longtext: 4HXF-pdb

content
HEADER    HYDROLASE/HYDROLASE INHIBITOR           09-NOV-12   4HXF
TITLE     ACYLAMINOACYL PEPTIDASE IN COMPLEX WITH Z-GLY-GLY-PHE-CHLOROMETHYL
TITLE    2 KETONE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PUTATIVE UNCHARACTERIZED PROTEIN PH0594;
COMPND   3 CHAIN: B;
COMPND   4 EC: 3.4.19.1;
COMPND   5 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PYROCOCCUS HORIKOSHII;
SOURCE   3 ORGANISM_TAXID: 70601;
SOURCE   4 STRAIN: ATCC 700860 / DSM 12428 / JCM 9974 / NBRC 100139 / OT-3;
SOURCE   5 GENE: PH0594;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   8 EXPRESSION_SYSTEM_STRAIN: ROSETTA(DE3);
SOURCE   9 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PET15B
KEYWDS    SELF-COMPARTMENTALIZATION, BETA-PROPELLER, ALPHA/BETA HYRDOLASE FOLD,
KEYWDS   2 HYDROLASE, HYDROLASE-HYDROLASE INHIBITOR COMPLEX
EXPDTA    X-RAY DIFFRACTION
AUTHOR    A.KISS-SZEMAN,D.K.MENYHARD,E.TICHY-RACS,B.HORNUNG,K.RADI,Z.SZELTNER,
AUTHOR   2 K.DOMOKOS,I.SZAMOSI,G.NARAY-SZABO,L.POLGAR,V.HARMAT
REVDAT   2   15-MAY-13 4HXF    1       JRNL
REVDAT   1   08-MAY-13 4HXF    0
JRNL        AUTH   D.K.MENYHARD,A.KISS-SZEMAN,E.TICHY-RACS,B.HORNUNG,K.RADI,
JRNL        AUTH 2 Z.SZELTNER,K.DOMOKOS,I.SZAMOSI,G.NARAY-SZABO,L.POLGAR,
JRNL        AUTH 3 V.HARMAT
JRNL        TITL   A SELF-COMPARTMENTALIZING HEXAMER SERINE PROTEASE FROM
JRNL        TITL 2 PYROCOCCUS HORIKOSHII - SUBSTRATE SELECTION ACHIEVED THROUGH
JRNL        TITL 3 MULTIMERIZATION.
JRNL        REF    J.BIOL.CHEM.                               2013
JRNL        REFN                   ESSN 1083-351X
JRNL        PMID   23632025
JRNL        DOI    10.1074/JBC.M113.451534
REMARK   1
REMARK   1 REFERENCE 1
REMARK   1  AUTH   Z.SZELTNER,A.L.KISS,K.DOMOKOS,V.HARMAT,G.NARAY-SZABO,
REMARK   1  AUTH 2 L.POLGAR
REMARK   1  TITL   CHARACTERIZATION OF A NOVEL ACYLAMINOACYL PEPTIDASE WITH
REMARK   1  TITL 2 HEXAMERIC STRUCTURE AND ENDOPEPTIDASE ACTIVITY.
REMARK   1  REF    BIOCHIM.BIOPHYS.ACTA          V.1794  1204 2009
REMARK   1  REFN                   ISSN 0006-3002
REMARK   1  PMID   19303951
REMARK   1  DOI    10.1016/J.BBAPAP.2009.03.004
REMARK   2
REMARK   2 RESOLUTION.    1.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.6.0117
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.60
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.74
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.2
REMARK   3   NUMBER OF REFLECTIONS             : 116219
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.156
REMARK   3   R VALUE            (WORKING SET) : 0.155
REMARK   3   FREE R VALUE                     : 0.186
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.100
REMARK   3   FREE R VALUE TEST SET COUNT      : 6218
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.60
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.64
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 7838
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 94.19
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2900
REMARK   3   BIN FREE R VALUE SET COUNT          : 447
REMARK   3   BIN FREE R VALUE                    : 0.2930
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 5019
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 102
REMARK   3   SOLVENT ATOMS            : 655
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 19.81
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.05000
REMARK   3    B22 (A**2) : 0.05000
REMARK   3    B33 (A**2) : -0.08000
REMARK   3    B12 (A**2) : 0.03000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.064
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.068
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.048
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 2.824
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.973
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.961
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  5327 ; 0.025 ; 0.020
REMARK   3   BOND LENGTHS OTHERS               (A):  3806 ; 0.001 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  7178 ; 2.248 ; 1.963
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  9201 ; 1.108 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   641 ; 7.019 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   263 ;34.562 ;23.650
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   919 ;13.597 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    33 ;18.017 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   736 ; 0.144 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  5867 ; 0.013 ; 0.020
REMARK   3   GENERAL PLANES OTHERS             (A):  1195 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 2
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B     4        B    12
REMARK   3    RESIDUE RANGE :   B   345        B   618
REMARK   3    ORIGIN FOR THE GROUP (A):  -7.8090 -44.7260 -14.5920
REMARK   3    T TENSOR
REMARK   3      T11:   0.0479 T22:   0.0364
REMARK   3      T33:   0.0249 T12:   0.0024
REMARK   3      T13:   0.0009 T23:  -0.0018
REMARK   3    L TENSOR
REMARK   3      L11:   0.5309 L22:   0.3278
REMARK   3      L33:   0.5254 L12:   0.0677
REMARK   3      L13:  -0.0801 L23:   0.0216
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0232 S12:   0.0281 S13:  -0.0906
REMARK   3      S21:  -0.0194 S22:   0.0063 S23:  -0.0621
REMARK   3      S31:   0.0702 S32:   0.0575 S33:   0.0169
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B    13        B   344
REMARK   3    ORIGIN FOR THE GROUP (A): -18.5000 -20.8950 -29.5520
REMARK   3    T TENSOR
REMARK   3      T11:   0.0851 T22:   0.0866
REMARK   3      T33:   0.0101 T12:  -0.0122
REMARK   3      T13:  -0.0187 T23:   0.0073
REMARK   3    L TENSOR
REMARK   3      L11:   0.2141 L22:   0.2842
REMARK   3      L33:   0.1288 L12:   0.0621
REMARK   3      L13:  -0.0337 L23:  -0.0214
REMARK   3    S TENSOR
REMARK   3      S11:   0.0030 S12:   0.0427 S13:   0.0218
REMARK   3      S21:  -0.0229 S22:   0.0165 S23:   0.0104
REMARK   3      S31:   0.0010 S32:  -0.0249 S33:  -0.0195
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : BABINET MODEL WITH MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS:
REMARK   3  HYDROGENS HAVE BEEN ADDED IN THE RIDING POSITIONS.
REMARK   3  HEZ B711 AND WATER B1336 LIE ON A TWOFOLD AXIS.
REMARK   4
REMARK   4 4HXF COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 14-NOV-12.
REMARK 100 THE RCSB ID CODE IS RCSB076040.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 15-NOV-10
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : ROTATING ANODE
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : RIGAKU RU200
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.5418
REMARK 200  MONOCHROMATOR                  : CONFOCAL MIRRORS
REMARK 200  OPTICS                         : CONFOCAL MIRRORS
REMARK 200
REMARK 200  DETECTOR TYPE                  : IMAGE PLATE
REMARK 200  DETECTOR MANUFACTURER          : RIGAKU RAXIS IV++
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 122438
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.600
REMARK 200  RESOLUTION RANGE LOW       (A) : 20.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.2
REMARK 200  DATA REDUNDANCY                : 7.030
REMARK 200  R MERGE                    (I) : 0.06600
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 22.2500
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.64
REMARK 200  COMPLETENESS FOR SHELL     (%) : 94.4
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.24
REMARK 200  R MERGE FOR SHELL          (I) : 0.73800
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.570
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: PDB ENTRY 4HXE
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 61.73
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.21
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 2.5 M 1,6-HEXANEDIOL, 0.20 M MGCL2,
REMARK 280  0.1 M TRIS/HCL , PH 8.5, VAPOR DIFFUSION, HANGING DROP,
REMARK 280  TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: H 3 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z
REMARK 290       3555   -X+Y,-X,Z
REMARK 290       4555   Y,X,-Z
REMARK 290       5555   X-Y,-Y,-Z
REMARK 290       6555   -X,-X+Y,-Z
REMARK 290       7555   X+2/3,Y+1/3,Z+1/3
REMARK 290       8555   -Y+2/3,X-Y+1/3,Z+1/3
REMARK 290       9555   -X+Y+2/3,-X+1/3,Z+1/3
REMARK 290      10555   Y+2/3,X+1/3,-Z+1/3
REMARK 290      11555   X-Y+2/3,-Y+1/3,-Z+1/3
REMARK 290      12555   -X+2/3,-X+Y+1/3,-Z+1/3
REMARK 290      13555   X+1/3,Y+2/3,Z+2/3
REMARK 290      14555   -Y+1/3,X-Y+2/3,Z+2/3
REMARK 290      15555   -X+Y+1/3,-X+2/3,Z+2/3
REMARK 290      16555   Y+1/3,X+2/3,-Z+2/3
REMARK 290      17555   X-Y+1/3,-Y+2/3,-Z+2/3
REMARK 290      18555   -X+1/3,-X+Y+2/3,-Z+2/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   4  0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   7  1.000000  0.000000  0.000000       92.02850
REMARK 290   SMTRY2   7  0.000000  1.000000  0.000000       53.13268
REMARK 290   SMTRY3   7  0.000000  0.000000  1.000000       48.50967
REMARK 290   SMTRY1   8 -0.500000 -0.866025  0.000000       92.02850
REMARK 290   SMTRY2   8  0.866025 -0.500000  0.000000       53.13268
REMARK 290   SMTRY3   8  0.000000  0.000000  1.000000       48.50967
REMARK 290   SMTRY1   9 -0.500000  0.866025  0.000000       92.02850
REMARK 290   SMTRY2   9 -0.866025 -0.500000  0.000000       53.13268
REMARK 290   SMTRY3   9  0.000000  0.000000  1.000000       48.50967
REMARK 290   SMTRY1  10 -0.500000  0.866025  0.000000       92.02850
REMARK 290   SMTRY2  10  0.866025  0.500000  0.000000       53.13268
REMARK 290   SMTRY3  10  0.000000  0.000000 -1.000000       48.50967
REMARK 290   SMTRY1  11  1.000000  0.000000  0.000000       92.02850
REMARK 290   SMTRY2  11  0.000000 -1.000000  0.000000       53.13268
REMARK 290   SMTRY3  11  0.000000  0.000000 -1.000000       48.50967
REMARK 290   SMTRY1  12 -0.500000 -0.866025  0.000000       92.02850
REMARK 290   SMTRY2  12 -0.866025  0.500000  0.000000       53.13268
REMARK 290   SMTRY3  12  0.000000  0.000000 -1.000000       48.50967
REMARK 290   SMTRY1  13  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  13  0.000000  1.000000  0.000000      106.26536
REMARK 290   SMTRY3  13  0.000000  0.000000  1.000000       97.01933
REMARK 290   SMTRY1  14 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  14  0.866025 -0.500000  0.000000      106.26536
REMARK 290   SMTRY3  14  0.000000  0.000000  1.000000       97.01933
REMARK 290   SMTRY1  15 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  15 -0.866025 -0.500000  0.000000      106.26536
REMARK 290   SMTRY3  15  0.000000  0.000000  1.000000       97.01933
REMARK 290   SMTRY1  16 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2  16  0.866025  0.500000  0.000000      106.26536
REMARK 290   SMTRY3  16  0.000000  0.000000 -1.000000       97.01933
REMARK 290   SMTRY1  17  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2  17  0.000000 -1.000000  0.000000      106.26536
REMARK 290   SMTRY3  17  0.000000  0.000000 -1.000000       97.01933
REMARK 290   SMTRY1  18 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2  18 -0.866025  0.500000  0.000000      106.26536
REMARK 290   SMTRY3  18  0.000000  0.000000 -1.000000       97.01933
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: HEXAMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: HEXAMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 48050 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 120550 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -145.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 350   BIOMT2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 350   BIOMT3   2  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 350   BIOMT2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 350   BIOMT3   3  0.000000  0.000000  1.000000        0.00000
REMARK 350   BIOMT1   4 -0.500000  0.866025  0.000000        0.00000
REMARK 350   BIOMT2   4  0.866025  0.500000  0.000000        0.00000
REMARK 350   BIOMT3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 350   BIOMT1   5  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   5  0.000000 -1.000000  0.000000        0.00000
REMARK 350   BIOMT3   5  0.000000  0.000000 -1.000000        0.00000
REMARK 350   BIOMT1   6 -0.500000 -0.866025  0.000000        0.00000
REMARK 350   BIOMT2   6 -0.866025  0.500000  0.000000        0.00000
REMARK 350   BIOMT3   6  0.000000  0.000000 -1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH B1429  LIES ON A SPECIAL POSITION.
REMARK 375      HOH B1120  LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET B     1
REMARK 465     THR B     2
REMARK 465     SER B     3
REMARK 465     ASN B    79
REMARK 465     GLU B    80
REMARK 465     GLU B   619
REMARK 465     GLU B   620
REMARK 465     LYS B   621
REMARK 465     LYS B   622
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     ILE B   4    CD1
REMARK 470     GLU B   5    CG   CD   OE1  OE2
REMARK 470     LYS B  13    CE   NZ
REMARK 470     ARG B  23    NE   CZ   NH1  NH2
REMARK 470     LYS B  40    CE   NZ
REMARK 470     GLU B  51    CB   CG   CD   OE1  OE2
REMARK 470     ASP B  52    CG   OD1  OD2
REMARK 470     GLU B  81    CG   CD   OE1  OE2
REMARK 470     LYS B  82    CG   CD   CE   NZ
REMARK 470     LYS B  83    CG   CD   CE   NZ
REMARK 470     THR B  84    OG1  CG2
REMARK 470     LYS B  93    CD   CE   NZ
REMARK 470     LYS B  99    CD   CE   NZ
REMARK 470     LYS B 104    CE   NZ
REMARK 470     LYS B 151    CE   NZ
REMARK 470     ASP B 259    CG   OD1  OD2
REMARK 470     ASN B 271    CG   OD1  ND2
REMARK 470     GLU B 292    CD   OE1  OE2
REMARK 470     ARG B 333    CZ   NH1  NH2
REMARK 470     LYS B 344    CD   CE   NZ
REMARK 470     GLU B 371    CG   CD   OE1  OE2
REMARK 470     LYS B 373    CG   CD   CE   NZ
REMARK 470     GLU B 374    CG   CD   OE1  OE2
REMARK 470     GLU B 520    CD   OE1  OE2
REMARK 470     LYS B 532    NZ
REMARK 470     LYS B 575    CE   NZ
REMARK 470     LYS B 606    CG   CD   CE   NZ
REMARK 470     GLU B 614    CG   CD   OE1  OE2
REMARK 470     LYS B 615    CD   CE   NZ
REMARK 470     LYS B 618    CG   CD   CE   NZ
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O6   HEZ B   714     O    HOH B  1433              2.06
REMARK 500   O    HOH B  1276     O    HOH B  1441              2.07
REMARK 500   NZ   LYS B   124     OE2  GLU B   169              2.09
REMARK 500   OG   SER B   466     O4   Y3A B   701              2.17
REMARK 500   O    HOH B  1086     O    HOH B  1423              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    GLU B 110   CD    GLU B 110   OE1    -0.084
REMARK 500    GLU B 163   CD    GLU B 163   OE2    -0.082
REMARK 500    HIS B 179   CG    HIS B 179   CD2     0.074
REMARK 500    HIS B 189   CG    HIS B 189   CD2     0.056
REMARK 500    GLU B 242   CD    GLU B 242   OE2    -0.085
REMARK 500    TYR B 412   CE1   TYR B 412   CZ     -0.079
REMARK 500    GLU B 458   CD    GLU B 458   OE1     0.067
REMARK 500    TRP B 506   NE1   TRP B 506   CE2    -0.082
REMARK 500    HIS B 542   CG    HIS B 542   CD2     0.059
REMARK 500    HIS B 559   CG    HIS B 559   CD2     0.081
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ASP B  19   CB  -  CG  -  OD2 ANGL. DEV. =  -6.0 DEGREES
REMARK 500    GLU B 102   O   -  C   -  N   ANGL. DEV. =  11.5 DEGREES
REMARK 500    ALA B 103   C   -  N   -  CA  ANGL. DEV. =  15.4 DEGREES
REMARK 500    ASP B 114   CB  -  CG  -  OD1 ANGL. DEV. =   6.2 DEGREES
REMARK 500    ASP B 136   CB  -  CG  -  OD2 ANGL. DEV. =  -7.0 DEGREES
REMARK 500    ASP B 148   CB  -  CG  -  OD2 ANGL. DEV. =  -8.9 DEGREES
REMARK 500    LYS B 198   CD  -  CE  -  NZ  ANGL. DEV. =  16.2 DEGREES
REMARK 500    ARG B 428   NE  -  CZ  -  NH1 ANGL. DEV. =   3.1 DEGREES
REMARK 500    PHE B 557   CB  -  CG  -  CD1 ANGL. DEV. =   4.3 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASP B   7     -164.77   -165.26
REMARK 500    LYS B  24     -126.98     55.05
REMARK 500    LYS B  83       43.53     38.94
REMARK 500    ASP B 136     -164.95   -122.32
REMARK 500    GLU B 162       51.25     70.38
REMARK 500    ASP B 191     -116.38   -109.06
REMARK 500    ARG B 195       59.96    -90.13
REMARK 500    LYS B 198      -54.86   -125.96
REMARK 500    ASP B 309     -106.86   -153.36
REMARK 500    ASP B 337       49.84   -146.37
REMARK 500    ILE B 359     -112.45     65.81
REMARK 500    LYS B 389       46.67    -85.65
REMARK 500    LEU B 435     -103.16    -99.65
REMARK 500    SER B 466     -119.55     65.91
REMARK 500    PHE B 611       86.30   -155.69
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: NON-CIS, NON-TRANS
REMARK 500
REMARK 500 THE FOLLOWING PEPTIDE BONDS DEVIATE SIGNIFICANTLY FROM BOTH
REMARK 500 CIS AND TRANS CONFORMATION.  CIS BONDS, IF ANY, ARE LISTED
REMARK 500 ON CISPEP RECORDS.  TRANS IS DEFINED AS 180 +/- 30 AND
REMARK 500 CIS IS DEFINED AS 0 +/- 30 DEGREES.
REMARK 500                                 MODEL     OMEGA
REMARK 500 GLU B  102     ALA B  103                 -146.00
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: MAIN CHAIN PLANARITY
REMARK 500
REMARK 500 THE FOLLOWING RESIDUES HAVE A PSEUDO PLANARITY
REMARK 500 TORSION ANGLE, C(I) - CA(I) - N(I+1) - O(I), GREATER
REMARK 500 10.0 DEGREES. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 500 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 500 I=INSERTION CODE).
REMARK 500
REMARK 500  M RES CSSEQI        ANGLE
REMARK 500    GLU B 371         11.34
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CHIRAL CENTERS
REMARK 500
REMARK 500 UNEXPECTED CONFIGURATION OF THE FOLLOWING CHIRAL
REMARK 500 CENTER(S) USING IMPROPER CA--C--CB--N CHIRALITY
REMARK 500 FOR AMINO ACIDS AND C1'--O4'--N1(N9)--C2' FOR
REMARK 500 NUCLEIC ACIDS OR EQUIVALENT ANGLE
REMARK 500 M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,6X,F5.1,6X,A1,10X,A1,3X,A16)
REMARK 500
REMARK 500  M RES CSSEQI    IMPROPER   EXPECTED   FOUND DETAILS
REMARK 500    LYS B 104        23.3      L          L   OUTSIDE RANGE
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH B1262        DISTANCE =  5.60 ANGSTROMS
REMARK 525    HOH B1276        DISTANCE =  6.12 ANGSTROMS
REMARK 525    HOH B1278        DISTANCE =  5.10 ANGSTROMS
REMARK 610
REMARK 610 MISSING HETEROATOM
REMARK 610 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 610 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 610 I=INSERTION CODE):
REMARK 610   M RES C SSEQI
REMARK 610     Y3A B  701
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG B 706  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH B 926   O
REMARK 620 2 HOH B1106   O    86.8
REMARK 620 3 GLU B 127   OE1  94.4  92.2
REMARK 620 4 HOH B1135   O   100.3 172.8  86.7
REMARK 620 5 HOH B1349   O    99.0  91.7 166.2  87.8
REMARK 620 6 HOH B1272   O   169.4  83.3  82.2  89.5  85.1
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG B 705  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 GLU B 162   OE1
REMARK 620 2 HOH B 876   O    88.8
REMARK 620 3 LYS B  99   O    87.3  86.6
REMARK 620 N                    1     2
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG B 703  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH B 975   O
REMARK 620 2 HOH B 978   O    90.7
REMARK 620 3 ASP B 244   OD2  92.2 176.7
REMARK 620 4 GLU B 263   OE2  89.5  89.1  92.3
REMARK 620 5 HOH B 865   O    86.4  91.5  87.4 175.8
REMARK 620 6 HOH B 862   O   179.0  88.9  88.1  91.5  92.7
REMARK 620 N                    1     2     3     4     5
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG B 704  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH B 956   O
REMARK 620 2 HOH B1229   O    86.8
REMARK 620 3 HOH B 915   O    88.9 173.0
REMARK 620 4 HOH B 901   O    93.9  92.5  93.3
REMARK 620 N                    1     2     3
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              MG B 702  MG
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 HOH B 980   O
REMARK 620 2 HOH B 891   O    92.0
REMARK 620 3 ASP B 424   OD1  95.1  87.0
REMARK 620 N                    1     2
REMARK 630
REMARK 630 MOLECULE TYPE: PEPTIDE-LIKE ENZYME INHIBITOR
REMARK 630 MOLECULE NAME: (PHENYLMETHYL) N-[2-OXIDANYLIDENE-2-[[2-
REMARK 630 OXIDANYLIDENE-2-[[(2S,3S)-3-OXIDANYL-1-PHENYL-BUTAN-2-YL]AMINO]
REMARK 630 ETHYL]AMINO]ETHYL]CARBAMATE
REMARK 630 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 630  SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 630
REMARK 630   M RES C SSSEQI
REMARK 630     Y3A B   701
REMARK 630 SOURCE: NULL
REMARK 630 TAXONOMY: NULL
REMARK 630 SUBCOMP:    PHQ GLY GLY HPH 0QE
REMARK 630 DETAILS: THE CHLORORMETHYL KETONE INHIBITOR LINKED TO THE PROTEIN
REMARK 630 THROUGH TWO COVALENT BONDS WITH THE ACTIVE SITE SERINE AND
REMARK 630 HISTIDINE
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE Y3A B 701
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MG B 702
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MG B 703
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MG B 704
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MG B 705
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MG B 706
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 707
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 708
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 709
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 710
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 711
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 712
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 713
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 714
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 715
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE HEZ B 716
REMARK 800
REMARK 800 SITE_IDENTIFIER: BC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CL B 717
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4HXE   RELATED DB: PDB
REMARK 900 RELATED ID: 4HXG   RELATED DB: PDB
DBREF  4HXF B    1   622  UNP    O58323   O58323_PYRHO     1    622
SEQRES   1 B  622  MET THR SER ILE GLU TRP ASP GLU LYS THR PHE THR LYS
SEQRES   2 B  622  PHE ALA TYR LEU SER ASP PRO ARG THR ARG LYS ASN LEU
SEQRES   3 B  622  VAL ALA TYR VAL LEU THR LYS ALA ASN LEU GLU SER ASN
SEQRES   4 B  622  LYS TYR GLU ASN THR ILE VAL ILE GLU ASN LEU GLU ASP
SEQRES   5 B  622  GLY SER ARG LYS PHE ILE GLU ASP ALA SER MET PRO ARG
SEQRES   6 B  622  ILE SER PRO ASP GLY LYS LYS ILE ALA PHE MET ARG PHE
SEQRES   7 B  622  ASN GLU GLU LYS LYS THR ALA GLN ILE TRP VAL ALA ASP
SEQRES   8 B  622  LEU LYS THR LEU SER ALA LYS LYS VAL LEU GLU ALA LYS
SEQRES   9 B  622  ASN ILE ARG SER ILE GLU TRP ASN GLN ASP SER ARG ARG
SEQRES  10 B  622  LEU LEU ALA VAL GLY PHE LYS ARG ARG GLU ASP GLU ASP
SEQRES  11 B  622  PHE ILE PHE GLU ASP ASP VAL PRO ALA TRP PHE ASP ASN
SEQRES  12 B  622  MET GLY PHE PHE ASP GLY GLU LYS THR THR PHE TRP VAL
SEQRES  13 B  622  ILE ASP THR GLU GLY GLU GLU VAL ILE GLU GLN PHE GLU
SEQRES  14 B  622  LYS PRO ARG PHE SER SER GLY ILE TRP HIS GLY ASP SER
SEQRES  15 B  622  ILE VAL VAL SER VAL PRO HIS ARG ASP VAL ILE PRO ARG
SEQRES  16 B  622  TYR PHE LYS TYR TRP ASP ILE TYR LEU TRP LYS ASP GLY
SEQRES  17 B  622  GLU GLU GLU LYS LEU PHE GLU LYS VAL SER PHE TYR ALA
SEQRES  18 B  622  ILE ASP SER ASP GLY GLU ARG ILE LEU LEU TYR GLY LYS
SEQRES  19 B  622  PRO GLU LYS LYS TYR VAL SER GLU HIS ASP LYS ILE TYR
SEQRES  20 B  622  ILE TYR ASP GLY GLU VAL LYS GLY ILE LEU ASP ASP ILE
SEQRES  21 B  622  ASP ARG GLU VAL ALA GLN ALA LYS ILE ARG ASN GLY LYS
SEQRES  22 B  622  VAL TYR PHE THR LEU PHE GLU GLU GLY SER VAL ASN LEU
SEQRES  23 B  622  TYR LEU TRP ASP GLY GLU VAL ARG GLU ILE ALA LYS GLY
SEQRES  24 B  622  LYS HIS TRP ILE MET GLY PHE ASP ALA ASP GLU ARG LEU
SEQRES  25 B  622  ILE TYR LEU LYS GLU THR ALA THR ARG PRO ALA GLU LEU
SEQRES  26 B  622  TYR LEU TRP ASP GLY GLU GLU ARG GLN LEU THR ASP TYR
SEQRES  27 B  622  ASN GLY LEU ILE PHE LYS LYS LEU LYS THR PHE GLU PRO
SEQRES  28 B  622  ARG HIS PHE ARG PHE LYS SER ILE ASP LEU GLU LEU ASP
SEQRES  29 B  622  GLY TRP TYR ILE LYS PRO GLU ILE LYS GLU GLY GLU LYS
SEQRES  30 B  622  ALA PRO VAL ILE VAL PHE VAL HIS GLY GLY PRO LYS GLY
SEQRES  31 B  622  MET TYR GLY TYR TYR PHE LYS TYR GLU MET GLN LEU MET
SEQRES  32 B  622  ALA SER LYS GLY TYR TYR ILE VAL TYR VAL ASN PRO ARG
SEQRES  33 B  622  GLY SER ASN GLY TYR SER GLU ASP PHE ALA LEU ARG VAL
SEQRES  34 B  622  LEU GLU ARG THR GLY LEU GLU ASP PHE GLN ASP ILE LEU
SEQRES  35 B  622  ASN GLY ILE GLU GLU PHE LEU ARG LEU GLU PRO GLN ALA
SEQRES  36 B  622  ASP ARG GLU ARG ILE GLY ILE THR GLY ILE SER TYR GLY
SEQRES  37 B  622  GLY TYR MET THR ASN TRP ALA LEU THR GLN SER ASP LEU
SEQRES  38 B  622  PHE LYS ALA GLY ILE SER GLU ASN GLY ILE SER TYR TRP
SEQRES  39 B  622  LEU THR SER TYR ALA PHE SER ASP ILE GLY LEU TRP PHE
SEQRES  40 B  622  ASP LYS GLU VAL ILE GLY ASP ASN PRO LEU GLU ASN GLU
SEQRES  41 B  622  ASN TYR ARG LYS LEU SER PRO LEU PHE TYR ALA LYS ASN
SEQRES  42 B  622  VAL LYS ALA PRO LEU LEU LEU ILE HIS SER LEU GLU ASP
SEQRES  43 B  622  TYR ARG CYS PRO LEU ASP GLN SER LEU MET PHE TYR HIS
SEQRES  44 B  622  VAL LEU LYS ASP LEU GLY LYS GLU VAL TYR ILE ALA ILE
SEQRES  45 B  622  PHE LYS LYS GLY ALA HIS GLY HIS SER ILE ARG GLY SER
SEQRES  46 B  622  PRO ARG HIS ARG MET LYS ARG TYR LYS LEU PHE MET GLU
SEQRES  47 B  622  PHE PHE GLU ARG LYS LEU LYS LYS TYR GLU GLU GLY PHE
SEQRES  48 B  622  ASP VAL GLU LYS ILE LEU LYS GLU GLU LYS LYS
HET    Y3A  B 701      16
HET     MG  B 702       1
HET     MG  B 703       1
HET     MG  B 704       1
HET     MG  B 705       1
HET     MG  B 706       1
HET    HEZ  B 707       8
HET    HEZ  B 708       8
HET    HEZ  B 709       8
HET    HEZ  B 710       8
HET    HEZ  B 711       8
HET    HEZ  B 712       8
HET    HEZ  B 713       8
HET    HEZ  B 714       8
HET    HEZ  B 715       8
HET    HEZ  B 716       8
HET     CL  B 717       1
HETNAM     Y3A (PHENYLMETHYL) N-[2-OXIDANYLIDENE-2-[[2-OXIDANYLIDENE-
HETNAM   2 Y3A  2-[[(2S,3S)-3-OXIDANYL-1-PHENYL-BUTAN-2-
HETNAM   3 Y3A  YL]AMINO]ETHYL]AMINO]ETHYL]CARBAMATE
HETNAM      MG MAGNESIUM ION
HETNAM     HEZ HEXANE-1,6-DIOL
HETNAM      CL CHLORIDE ION
HETSYN     Y3A Z-GLY-GLY-PHE-CHLOROMETHYL KETONE (BOUND FORM)
FORMUL   2  Y3A    C22 H27 N3 O5
FORMUL   3   MG    5(MG 2+)
FORMUL   8  HEZ    10(C6 H14 O2)
FORMUL  18   CL    CL 1-
FORMUL  19  HOH   *655(H2 O)
HELIX    1   1 LYS B    9  PHE B   14  5                                   6
HELIX    2   2 ASN B  339  LYS B  344  1                                   6
HELIX    3   3 LYS B  397  LYS B  406  1                                  10
HELIX    4   4 SER B  422  ARG B  428  1                                   7
HELIX    5   5 LEU B  435  GLU B  452  1                                  18
HELIX    6   6 SER B  466  SER B  479  1                                  14
HELIX    7   7 TYR B  493  SER B  501  1                                   9
HELIX    8   8 ILE B  503  ILE B  512  1                                  10
HELIX    9   9 GLU B  520  LEU B  525  1                                   6
HELIX   10  10 SER B  526  VAL B  534  5                                   9
HELIX   11  11 LEU B  551  LEU B  564  1                                  14
HELIX   12  12 GLY B  579  GLY B  584  1                                   6
HELIX   13  13 SER B  585  LEU B  604  1                                  20
HELIX   14  14 ASP B  612  LYS B  618  1                                   7
SHEET    1   A 4 TYR B  16  ARG B  23  0
SHEET    2   A 4 LEU B  26  ASN B  35 -1  O  ALA B  28   N  ARG B  21
SHEET    3   A 4 LYS B  40  ASN B  49 -1  O  GLU B  48   N  VAL B  27
SHEET    4   A 4 ARG B  55  GLU B  59 -1  O  LYS B  56   N  ILE B  47
SHEET    1   B 4 SER B  62  ILE B  66  0
SHEET    2   B 4 LYS B  72  ARG B  77 -1  O  ALA B  74   N  ARG B  65
SHEET    3   B 4 GLN B  86  ASP B  91 -1  O  TRP B  88   N  PHE B  75
SHEET    4   B 4 ALA B  97  GLU B 102 -1  O  VAL B 100   N  ILE B  87
SHEET    1   C 4 ILE B 106  TRP B 111  0
SHEET    2   C 4 ARG B 117  LYS B 124 -1  O  VAL B 121   N  ARG B 107
SHEET    3   C 4 LYS B 151  ASP B 158 -1  O  TRP B 155   N  ALA B 120
SHEET    4   C 4 GLU B 163  PRO B 171 -1  O  GLU B 163   N  ASP B 158
SHEET    1   D 2 TRP B 140  PHE B 141  0
SHEET    2   D 2 GLY B 145  PHE B 146 -1  O  GLY B 145   N  PHE B 141
SHEET    1   E 7 SER B 175  HIS B 179  0
SHEET    2   E 7 SER B 182  PRO B 188 -1  O  VAL B 184   N  ILE B 177
SHEET    3   E 7 TRP B 200  LYS B 206 -1  O  TRP B 205   N  ILE B 183
SHEET    4   E 7 GLU B 209  SER B 224 -1  O  LEU B 213   N  ILE B 202
SHEET    5   E 7 ILE B 229  LYS B 234 -1  O  LEU B 230   N  ILE B 222
SHEET    6   E 7 LYS B 245  TYR B 249 -1  O  TYR B 249   N  ILE B 229
SHEET    7   E 7 VAL B 253  GLY B 255 -1  O  LYS B 254   N  ILE B 248
SHEET    1   F 4 VAL B 264  ARG B 270  0
SHEET    2   F 4 LYS B 273  GLU B 280 -1  O  LYS B 273   N  ARG B 270
SHEET    3   F 4 SER B 283  TRP B 289 -1  O  SER B 283   N  GLU B 280
SHEET    4   F 4 VAL B 293  ALA B 297 -1  O  ARG B 294   N  LEU B 288
SHEET    1   G 4 TRP B 302  ALA B 308  0
SHEET    2   G 4 LEU B 312  GLU B 317 -1  O  GLU B 317   N  TRP B 302
SHEET    3   G 4 GLU B 324  TRP B 328 -1  O  TYR B 326   N  TYR B 314
SHEET    4   G 4 GLU B 332  GLN B 334 -1  O  ARG B 333   N  LEU B 327
SHEET    1   H 8 ARG B 352  SER B 358  0
SHEET    2   H 8 LEU B 361  ILE B 368 -1  O  LEU B 363   N  PHE B 356
SHEET    3   H 8 TYR B 409  VAL B 413 -1  O  TYR B 412   N  TRP B 366
SHEET    4   H 8 ALA B 378  VAL B 384  1  N  PRO B 379   O  TYR B 409
SHEET    5   H 8 ALA B 455  ILE B 465  1  O  GLY B 461   N  VAL B 382
SHEET    6   H 8 ALA B 484  GLU B 488  1  O  GLU B 488   N  GLY B 464
SHEET    7   H 8 LEU B 538  SER B 543  1  O  LEU B 539   N  SER B 487
SHEET    8   H 8 VAL B 568  PHE B 573  1  O  PHE B 573   N  HIS B 542
LINK         OG  SER B 466                 C4  Y3A B 701     1555   1555  1.50
LINK         NE2 HIS B 578                 C15 Y3A B 701     1555   1555  1.83
LINK        MG    MG B 706                 O   HOH B 926     1555   1555  2.02
LINK         OE1 GLU B 162                MG    MG B 705     1555   1555  2.04
LINK        MG    MG B 703                 O   HOH B 975     1555   1555  2.07
LINK        MG    MG B 703                 O   HOH B 978     1555   1555  2.07
LINK        MG    MG B 704                 O   HOH B 956     1555   1555  2.07
LINK        MG    MG B 702                 O   HOH B 980     1555   1555  2.08
LINK        MG    MG B 702                 O   HOH B 891     1555   1555  2.08
LINK        MG    MG B 706                 O   HOH B1106     1555   1555  2.09
LINK         OE1 GLU B 127                MG    MG B 706     1555   1555  2.09
LINK        MG    MG B 705                 O   HOH B 876     1555   1555  2.10
LINK        MG    MG B 704                 O   HOH B1229     1555   1555  2.11
LINK         OD2 ASP B 244                MG    MG B 703     1555   1555  2.11
LINK         OE2 GLU B 263                MG    MG B 703     1555   1555  2.12
LINK         OD1 ASP B 424                MG    MG B 702     1555   1555  2.12
LINK         O   LYS B  99                MG    MG B 705     1555   1555  2.16
LINK        MG    MG B 703                 O   HOH B 865     1555   1555  2.16
LINK        MG    MG B 703                 O   HOH B 862     1555   1555  2.18
LINK        MG    MG B 706                 O   HOH B1135     1555   1555  2.23
LINK        MG    MG B 704                 O   HOH B 915     1555   1555  2.25
LINK        MG    MG B 704                 O   HOH B 901     1555   1555  2.28
LINK        MG    MG B 706                 O   HOH B1349     1555   1555  2.30
LINK        MG    MG B 706                 O   HOH B1272     1555   1555  2.32
CISPEP   1 VAL B  137    PRO B  138          0        -9.74
CISPEP   2 GLY B  387    PRO B  388          0        26.05
SITE     1 AC1 11 GLY B 386  GLY B 387  SER B 466  TYR B 467
SITE     2 AC1 11 PHE B 507  ASP B 508  ARG B 548  HIS B 578
SITE     3 AC1 11 HEZ B 710  HEZ B 715  HOH B1455
SITE     1 AC2  6 ASP B 424  HOH B 843  HOH B 891  HOH B 902
SITE     2 AC2  6 HOH B 972  HOH B 980
SITE     1 AC3  6 ASP B 244  GLU B 263  HOH B 862  HOH B 865
SITE     2 AC3  6 HOH B 975  HOH B 978
SITE     1 AC4  6 HOH B 901  HOH B 915  HOH B 942  HOH B 956
SITE     2 AC4  6 HOH B1028  HOH B1229
SITE     1 AC5  6 LYS B  99  GLU B 162  HOH B 849  HOH B 876
SITE     2 AC5  6 HOH B 976  HOH B 977
SITE     1 AC6  6 GLU B 127  HOH B 926  HOH B1106  HOH B1135
SITE     2 AC6  6 HOH B1272  HOH B1349
SITE     1 AC7  6 TYR B 239  ASP B 424  GLU B 431  HOH B 908
SITE     2 AC7  6 HOH B 972  HOH B 984
SITE     1 AC8  5 ASP B 424  GLU B 431  ASN B 521  HOH B 884
SITE     2 AC8  5 HOH B 911
SITE     1 AC9  6 TYR B 220  ILE B 222  TYR B 232  ALA B 265
SITE     2 AC9  6 GLN B 266  ALA B 267
SITE     1 BC1  6 TYR B 395  LYS B 397  HIS B 578  Y3A B 701
SITE     2 BC1  6 HOH B1125  HOH B1403
SITE     1 BC2  3 ASP B 480  LYS B 535  HOH B 998
SITE     1 BC3  6 SER B 543  LEU B 544  ASP B 546  PRO B 550
SITE     2 BC3  6 LEU B 551  HOH B1336
SITE     1 BC4  5 GLU B 518  ARG B 523  PHE B 529  TYR B 530
SITE     2 BC4  5 LYS B 532
SITE     1 BC5  6 GLU B 350  ARG B 352  ILE B 368  LYS B 369
SITE     2 BC5  6 GLU B 452  HOH B1433
SITE     1 BC6  6 PHE B 197  VAL B 240  ARG B 548  Y3A B 701
SITE     2 BC6  6 HOH B1133  HOH B1416
SITE     1 BC7  4 ILE B 260  GLU B 280  TYR B 287  VAL B 293
SITE     1 BC8  3 ARG B  55  HOH B 813  HOH B1182
CRYST1  184.057  184.057  145.529  90.00  90.00 120.00 H 3 2        18
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.005433  0.003137  0.000000        0.00000
SCALE2      0.000000  0.006274  0.000000        0.00000
SCALE3      0.000000  0.000000  0.006871        0.00000
TER    5094      LYS B 618
MASTER      721    0   17   14   37    0   32    6 5776    1  127   48
END