longtext: 4GBG-pdb

content
HEADER    HYDROLASE                               27-JUL-12   4GBG
TITLE     CRYSTAL STRUCTURE OF ETHYL ACETOACETATE TREATED LIPASE FROM
TITLE    2 THERMOMYCES LANUGINOSA AT 2.9 A RESOLUTION
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: A;
COMPND   5 SYNONYM: TRIACYLGLYCEROL LIPASE;
COMPND   6 EC: 3.1.1.3
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: THERMOMYCES LANUGINOSA;
SOURCE   3 ORGANISM_TAXID: 5541
KEYWDS    HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.YAMINI,J.MUKHERJEE,M.N.GUPTA,M.SINHA,P.KAUR,S.SHARMA,T.P.SINGH
REVDAT   1   22-AUG-12 4GBG    0
JRNL        AUTH   S.YAMINI,J.MUKHERJEE,M.N.GUPTA,M.SINHA,P.KAUR,S.SHARMA,
JRNL        AUTH 2 T.P.SINGH
JRNL        TITL   CRYSTAL STRUCTURE OF ETHYL ACETOACETATE TREATED LIPASE FROM
JRNL        TITL 2 THERMOMYCES LANUGINOSA AT 2.9 A RESOLUTION
JRNL        REF    TO BE PUBLISHED
JRNL        REFN
REMARK   2
REMARK   2 RESOLUTION.    2.90 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : CNS 1.1
REMARK   3   AUTHORS     : BRUNGER,ADAMS,CLORE,DELANO,GROS,GROSSE-
REMARK   3               : KUNSTLEVE,JIANG,KUSZEWSKI,NILGES,PANNU,
REMARK   3               : READ,RICE,SIMONSON,WARREN
REMARK   3
REMARK   3  REFINEMENT TARGET : NULL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.90
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 40.56
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   DATA CUTOFF HIGH         (ABS(F)) : 2537552.830
REMARK   3   DATA CUTOFF LOW          (ABS(F)) : 0.0000
REMARK   3   COMPLETENESS (WORKING+TEST)   (%) : 99.9
REMARK   3   NUMBER OF REFLECTIONS             : 20259
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE            (WORKING SET) : 0.226
REMARK   3   FREE R VALUE                     : 0.247
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.900
REMARK   3   FREE R VALUE TEST SET COUNT      : 985
REMARK   3   ESTIMATED ERROR OF FREE R VALUE  : 0.008
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 6
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.90
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 3.08
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.90
REMARK   3   REFLECTIONS IN BIN    (WORKING SET) : 3161
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2800
REMARK   3   BIN FREE R VALUE                    : 0.2940
REMARK   3   BIN FREE R VALUE TEST SET SIZE  (%) : 5.20
REMARK   3   BIN FREE R VALUE TEST SET COUNT     : 175
REMARK   3   ESTIMATED ERROR OF BIN FREE R VALUE : 0.022
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4140
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 37
REMARK   3   SOLVENT ATOMS            : 350
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 55.30
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 50.60
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 2.00000
REMARK   3    B22 (A**2) : 2.00000
REMARK   3    B33 (A**2) : -4.00000
REMARK   3    B12 (A**2) : 3.55000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM LUZZATI PLOT        (A) : 0.36
REMARK   3   ESD FROM SIGMAA              (A) : 0.36
REMARK   3   LOW RESOLUTION CUTOFF        (A) : 5.00
REMARK   3
REMARK   3  CROSS-VALIDATED ESTIMATED COORDINATE ERROR.
REMARK   3   ESD FROM C-V LUZZATI PLOT    (A) : 0.42
REMARK   3   ESD FROM C-V SIGMAA          (A) : 0.44
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES.
REMARK   3   BOND LENGTHS                 (A) : 0.010
REMARK   3   BOND ANGLES            (DEGREES) : 1.70
REMARK   3   DIHEDRAL ANGLES        (DEGREES) : 25.70
REMARK   3   IMPROPER ANGLES        (DEGREES) : 1.13
REMARK   3
REMARK   3  ISOTROPIC THERMAL MODEL : RESTRAINED
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.    RMS    SIGMA
REMARK   3   MAIN-CHAIN BOND              (A**2) : 1.680 ; 1.500
REMARK   3   MAIN-CHAIN ANGLE             (A**2) : 2.800 ; 2.000
REMARK   3   SIDE-CHAIN BOND              (A**2) : 2.170 ; 2.000
REMARK   3   SIDE-CHAIN ANGLE             (A**2) : 3.350 ; 2.500
REMARK   3
REMARK   3  BULK SOLVENT MODELING.
REMARK   3   METHOD USED : FLAT MODEL
REMARK   3   KSOL        : 0.33
REMARK   3   BSOL        : 50.03
REMARK   3
REMARK   3  NCS MODEL : NULL
REMARK   3
REMARK   3  NCS RESTRAINTS.                         RMS   SIGMA/WEIGHT
REMARK   3   GROUP  1  POSITIONAL            (A) : NULL  ; NULL
REMARK   3   GROUP  1  B-FACTOR           (A**2) : NULL  ; NULL
REMARK   3
REMARK   3  PARAMETER FILE  1  : PROTEIN_REP.PARAM
REMARK   3  PARAMETER FILE  2  : ION.PARAM
REMARK   3  PARAMETER FILE  3  : WATER_REP.PARAM
REMARK   3  PARAMETER FILE  4  : CARBOHYDRATE.PARAM
REMARK   3  PARAMETER FILE  5  : XXX.PARAM
REMARK   3  PARAMETER FILE  6  : NULL
REMARK   3  TOPOLOGY FILE  1   : PROTEIN.TOP
REMARK   3  TOPOLOGY FILE  2   : ION.TOP
REMARK   3  TOPOLOGY FILE  3   : WATER.TOP
REMARK   3  TOPOLOGY FILE  4   : CARBOHYDRATE.TOP
REMARK   3  TOPOLOGY FILE  5   : XXX.TOP
REMARK   3  TOPOLOGY FILE  6   : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4GBG COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 10-AUG-12.
REMARK 100 THE RCSB ID CODE IS RCSB073975.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 05-MAY-12
REMARK 200  TEMPERATURE           (KELVIN) : 77
REMARK 200  PH                             : 7.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : BM14
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97
REMARK 200  MONOCHROMATOR                  : GRAPHITE
REMARK 200  OPTICS                         : MIRROR
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MARRESEARCH
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : DENZO
REMARK 200  DATA SCALING SOFTWARE          : SCALEPACK
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 20267
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.900
REMARK 200  RESOLUTION RANGE LOW       (A) : 40.560
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : NULL
REMARK 200  R MERGE                    (I) : 0.10300
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 11.0000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.90
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 3.06
REMARK 200  COMPLETENESS FOR SHELL     (%) : 100.0
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : 0.50000
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 3.100
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: AMORE
REMARK 200 STARTING MODEL: 4FLF
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 68.60
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.92
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M HEPES, 0.1M NACL, 1.6M AMMONIUM
REMARK 280  SULPHATE, PH 7.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 298K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 61
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+1/3
REMARK 290       3555   -X+Y,-X,Z+2/3
REMARK 290       4555   -X,-Y,Z+1/2
REMARK 290       5555   Y,-X+Y,Z+5/6
REMARK 290       6555   X-Y,X,Z+1/6
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       26.89667
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       53.79333
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000       40.34500
REMARK 290   SMTRY1   5  0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   5 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000  1.000000       67.24167
REMARK 290   SMTRY1   6  0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000  1.000000       13.44833
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1520 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 21170 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -9.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OD1  ASN A   101     O    CYS A   104              1.80
REMARK 500   OD1  ASN A   101     N    GLY A   106              2.09
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ILE A  76   N   -  CA  -  C   ANGL. DEV. = -16.2 DEGREES
REMARK 500    CYS A 104   CA  -  CB  -  SG  ANGL. DEV. =   9.2 DEGREES
REMARK 500    PHE A 262   CB  -  CA  -  C   ANGL. DEV. = -13.7 DEGREES
REMARK 500    VAL B 187   CB  -  CA  -  C   ANGL. DEV. = -21.3 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP A  62       76.67     46.47
REMARK 500    VAL A 120      -27.67   -144.95
REMARK 500    SER A 146     -118.02     52.66
REMARK 500    THR A 199     -125.95     52.82
REMARK 500    ASP A 201      142.15    -36.22
REMARK 500    PRO A 204        4.93    -57.72
REMARK 500    LEU A 227       -0.46     79.34
REMARK 500    PRO A 253     -169.15    -71.96
REMARK 500    PHE A 262       -6.75     63.90
REMARK 500    TYR B  21      -64.96   -101.35
REMARK 500    LYS B  24       62.67   -106.30
REMARK 500    ASN B  25      -70.94    -85.91
REMARK 500    ASN B  26     -126.22     53.03
REMARK 500    CYS B  41       56.76   -144.08
REMARK 500    ILE B 100       62.25   -155.97
REMARK 500    SER B 146     -133.80     59.86
REMARK 500    ASN B 162       26.65   -148.47
REMARK 500    VAL B 187       29.22   -149.77
REMARK 500    THR B 199     -106.90     17.24
REMARK 500    LEU B 227       13.56     82.32
REMARK 500    PHE B 262      -57.92     72.35
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: PLANAR GROUPS
REMARK 500
REMARK 500 PLANAR GROUPS IN THE FOLLOWING RESIDUES HAVE A TOTAL
REMARK 500 RMS DISTANCE OF ALL ATOMS FROM THE BEST-FIT PLANE
REMARK 500 BY MORE THAN AN EXPECTED VALUE OF 6*RMSD, WITH AN
REMARK 500 RMSD 0.02 ANGSTROMS, OR AT LEAST ONE ATOM HAS
REMARK 500 AN RMSD GREATER THAN THIS VALUE
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        RMS     TYPE
REMARK 500    TYR B 171         0.07    SIDE CHAIN
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EAC A 301
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG A 302
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NAG B 301
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4FLF   RELATED DB: PDB
DBREF  4GBG A    1   269  UNP    O59952   LIP_THELA       23    291
DBREF  4GBG B    1   269  UNP    O59952   LIP_THELA       23    291
SEQRES   1 A  269  GLU VAL SER GLN ASP LEU PHE ASN GLN PHE ASN LEU PHE
SEQRES   2 A  269  ALA GLN TYR SER ALA ALA ALA TYR CYS GLY LYS ASN ASN
SEQRES   3 A  269  ASP ALA PRO ALA GLY THR ASN ILE THR CYS THR GLY ASN
SEQRES   4 A  269  ALA CYS PRO GLU VAL GLU LYS ALA ASP ALA THR PHE LEU
SEQRES   5 A  269  TYR SER PHE GLU ASP SER GLY VAL GLY ASP VAL THR GLY
SEQRES   6 A  269  PHE LEU ALA LEU ASP ASN THR ASN LYS LEU ILE VAL LEU
SEQRES   7 A  269  SER PHE ARG GLY SER ARG SER ILE GLU ASN TRP ILE GLY
SEQRES   8 A  269  ASN LEU ASN PHE ASP LEU LYS GLU ILE ASN ASP ILE CYS
SEQRES   9 A  269  SER GLY CYS ARG GLY HIS ASP GLY PHE THR SER SER TRP
SEQRES  10 A  269  ARG SER VAL ALA ASP THR LEU ARG GLN LYS VAL GLU ASP
SEQRES  11 A  269  ALA VAL ARG GLU HIS PRO ASP TYR ARG VAL VAL PHE THR
SEQRES  12 A  269  GLY HIS SER LEU GLY GLY ALA LEU ALA THR VAL ALA GLY
SEQRES  13 A  269  ALA ASP LEU ARG GLY ASN GLY TYR ASP ILE ASP VAL PHE
SEQRES  14 A  269  SER TYR GLY ALA PRO ARG VAL GLY ASN ARG ALA PHE ALA
SEQRES  15 A  269  GLU PHE LEU THR VAL GLN THR GLY GLY THR LEU TYR ARG
SEQRES  16 A  269  ILE THR HIS THR ASN ASP ILE VAL PRO ARG LEU PRO PRO
SEQRES  17 A  269  ARG GLU PHE GLY TYR SER HIS SER SER PRO GLU TYR TRP
SEQRES  18 A  269  ILE LYS SER GLY THR LEU VAL PRO VAL THR ARG ASN ASP
SEQRES  19 A  269  ILE VAL LYS ILE GLU GLY ILE ASP ALA THR GLY GLY ASN
SEQRES  20 A  269  ASN GLN PRO ASN ILE PRO ASP ILE PRO ALA HIS LEU TRP
SEQRES  21 A  269  TYR PHE GLY LEU ILE GLY THR CYS LEU
SEQRES   1 B  269  GLU VAL SER GLN ASP LEU PHE ASN GLN PHE ASN LEU PHE
SEQRES   2 B  269  ALA GLN TYR SER ALA ALA ALA TYR CYS GLY LYS ASN ASN
SEQRES   3 B  269  ASP ALA PRO ALA GLY THR ASN ILE THR CYS THR GLY ASN
SEQRES   4 B  269  ALA CYS PRO GLU VAL GLU LYS ALA ASP ALA THR PHE LEU
SEQRES   5 B  269  TYR SER PHE GLU ASP SER GLY VAL GLY ASP VAL THR GLY
SEQRES   6 B  269  PHE LEU ALA LEU ASP ASN THR ASN LYS LEU ILE VAL LEU
SEQRES   7 B  269  SER PHE ARG GLY SER ARG SER ILE GLU ASN TRP ILE GLY
SEQRES   8 B  269  ASN LEU ASN PHE ASP LEU LYS GLU ILE ASN ASP ILE CYS
SEQRES   9 B  269  SER GLY CYS ARG GLY HIS ASP GLY PHE THR SER SER TRP
SEQRES  10 B  269  ARG SER VAL ALA ASP THR LEU ARG GLN LYS VAL GLU ASP
SEQRES  11 B  269  ALA VAL ARG GLU HIS PRO ASP TYR ARG VAL VAL PHE THR
SEQRES  12 B  269  GLY HIS SER LEU GLY GLY ALA LEU ALA THR VAL ALA GLY
SEQRES  13 B  269  ALA ASP LEU ARG GLY ASN GLY TYR ASP ILE ASP VAL PHE
SEQRES  14 B  269  SER TYR GLY ALA PRO ARG VAL GLY ASN ARG ALA PHE ALA
SEQRES  15 B  269  GLU PHE LEU THR VAL GLN THR GLY GLY THR LEU TYR ARG
SEQRES  16 B  269  ILE THR HIS THR ASN ASP ILE VAL PRO ARG LEU PRO PRO
SEQRES  17 B  269  ARG GLU PHE GLY TYR SER HIS SER SER PRO GLU TYR TRP
SEQRES  18 B  269  ILE LYS SER GLY THR LEU VAL PRO VAL THR ARG ASN ASP
SEQRES  19 B  269  ILE VAL LYS ILE GLU GLY ILE ASP ALA THR GLY GLY ASN
SEQRES  20 B  269  ASN GLN PRO ASN ILE PRO ASP ILE PRO ALA HIS LEU TRP
SEQRES  21 B  269  TYR PHE GLY LEU ILE GLY THR CYS LEU
MODRES 4GBG ASN B   33  ASN  GLYCOSYLATION SITE
MODRES 4GBG ASN A   33  ASN  GLYCOSYLATION SITE
HET    EAC  A 301       9
HET    NAG  A 302      14
HET    NAG  B 301      14
HETNAM     EAC ETHYL 3-OXOBUTANOATE
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   3  EAC    C6 H10 O3
FORMUL   4  NAG    2(C8 H15 N O6)
FORMUL   6  HOH   *350(H2 O)
HELIX    1   1 SER A    3  GLY A   23  1                                  21
HELIX    2   2 CYS A   41  ALA A   47  1                                   7
HELIX    3   3 SER A   85  ASN A   92  1                                   8
HELIX    4   4 ASP A  111  SER A  119  1                                   9
HELIX    5   5 VAL A  120  HIS A  135  1                                  16
HELIX    6   6 SER A  146  ARG A  160  1                                  15
HELIX    7   7 ARG A  179  GLN A  188  1                                  10
HELIX    8   8 PRO A  208  PHE A  211  5                                   4
HELIX    9   9 THR A  231  ASN A  233  5                                   3
HELIX   10  10 PRO A  256  TRP A  260  5                                   5
HELIX   11  11 SER B    3  GLY B   23  1                                  21
HELIX   12  12 CYS B   41  ALA B   47  1                                   7
HELIX   13  13 SER B   85  ASN B   92  1                                   8
HELIX   14  14 HIS B  110  HIS B  135  1                                  26
HELIX   15  15 SER B  146  LEU B  159  1                                  14
HELIX   16  16 ASN B  178  GLN B  188  1                                  11
HELIX   17  17 ILE B  202  LEU B  206  5                                   5
HELIX   18  18 PRO B  208  GLY B  212  5                                   5
HELIX   19  19 THR B  231  ASN B  233  5                                   3
HELIX   20  20 ILE B  255  TRP B  260  5                                   6
SHEET    1   A 8 ALA A  49  SER A  58  0
SHEET    2   A 8 VAL A  63  ASP A  70 -1  O  VAL A  63   N  SER A  58
SHEET    3   A 8 LEU A  75  PHE A  80 -1  O  VAL A  77   N  ALA A  68
SHEET    4   A 8 ARG A 139  HIS A 145  1  O  VAL A 141   N  ILE A  76
SHEET    5   A 8 ILE A 166  TYR A 171  1  O  TYR A 171   N  GLY A 144
SHEET    6   A 8 LEU A 193  HIS A 198  1  O  TYR A 194   N  VAL A 168
SHEET    7   A 8 GLU A 219  ILE A 222  1  O  ILE A 222   N  THR A 197
SHEET    8   A 8 ILE A 235  ILE A 238 -1  O  ILE A 238   N  GLU A 219
SHEET    1   B 2 LEU A  97  GLU A  99  0
SHEET    2   B 2 ARG A 108  HIS A 110 -1  O  GLY A 109   N  LYS A  98
SHEET    1   C 2 GLY A 177  ASN A 178  0
SHEET    2   C 2 TYR A 213  SER A 214 -1  O  SER A 214   N  GLY A 177
SHEET    1   D 8 ALA B  49  SER B  58  0
SHEET    2   D 8 VAL B  63  ASP B  70 -1  O  LEU B  67   N  LEU B  52
SHEET    3   D 8 LEU B  75  PHE B  80 -1  O  VAL B  77   N  ALA B  68
SHEET    4   D 8 ARG B 139  HIS B 145  1  O  VAL B 141   N  ILE B  76
SHEET    5   D 8 ILE B 166  TYR B 171  1  O  ASP B 167   N  PHE B 142
SHEET    6   D 8 LEU B 193  HIS B 198  1  O  TYR B 194   N  VAL B 168
SHEET    7   D 8 GLU B 219  ILE B 222  1  O  ILE B 222   N  THR B 197
SHEET    8   D 8 ILE B 235  ILE B 238 -1  O  VAL B 236   N  TRP B 221
SSBOND   1 CYS A   22    CYS A  268                          1555   1555  2.03
SSBOND   2 CYS A   36    CYS A   41                          1555   1555  2.03
SSBOND   3 CYS A  104    CYS A  107                          1555   1555  2.04
SSBOND   4 CYS B   22    CYS B  268                          1555   1555  2.02
SSBOND   5 CYS B   36    CYS B   41                          1555   1555  2.03
SSBOND   6 CYS B  104    CYS B  107                          1555   1555  2.04
LINK         ND2 ASN B  33                 C1  NAG B 301     1555   1555  1.44
LINK         ND2 ASN A  33                 C1  NAG A 302     1555   1555  1.45
CISPEP   1 LEU A  206    PRO A  207          0         0.08
CISPEP   2 SER A  217    PRO A  218          0         0.14
CISPEP   3 LEU B  206    PRO B  207          0         0.66
CISPEP   4 SER B  217    PRO B  218          0         0.88
SITE     1 AC1  3 ILE A  86  GLU A  87  HOH A 453
SITE     1 AC2  1 ASN A  33
SITE     1 AC3  2 ASN B  33  HOH B 557
CRYST1  140.489  140.489   80.690  90.00  90.00 120.00 P 61         12
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.007118  0.004110  0.000000        0.00000
SCALE2      0.000000  0.008219  0.000000        0.00000
SCALE3      0.000000  0.000000  0.012393        0.00000
TER    2071      LEU A 269
TER    4142      LEU B 269
MASTER      334    0    3   20   20    0    3    6 4527    2   51   42
END