP SER VAL GLU
SEQRES   9 A  433  ALA SER LEU SER GLY ASN SER ILE THR VAL ARG VAL THR
SEQRES  10 A  433  VAL GLY SER LYS SER ILE SER PHE SER ALA SER ILE ARG
SEQRES  11 A  433  LYS PRO SER GLY ALA GLY PRO PHE PRO ALA ILE ILE GLY
SEQRES  12 A  433  ILE GLY GLY ALA SER ILE PRO ILE PRO SER ASN VAL ALA
SEQRES  13 A  433  THR ILE THR PHE ASN ASN ASP GLU PHE GLY ALA GLN MET
SEQRES  14 A  433  GLY SER GLY SER ARG GLY GLN GLY LYS PHE TYR ASP LEU
SEQRES  15 A  433  PHE GLY ARG ASP HIS SER ALA GLY SER LEU THR ALA TRP
SEQRES  16 A  433  ALA TRP GLY VAL ASP ARG LEU ILE ASP GLY LEU GLU GLN
SEQRES  17 A  433  VAL GLY ALA GLN ALA SER GLY ILE ASP THR LYS ARG LEU
SEQRES  18 A  433  GLY VAL THR GLY CYS ALA ARG ASN GLY LYS GLY ALA PHE
SEQRES  19 A  433  ILE THR GLY ALA LEU VAL ASP ARG ILE ALA LEU THR ILE
SEQRES  20 A  433  PRO GLN GLU SER GLY ALA GLY GLY ALA ALA CYS TRP ARG
SEQRES  21 A  433  ILE SER ASP GLN GLN LYS ALA ALA GLY ALA ASN ILE GLN
SEQRES  22 A  433  THR ALA ALA GLN ILE ILE THR GLU ASN PRO TRP PHE SER
SEQRES  23 A  433  ARG ASN PHE ASP PRO HIS VAL ASN SER ILE THR SER VAL
SEQRES  24 A  433  PRO GLN ASP HIS HIS LEU LEU ALA ALA LEU ILE VAL PRO
SEQRES  25 A  433  ARG GLY LEU ALA VAL PHE GLU ASN ASN ILE ASP TRP LEU
SEQRES  26 A  433  GLY PRO VAL SER THR THR GLY CYS MET ALA ALA GLY ARG
SEQRES  27 A  433  LEU ILE TYR LYS ALA TYR GLY VAL PRO ASN ASN MET GLY
SEQRES  28 A  433  PHE SER LEU VAL GLY GLY HIS ASN HIS CYS GLN PHE PRO
SEQRES  29 A  433  SER SER GLN ASN GLN ASP LEU ASN SER TYR ILE ASN TYR
SEQRES  30 A  433  PHE LEU LEU GLY GLN GLY SER PRO SER GLY VAL GLU HIS
SEQRES  31 A  433  SER ASP VAL ASN VAL ASN VAL ALA GLU TRP ALA PRO TRP
SEQRES  32 A  433  GLY ALA GLY ALA PRO THR LEU ALA LEU GLU GLN LYS LEU
SEQRES  33 A  433  ILE SER GLU GLU ASP LEU ASN SER ALA VAL ASP HIS HIS
SEQRES  34 A  433  HIS HIS HIS HIS
HET    MCU  A 501      15
HET    EDO  A 502       4
HET    EDO  A 503       4
HET    EDO  A 504       4
HET    EDO  A 505       4
HET    EDO  A 506       4
HET    EDO  A 507       4
HET    EDO  A 508       4
HET    EDO  A 509       4
HET    EDO  A 510       4
HET    EDO  A 511       4
HET    EDO  A 512       4
HET    EDO  A 513       4
HET    EDO  A 514       4
HET    EDO  A 515       4
HET    EDO  A 516       4
HET    EDO  A 517       4
HET    EDO  A 518       4
HET    EDO  A 519       4
HET    EDO  A 520       4
HET    EDO  A 521       4
HET    GOL  A 522       6
HET    GOL  A 523       6
HETNAM     MCU METHYL 4-O-METHYL-BETA-D-GLUCOPYRANURONATE
HETNAM     EDO 1,2-ETHANEDIOL
HETNAM     GOL GLYCEROL
HETSYN     EDO ETHYLENE GLYCOL
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
FORMUL   2  MCU    C8 H14 O7
FORMUL   3  EDO    20(C2 H6 O2)
FORMUL  23  GOL    2(C3 H8 O3)
FORMUL  25  HOH   *212(H2 O)
HELIX    1   1 THR A   59  GLU A   78  1                                  20
HELIX    2   2 ASN A  147  GLY A  152  1                                   6
HELIX    3   3 GLY A  156  ARG A  160  5                                   5
HELIX    4   4 GLY A  163  GLY A  170  1                                   8
HELIX    5   5 GLY A  176  GLY A  196  1                                  21
HELIX    6   6 GLY A  196  GLY A  201  1                                   6
HELIX    7   7 ALA A  213  VAL A  226  1                                  14
HELIX    8   8 CYS A  244  ALA A  254  1                                  11
HELIX    9   9 THR A  260  ILE A  265  1                                   6
HELIX   10  10 PHE A  275  VAL A  279  5                                   5
HELIX   11  11 SER A  281  VAL A  285  5                                   5
HELIX   12  12 ASP A  288  HIS A  290  5                                   3
HELIX   13  13 LEU A  291  VAL A  297  1                                   7
HELIX   14  14 ILE A  308  LEU A  311  5                                   4
HELIX   15  15 GLY A  312  GLY A  331  1                                  20
HELIX   16  16 VAL A  332  ASN A  334  5                                   3
HELIX   17  17 PRO A  350  SER A  352  5                                   3
HELIX   18  18 GLN A  353  LEU A  365  1                                  13
HELIX   19  19 ASN A  382  ALA A  387  1                                   6
SHEET    1   A10 SER A  88  SER A  94  0
SHEET    2   A10 SER A  97  VAL A 104 -1  O  ARG A 101   N  GLU A  90
SHEET    3   A10 LYS A 107  ARG A 116 -1  O  PHE A 111   N  VAL A 100
SHEET    4   A10 ALA A 142  PHE A 146 -1  O  THR A 143   N  ARG A 116
SHEET    5   A10 PHE A 124  ILE A 130  1  N  GLY A 129   O  ILE A 144
SHEET    6   A10 ILE A 202  CYS A 212  1  O  GLY A 208   N  ILE A 128
SHEET    7   A10 LEU A 231  GLN A 235  1  O  ILE A 233   N  VAL A 209
SHEET    8   A10 GLY A 300  GLU A 305  1  O  ALA A 302   N  THR A 232
SHEET    9   A10 MET A 336  LEU A 340  1  O  SER A 339   N  GLU A 305
SHEET   10   A10 GLU A 375  HIS A 376  1  O  HIS A 376   N  LEU A 340
SSBOND   1 CYS A   65    CYS A   31                          1555   1555  2.03
SSBOND   2 CYS A  319    CYS A  244                          1555   1555  2.04
SSBOND   3 CYS A  347    CYS A  212                          1555   1555  2.03
CISPEP   1 GLY A  122    PRO A  123          0         3.02
CISPEP   2 VAL A  297    PRO A  298          0        -0.68
SITE     1 AC1  9 ALA A 213  ARG A 214  LYS A 217  GLN A 259
SITE     2 AC1  9 GLU A 267  TRP A 310  LEU A 311  HIS A 346
SITE     3 AC1  9 HOH A 793
SITE     1 AC2  6 VAL A 332  PRO A 333  ASN A 334  ASN A 335
SITE     2 AC2  6 HOH A 735  HOH A 744
SITE     1 AC3  6 ASP A  47  LYS A  56  TRP A 389  GLY A 392
SITE     2 AC3  6 ALA A 393  EDO A 513
SITE     1 AC4  8 ARG A  67  ALA A 224  LEU A 225  ASP A 227
SITE     2 AC4  8 LEU A 295  ARG A 299  HOH A 699  HOH A 798
SITE     1 AC5  8 ARG A 246  ASP A 249  GLN A 250  ILE A 282
SITE     2 AC5  8 THR A 283  HOH A 620  HOH A 649  HOH A 749
SITE     1 AC6  4 GLN A 250  VAL A 381  ASN A 382  TRP A 386
SITE     1 AC7  6 PHE A 111  SER A 112  ALA A 113  PHE A 146
SITE     2 AC7  6 ASN A 147  GLU A 150
SITE     1 AC8  8 PRO A 138  ASN A 140  ASN A 307  GLY A 342
SITE     2 AC8  8 GLY A 343  ASN A 362  LEU A 366  ASP A 378
SITE     1 AC9  3 PRO A 138  VAL A 141  THR A 143
SITE     1 BC1  3 ASN A  41  ALA A  43  HOH A 757
SITE     1 BC2  3 ASN A 140  GOL A 523  HOH A 723
SITE     1 BC3  5 GLU A  90  SER A 108  ILE A 109  SER A 110
SITE     2 BC3  5 HOH A 779
SITE     1 BC4  5 ASP A  47  THR A  50  LYS A  56  ASP A  87
SITE     2 BC4  5 EDO A 503
SITE     1 BC5  5 VAL A  40  ASN A  41  SER A  42  PRO A  85
SITE     2 BC5  5 HOH A 804
SITE     1 BC6  6 ASN A 307  ASN A 358  VAL A 379  EDO A 517
SITE     2 BC6  6 HOH A 611  HOH A 619
SITE     1 BC7  8 ARG A 160  ILE A 265  ASN A 268  PRO A 269
SITE     2 BC7  8 TRP A 270  PHE A 271  SER A 272  ASP A 276
SITE     1 BC8  7 ASN A 307  ASP A 309  PRO A 313  ASN A 354
SITE     2 BC8  7 GLN A 355  ASN A 358  EDO A 515
SITE     1 BC9  8 ALA A  52  SER A  53  GLY A  54  PRO A  85
SITE     2 BC9  8 PRO A  86  ARG A 187  ASP A 190  EDO A 519
SITE     1 CC1  6 PRO A  38  VAL A  40  ALA A  52  PRO A  85
SITE     2 CC1  6 PRO A 286  EDO A 518
SITE     1 CC2  1 ASP A 203
SITE     1 CC3  4 LYS A  44  GLU A 385  TRP A 386  HOH A 763
SITE     1 CC4  3 ASP A 249  LYS A 252  ALA A 253
SITE     1 CC5  6 ASN A 140  ALA A 253  GLY A 255  LEU A 366
SITE     2 CC5  6 EDO A 511  HOH A 712
CRYST1   52.096   69.766  103.950  90.00  90.00  90.00 P 21 2 21     4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.019195  0.000000  0.000000        0.00000
SCALE2      0.000000  0.014334  0.000000        0.00000
SCALE3      0.000000  0.000000  0.009620        0.00000
TER    2737      ALA A 397
MASTER      427    0   23   19   10    0   40    6 3055    1  113   34
END