longtext: 4J0K-pdb

content
HEADER    HYDROLASE                               31-JAN-13   4J0K
TITLE     TANNIN ACYL HYDROLASE IN COMPLEX WITH ETHYL GALLATE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: TANNASE;
COMPND   3 CHAIN: A, B;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: LACTOBACILLUS PLANTARUM;
SOURCE   3 ORGANISM_TAXID: 1590;
SOURCE   4 GENE: TANLPL;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   7 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID
KEYWDS    TANNIN, HYDROLASE, HYDROLYSIS
EXPDTA    X-RAY DIFFRACTION
AUTHOR    B.REN,M.WU,Q.WANG,X.PENG,H.WEN,Q.CHEN,W.J.MCKINSTRY
REVDAT   1   22-MAY-13 4J0K    0
JRNL        AUTH   B.REN,M.WU,Q.WANG,X.PENG,H.WEN,W.J.MCKINSTRY,Q.CHEN
JRNL        TITL   CRYSTAL STRUCTURE OF TANNASE FROM LACTOBACILLUS PLANTARUM
JRNL        REF    J.MOL.BIOL.                                2013
JRNL        REFN                   ESSN 1089-8638
JRNL        PMID   23648840
JRNL        DOI    10.1016/J.JMB.2013.04.032
REMARK   2
REMARK   2 RESOLUTION.    2.05 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE: 1.7.1_743)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VICENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-
REMARK   3               : KUNSTLEVE,LI-WEI HUNG,ROBERT IMMORMINO,
REMARK   3               : TOM IOERGER,AIRLIE MCCOY,ERIK MCKEE,NIGEL
REMARK   3               : MORIARTY,REETAL PAI,RANDY READ,JANE
REMARK   3               : RICHARDSON,DAVID RICHARDSON,TOD ROMO,JIM
REMARK   3               : SACCHETTINI,NICHOLAS SAUTER,JACOB SMITH,
REMARK   3               : LAURENT STORONI,TOM TERWILLIGER,PETER
REMARK   3               : ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.05
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.89
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.980
REMARK   3   COMPLETENESS FOR RANGE        (%) : 97.1
REMARK   3   NUMBER OF REFLECTIONS             : 52453
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.199
REMARK   3   R VALUE            (WORKING SET) : 0.197
REMARK   3   FREE R VALUE                     : 0.240
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.080
REMARK   3   FREE R VALUE TEST SET COUNT      : 2663
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 19.8933 -  5.4348    0.98     2634   143  0.2006 0.2342
REMARK   3     2  5.4348 -  4.3284    0.97     2612   143  0.1685 0.1606
REMARK   3     3  4.3284 -  3.7855    0.97     2608   132  0.1599 0.1677
REMARK   3     4  3.7855 -  3.4413    0.97     2627   139  0.1626 0.2030
REMARK   3     5  3.4413 -  3.1958    0.98     2649   138  0.1687 0.2568
REMARK   3     6  3.1958 -  3.0080    0.98     2640   131  0.1843 0.2304
REMARK   3     7  3.0080 -  2.8578    0.98     2643   129  0.1798 0.2276
REMARK   3     8  2.8578 -  2.7338    0.98     2638   141  0.1826 0.2450
REMARK   3     9  2.7338 -  2.6288    0.98     2665   151  0.1803 0.2394
REMARK   3    10  2.6288 -  2.5382    0.98     2612   149  0.1903 0.2566
REMARK   3    11  2.5382 -  2.4590    0.98     2639   147  0.2019 0.2284
REMARK   3    12  2.4590 -  2.3888    0.97     2640   123  0.2040 0.2937
REMARK   3    13  2.3888 -  2.3261    0.97     2620   134  0.2219 0.3019
REMARK   3    14  2.3261 -  2.2694    0.97     2570   155  0.2315 0.2948
REMARK   3    15  2.2694 -  2.2179    0.96     2610   143  0.2421 0.3059
REMARK   3    16  2.2179 -  2.1707    0.97     2601   150  0.2522 0.2752
REMARK   3    17  2.1707 -  2.1273    0.96     2600   133  0.2670 0.3126
REMARK   3    18  2.1273 -  2.0872    0.96     2541   149  0.2917 0.3698
REMARK   3    19  2.0872 -  2.0500    0.95     2641   133  0.3005 0.3327
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.20
REMARK   3   SHRINKAGE RADIUS   : 0.95
REMARK   3   K_SOL              : 0.38
REMARK   3   B_SOL              : 47.24
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.460
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 26.630
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 27.36
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 11.86580
REMARK   3    B22 (A**2) : -11.50130
REMARK   3    B33 (A**2) : -0.36450
REMARK   3    B12 (A**2) : -5.20870
REMARK   3    B13 (A**2) : 8.91240
REMARK   3    B23 (A**2) : -5.00290
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           7353
REMARK   3   ANGLE     :  0.934          10036
REMARK   3   CHIRALITY :  0.065           1124
REMARK   3   PLANARITY :  0.004           1313
REMARK   3   DIHEDRAL  : 15.552           2626
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 6
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: chain 'A' and (resseq 1:228)
REMARK   3    ORIGIN FOR THE GROUP (A):  26.6857 105.1422  80.3327
REMARK   3    T TENSOR
REMARK   3      T11:   0.2629 T22:   0.1501
REMARK   3      T33:   0.1729 T12:  -0.0176
REMARK   3      T13:   0.0215 T23:   0.0016
REMARK   3    L TENSOR
REMARK   3      L11:   0.2837 L22:   0.7214
REMARK   3      L33:   0.5184 L12:  -0.0089
REMARK   3      L13:  -0.0640 L23:   0.2930
REMARK   3    S TENSOR
REMARK   3      S11:   0.0587 S12:  -0.0288 S13:   0.0791
REMARK   3      S21:   0.1519 S22:  -0.0203 S23:  -0.0668
REMARK   3      S31:  -0.0823 S32:  -0.0108 S33:  -0.0432
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: chain 'A' and (resseq 243:340)
REMARK   3    ORIGIN FOR THE GROUP (A):  19.8927  75.1466  70.9977
REMARK   3    T TENSOR
REMARK   3      T11:   0.3263 T22:   0.1368
REMARK   3      T33:   0.1431 T12:  -0.0049
REMARK   3      T13:   0.0421 T23:  -0.0017
REMARK   3    L TENSOR
REMARK   3      L11:   0.4728 L22:   0.4182
REMARK   3      L33:   1.5031 L12:   0.3769
REMARK   3      L13:  -0.3449 L23:  -0.1669
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0076 S12:   0.0066 S13:  -0.0414
REMARK   3      S21:  -0.0281 S22:  -0.0390 S23:   0.0050
REMARK   3      S31:  -0.0112 S32:   0.0022 S33:   0.0570
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: chain 'A' and (resseq 341:469)
REMARK   3    ORIGIN FOR THE GROUP (A):  25.4166  91.1528  70.2757
REMARK   3    T TENSOR
REMARK   3      T11:   0.2840 T22:   0.1355
REMARK   3      T33:   0.1458 T12:  -0.0161
REMARK   3      T13:   0.0115 T23:  -0.0151
REMARK   3    L TENSOR
REMARK   3      L11:   0.3187 L22:   0.4013
REMARK   3      L33:   1.0222 L12:   0.0045
REMARK   3      L13:  -0.0960 L23:  -0.2797
REMARK   3    S TENSOR
REMARK   3      S11:   0.0167 S12:   0.0505 S13:   0.0233
REMARK   3      S21:  -0.0166 S22:   0.0251 S23:  -0.0717
REMARK   3      S31:   0.0335 S32:  -0.0133 S33:  -0.0383
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: chain 'B' and (resseq 1:209)
REMARK   3    ORIGIN FOR THE GROUP (A):  39.7159  42.0041  33.4135
REMARK   3    T TENSOR
REMARK   3      T11:   0.0639 T22:   0.1911
REMARK   3      T33:   0.1931 T12:   0.0244
REMARK   3      T13:  -0.0166 T23:   0.0000
REMARK   3    L TENSOR
REMARK   3      L11:   0.2813 L22:   0.4960
REMARK   3      L33:   0.1794 L12:   0.0947
REMARK   3      L13:  -0.2218 L23:  -0.0877
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0186 S12:   0.0150 S13:  -0.1021
REMARK   3      S21:  -0.1208 S22:  -0.0080 S23:  -0.1193
REMARK   3      S31:   0.0544 S32:  -0.0216 S33:  -0.0027
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: chain 'B' and (resseq 210:311)
REMARK   3    ORIGIN FOR THE GROUP (A):  30.6465  71.4168  40.0091
REMARK   3    T TENSOR
REMARK   3      T11:   0.2790 T22:   0.1741
REMARK   3      T33:   0.2070 T12:  -0.0390
REMARK   3      T13:   0.0400 T23:  -0.0472
REMARK   3    L TENSOR
REMARK   3      L11:   0.9748 L22:   0.6208
REMARK   3      L33:   0.6196 L12:  -0.0766
REMARK   3      L13:  -0.2950 L23:  -0.2007
REMARK   3    S TENSOR
REMARK   3      S11:   0.0106 S12:   0.0811 S13:  -0.0287
REMARK   3      S21:  -0.1741 S22:   0.0194 S23:   0.0376
REMARK   3      S31:   0.0002 S32:  -0.0265 S33:  -0.0555
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: chain 'B' and (resseq 312:469)
REMARK   3    ORIGIN FOR THE GROUP (A):  35.1107  63.6820  40.0282
REMARK   3    T TENSOR
REMARK   3      T11:   0.1118 T22:   0.1823
REMARK   3      T33:   0.1882 T12:   0.0064
REMARK   3      T13:   0.0021 T23:  -0.0003
REMARK   3    L TENSOR
REMARK   3      L11:   0.4743 L22:   0.4431
REMARK   3      L33:   0.3452 L12:   0.0372
REMARK   3      L13:  -0.2231 L23:   0.1654
REMARK   3    S TENSOR
REMARK   3      S11:   0.0544 S12:   0.0061 S13:   0.0714
REMARK   3      S21:  -0.1494 S22:   0.0418 S23:  -0.1022
REMARK   3      S31:  -0.1028 S32:   0.0126 S33:  -0.0508
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4J0K COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 14-FEB-13.
REMARK 100 THE RCSB ID CODE IS RCSB077443.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : NULL
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 5.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : AUSTRALIAN SYNCHROTRON
REMARK 200  BEAMLINE                       : MX2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9537
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 52480
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.050
REMARK 200  RESOLUTION RANGE LOW       (A) : NULL
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.0
REMARK 200  DATA REDUNDANCY                : 2.900
REMARK 200  R MERGE                    (I) : 0.10700
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 8.1000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.05
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.16
REMARK 200  COMPLETENESS FOR SHELL     (%) : 96.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.90
REMARK 200  R MERGE FOR SHELL          (I) : 0.39600
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.500
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: FOURIER SYNTHESIS
REMARK 200 SOFTWARE USED: PHENIX
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 43.76
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.19
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M TRI-SODIUM CITRATE PH 5.5, 20%
REMARK 280  PEG 3000, VAPOR DIFFUSION, SITTING DROP, TEMPERATURE 281K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     TYR A   229
REMARK 465     GLN A   230
REMARK 465     PRO A   231
REMARK 465     VAL A   232
REMARK 465     ALA A   233
REMARK 465     GLY A   234
REMARK 465     THR A   235
REMARK 465     THR A   236
REMARK 465     LYS A   237
REMARK 465     ASN A   238
REMARK 465     GLY A   239
REMARK 465     ARG A   240
REMARK 465     PRO A   241
REMARK 465     LYS A   242
REMARK 465     THR A   402
REMARK 465     THR A   403
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    MET A     1     O    HOH A   799              1.85
REMARK 500   OE1  GLN B   384     O    HOH B   844              1.88
REMARK 500   O    HOH A   813     O    HOH A   835              1.91
REMARK 500   O    HOH A   792     O    HOH A   816              1.93
REMARK 500   O    GLY B   362     O    HOH B   710              1.94
REMARK 500   NZ   LYS A   121     O    HOH A   855              1.96
REMARK 500   O    HOH A   758     O    HOH A   802              2.00
REMARK 500   O    HOH A   813     O    HOH A   814              2.00
REMARK 500   O    HOH A   714     O    HOH A   746              2.01
REMARK 500   O    HOH B   757     O    HOH B   758              2.01
REMARK 500   O    HOH A   827     O    HOH A   828              2.03
REMARK 500   O    HOH A   829     O    HOH A   830              2.08
REMARK 500   NH2  ARG A   378     O    HOH A   694              2.08
REMARK 500   O    GLY B   458     OG   SER B   462              2.09
REMARK 500   O    HOH A   693     O    HOH A   702              2.09
REMARK 500   O    LEU B   150     O    HOH B   717              2.11
REMARK 500   NZ   LYS B   237     O    HOH B   843              2.12
REMARK 500   O    HOH A   709     O    HOH A   817              2.13
REMARK 500   O    HOH B   826     O    HOH B   827              2.13
REMARK 500   O    HOH B   673     O    HOH B   680              2.14
REMARK 500   N    SER B   277     O    HOH B   811              2.14
REMARK 500   O    HOH A   672     O    HOH A   709              2.15
REMARK 500   O    HOH A   614     O    HOH A   719              2.15
REMARK 500   N    ALA B     9     O    HOH B   741              2.16
REMARK 500   O    HOH A   746     O    HOH A   847              2.16
REMARK 500   O    HOH A   693     O    HOH B   863              2.16
REMARK 500   OD2  ASP B   442     O    HOH B   860              2.17
REMARK 500   OE1  GLN B   469     O    HOH B   782              2.17
REMARK 500   NH1  ARG A     4     O    HOH A   799              2.17
REMARK 500   O    HOH A   644     O    HOH A   833              2.17
REMARK 500   OE1  GLN A   147     O    HOH A   812              2.18
REMARK 500   NZ   LYS A   144     O    HOH A   761              2.18
REMARK 500   NE2  GLN B   298     O    HOH B   700              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O1   PEG B   503     O1   PEG B   504     1655     2.11
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    MET A  71       79.64   -116.02
REMARK 500    ASP A  85      129.33   -171.11
REMARK 500    SER A 163     -123.09     61.34
REMARK 500    CYS A 204       61.24     32.01
REMARK 500    THR A 340      145.51     79.78
REMARK 500    ALA A 344     -158.71    -77.19
REMARK 500    ASP A 349       77.48   -100.38
REMARK 500    PHE A 371       21.29   -143.63
REMARK 500    ASN B   3      104.70    -55.28
REMARK 500    PRO B 130       38.79    -96.10
REMARK 500    SER B 163     -123.71     67.67
REMARK 500    CYS B 204       53.82     32.20
REMARK 500    ARG B 228     -156.31   -144.20
REMARK 500    HIS B 315        0.63    -52.89
REMARK 500    THR B 340      155.32     71.94
REMARK 500    ALA B 344     -168.33    -79.17
REMARK 500    PHE B 348      -51.13   -120.35
REMARK 500    SER B 452     -179.76   -171.94
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A 807        DISTANCE =  6.47 ANGSTROMS
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EGR A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PGE A 502
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EGR B 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PG4 B 502
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B 503
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE PEG B 504
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4J0C   RELATED DB: PDB
REMARK 900 RELATED ID: 4J0D   RELATED DB: PDB
REMARK 900 RELATED ID: 4J0G   RELATED DB: PDB
REMARK 900 RELATED ID: 4J0H   RELATED DB: PDB
REMARK 900 RELATED ID: 4J0I   RELATED DB: PDB
REMARK 900 RELATED ID: 4J0J   RELATED DB: PDB
REMARK 900 RELATED ID: 4JUI   RELATED DB: PDB
DBREF  4J0K A    1   469  UNP    B3Y018   B3Y018_LACPN     1    469
DBREF  4J0K B    1   469  UNP    B3Y018   B3Y018_LACPN     1    469
SEQADV 4J0K SER A    0  UNP  B3Y018              EXPRESSION TAG
SEQADV 4J0K SER B    0  UNP  B3Y018              EXPRESSION TAG
SEQRES   1 A  470  SER MET SER ASN ARG LEU ILE PHE ASP ALA ASP TRP LEU
SEQRES   2 A  470  VAL PRO GLU GLN VAL GLN VAL ALA GLY GLN ALA ILE GLN
SEQRES   3 A  470  TYR TYR ALA ALA ARG ASN ILE GLN TYR VAL GLN HIS PRO
SEQRES   4 A  470  VAL ALA ALA ILE GLN VAL LEU ASN VAL PHE VAL PRO ALA
SEQRES   5 A  470  ALA TYR LEU HIS GLY SER SER VAL ASN GLY TYR GLN ARG
SEQRES   6 A  470  ALA THR ALA PRO ILE LEU MET PRO ASN THR VAL GLY GLY
SEQRES   7 A  470  TYR LEU PRO GLY PRO ALA ASP ASP PRO GLN ARG VAL THR
SEQRES   8 A  470  TRP PRO THR ASN ALA GLY THR ILE GLN GLN ALA LEU LYS
SEQRES   9 A  470  ARG GLY TYR VAL VAL VAL ALA ALA GLY ILE ARG GLY ARG
SEQRES  10 A  470  THR THR VAL ASP LYS SER GLY GLN ARG VAL GLY GLN ALA
SEQRES  11 A  470  PRO ALA PHE ILE VAL ASP MET LYS ALA ALA ILE ARG TYR
SEQRES  12 A  470  VAL LYS TYR ASN GLN GLY ARG LEU PRO GLY ASP ALA ASN
SEQRES  13 A  470  ARG ILE ILE THR ASN GLY THR SER ALA GLY GLY ALA THR
SEQRES  14 A  470  SER ALA LEU ALA GLY ALA SER GLY ASN SER ALA TYR PHE
SEQRES  15 A  470  GLU PRO ALA LEU THR ALA LEU GLY ALA ALA PRO ALA THR
SEQRES  16 A  470  ASP ASP ILE PHE ALA VAL SER ALA TYR CYS PRO ILE HIS
SEQRES  17 A  470  ASN LEU GLU HIS ALA ASP MET ALA TYR GLU TRP GLN PHE
SEQRES  18 A  470  ASN GLY ILE ASN ASP TRP HIS ARG TYR GLN PRO VAL ALA
SEQRES  19 A  470  GLY THR THR LYS ASN GLY ARG PRO LYS PHE GLU PRO VAL
SEQRES  20 A  470  SER GLY GLN LEU THR VAL GLU GLU GLN ALA LEU SER LEU
SEQRES  21 A  470  ALA LEU LYS ALA GLN PHE SER THR TYR LEU ASN GLN LEU
SEQRES  22 A  470  LYS LEU THR ALA SER ASP GLY THR HIS LEU THR LEU ASN
SEQRES  23 A  470  GLU ALA GLY MET GLY SER PHE ARG ASP VAL VAL ARG GLN
SEQRES  24 A  470  LEU LEU ILE SER SER ALA GLN THR ALA PHE ASP GLN GLY
SEQRES  25 A  470  THR ASP ILE HIS LYS TYR ALA GLY PHE VAL VAL THR GLY
SEQRES  26 A  470  ASN GLN VAL THR ASP LEU ASP LEU SER ALA TYR LEU LYS
SEQRES  27 A  470  SER LEU THR ARG MET LYS ALA VAL PRO ALA PHE ASP GLN
SEQRES  28 A  470  LEU ASP LEU THR SER PRO GLU ASN ASN LEU PHE GLY ASP
SEQRES  29 A  470  ALA THR ALA LYS ALA LYS HIS PHE THR ALA LEU ALA GLN
SEQRES  30 A  470  THR ARG SER THR VAL THR ALA GLN LEU ALA ASP ALA GLU
SEQRES  31 A  470  LEU ILE GLN ALA ILE ASN PRO LEU SER TYR LEU THR THR
SEQRES  32 A  470  THR SER SER GLN VAL ALA LYS HIS TRP ARG ILE ARG HIS
SEQRES  33 A  470  GLY ALA ALA ASP ARG ASP THR SER PHE ALA ILE PRO ILE
SEQRES  34 A  470  ILE LEU ALA ILE MET LEU GLU ASN HIS GLY TYR GLY ILE
SEQRES  35 A  470  ASP PHE ALA LEU PRO TRP ASP ILE PRO HIS SER GLY ASP
SEQRES  36 A  470  TYR ASP LEU GLY ASP LEU PHE SER TRP ILE ASP GLY LEU
SEQRES  37 A  470  CYS GLN
SEQRES   1 B  470  SER MET SER ASN ARG LEU ILE PHE ASP ALA ASP TRP LEU
SEQRES   2 B  470  VAL PRO GLU GLN VAL GLN VAL ALA GLY GLN ALA ILE GLN
SEQRES   3 B  470  TYR TYR ALA ALA ARG ASN ILE GLN TYR VAL GLN HIS PRO
SEQRES   4 B  470  VAL ALA ALA ILE GLN VAL LEU ASN VAL PHE VAL PRO ALA
SEQRES   5 B  470  ALA TYR LEU HIS GLY SER SER VAL ASN GLY TYR GLN ARG
SEQRES   6 B  470  ALA THR ALA PRO ILE LEU MET PRO ASN THR VAL GLY GLY
SEQRES   7 B  470  TYR LEU PRO GLY PRO ALA ASP ASP PRO GLN ARG VAL THR
SEQRES   8 B  470  TRP PRO THR ASN ALA GLY THR ILE GLN GLN ALA LEU LYS
SEQRES   9 B  470  ARG GLY TYR VAL VAL VAL ALA ALA GLY ILE ARG GLY ARG
SEQRES  10 B  470  THR THR VAL ASP LYS SER GLY GLN ARG VAL GLY GLN ALA
SEQRES  11 B  470  PRO ALA PHE ILE VAL ASP MET LYS ALA ALA ILE ARG TYR
SEQRES  12 B  470  VAL LYS TYR ASN GLN GLY ARG LEU PRO GLY ASP ALA ASN
SEQRES  13 B  470  ARG ILE ILE THR ASN GLY THR SER ALA GLY GLY ALA THR
SEQRES  14 B  470  SER ALA LEU ALA GLY ALA SER GLY ASN SER ALA TYR PHE
SEQRES  15 B  470  GLU PRO ALA LEU THR ALA LEU GLY ALA ALA PRO ALA THR
SEQRES  16 B  470  ASP ASP ILE PHE ALA VAL SER ALA TYR CYS PRO ILE HIS
SEQRES  17 B  470  ASN LEU GLU HIS ALA ASP MET ALA TYR GLU TRP GLN PHE
SEQRES  18 B  470  ASN GLY ILE ASN ASP TRP HIS ARG TYR GLN PRO VAL ALA
SEQRES  19 B  470  GLY THR THR LYS ASN GLY ARG PRO LYS PHE GLU PRO VAL
SEQRES  20 B  470  SER GLY GLN LEU THR VAL GLU GLU GLN ALA LEU SER LEU
SEQRES  21 B  470  ALA LEU LYS ALA GLN PHE SER THR TYR LEU ASN GLN LEU
SEQRES  22 B  470  LYS LEU THR ALA SER ASP GLY THR HIS LEU THR LEU ASN
SEQRES  23 B  470  GLU ALA GLY MET GLY SER PHE ARG ASP VAL VAL ARG GLN
SEQRES  24 B  470  LEU LEU ILE SER SER ALA GLN THR ALA PHE ASP GLN GLY
SEQRES  25 B  470  THR ASP ILE HIS LYS TYR ALA GLY PHE VAL VAL THR GLY
SEQRES  26 B  470  ASN GLN VAL THR ASP LEU ASP LEU SER ALA TYR LEU LYS
SEQRES  27 B  470  SER LEU THR ARG MET LYS ALA VAL PRO ALA PHE ASP GLN
SEQRES  28 B  470  LEU ASP LEU THR SER PRO GLU ASN ASN LEU PHE GLY ASP
SEQRES  29 B  470  ALA THR ALA LYS ALA LYS HIS PHE THR ALA LEU ALA GLN
SEQRES  30 B  470  THR ARG SER THR VAL THR ALA GLN LEU ALA ASP ALA GLU
SEQRES  31 B  470  LEU ILE GLN ALA ILE ASN PRO LEU SER TYR LEU THR THR
SEQRES  32 B  470  THR SER SER GLN VAL ALA LYS HIS TRP ARG ILE ARG HIS
SEQRES  33 B  470  GLY ALA ALA ASP ARG ASP THR SER PHE ALA ILE PRO ILE
SEQRES  34 B  470  ILE LEU ALA ILE MET LEU GLU ASN HIS GLY TYR GLY ILE
SEQRES  35 B  470  ASP PHE ALA LEU PRO TRP ASP ILE PRO HIS SER GLY ASP
SEQRES  36 B  470  TYR ASP LEU GLY ASP LEU PHE SER TRP ILE ASP GLY LEU
SEQRES  37 B  470  CYS GLN
HET    EGR  A 501      14
HET    PGE  A 502      10
HET    EGR  B 501      14
HET    PG4  B 502      13
HET    PEG  B 503       7
HET    PEG  B 504       7
HETNAM     EGR ETHYL 3,4,5-TRIHYDROXYBENZOATE
HETNAM     PGE TRIETHYLENE GLYCOL
HETNAM     PG4 TETRAETHYLENE GLYCOL
HETNAM     PEG DI(HYDROXYETHYL)ETHER
HETSYN     EGR ETHYL GALLATE
FORMUL   3  EGR    2(C9 H10 O5)
FORMUL   4  PGE    C6 H14 O4
FORMUL   6  PG4    C8 H18 O5
FORMUL   7  PEG    2(C4 H10 O3)
FORMUL   9  HOH   *527(H2 O)
HELIX    1   1 ASP A    8  LEU A   12  5                                   5
HELIX    2   2 ALA A   52  HIS A   55  5                                   4
HELIX    3   3 ASN A   94  ARG A  104  1                                  11
HELIX    4   4 PRO A  130  ASN A  146  1                                  17
HELIX    5   5 SER A  163  SER A  175  1                                  13
HELIX    6   6 SER A  178  TYR A  180  5                                   3
HELIX    7   7 PHE A  181  GLY A  189  1                                   9
HELIX    8   8 ASN A  208  GLU A  210  5                                   3
HELIX    9   9 HIS A  211  ASN A  221  1                                  11
HELIX   10  10 THR A  251  LYS A  273  1                                  23
HELIX   11  11 GLY A  290  ASP A  309  1                                  20
HELIX   12  12 ASP A  313  TYR A  317  5                                   5
HELIX   13  13 ASP A  331  THR A  340  1                                  10
HELIX   14  14 SER A  355  PHE A  361  1                                   7
HELIX   15  15 THR A  372  ARG A  378  1                                   7
HELIX   16  16 ASP A  387  ILE A  394  1                                   8
HELIX   17  17 ASN A  395  LEU A  400  5                                   6
HELIX   18  18 PHE A  424  HIS A  437  1                                  14
HELIX   19  19 ASP A  456  GLN A  469  1                                  14
HELIX   20  20 ASP B    8  LEU B   12  5                                   5
HELIX   21  21 ALA B   52  HIS B   55  5                                   4
HELIX   22  22 ASN B   94  GLY B  105  1                                  12
HELIX   23  23 PRO B  130  ASN B  146  1                                  17
HELIX   24  24 ASP B  153  ASN B  155  5                                   3
HELIX   25  25 SER B  163  SER B  175  1                                  13
HELIX   26  26 SER B  178  TYR B  180  5                                   3
HELIX   27  27 PHE B  181  GLY B  189  1                                   9
HELIX   28  28 ASN B  208  ASN B  221  1                                  14
HELIX   29  29 THR B  251  LEU B  272  1                                  22
HELIX   30  30 GLY B  290  ASP B  309  1                                  20
HELIX   31  31 ASP B  313  TYR B  317  5                                   5
HELIX   32  32 ASP B  331  THR B  340  1                                  10
HELIX   33  33 SER B  355  PHE B  361  1                                   7
HELIX   34  34 THR B  372  ARG B  378  1                                   7
HELIX   35  35 ASP B  387  ILE B  394  1                                   8
HELIX   36  36 PRO B  396  THR B  401  1                                   6
HELIX   37  37 PHE B  424  HIS B  437  1                                  14
HELIX   38  38 ASP B  456  GLN B  469  1                                  14
SHEET    1   A 9 VAL A  13  VAL A  19  0
SHEET    2   A 9 GLN A  22  GLN A  33 -1  O  TYR A  26   N  GLU A  15
SHEET    3   A 9 VAL A  44  PRO A  50 -1  O  VAL A  49   N  TYR A  27
SHEET    4   A 9 VAL A 107  ALA A 111 -1  O  VAL A 108   N  PHE A  48
SHEET    5   A 9 ILE A  69  PRO A  72  1  N  LEU A  70   O  VAL A 107
SHEET    6   A 9 ILE A 157  ASN A 160  1  O  ILE A 158   N  MET A  71
SHEET    7   A 9 ALA A 199  TYR A 203  1  O  ALA A 199   N  ILE A 157
SHEET    8   A 9 HIS A 410  ARG A 414  1  O  HIS A 410   N  VAL A 200
SHEET    9   A 9 GLY A 440  ALA A 444  1  O  ASP A 442   N  ILE A 413
SHEET    1   B 2 ASP A 225  ARG A 228  0
SHEET    2   B 2 VAL A 246  GLN A 249 -1  O  VAL A 246   N  ARG A 228
SHEET    1   C 2 PHE A 320  THR A 323  0
SHEET    2   C 2 GLN A 326  LEU A 330 -1  O  GLN A 326   N  THR A 323
SHEET    1   D 9 VAL B  13  VAL B  19  0
SHEET    2   D 9 GLN B  22  GLN B  33 -1  O  TYR B  26   N  GLU B  15
SHEET    3   D 9 VAL B  44  PRO B  50 -1  O  VAL B  47   N  ALA B  29
SHEET    4   D 9 VAL B 107  ALA B 111 -1  O  VAL B 108   N  PHE B  48
SHEET    5   D 9 ILE B  69  PRO B  72  1  N  LEU B  70   O  VAL B 107
SHEET    6   D 9 ILE B 157  THR B 162  1  O  ILE B 158   N  MET B  71
SHEET    7   D 9 ALA B 199  TYR B 203  1  O  TYR B 203   N  GLY B 161
SHEET    8   D 9 HIS B 410  ARG B 414  1  O  HIS B 410   N  VAL B 200
SHEET    9   D 9 GLY B 440  ALA B 444  1  O  ASP B 442   N  TRP B 411
SHEET    1   E 2 ASP B 225  THR B 235  0
SHEET    2   E 2 PRO B 241  GLN B 249 -1  O  LYS B 242   N  VAL B 232
SHEET    1   F 2 PHE B 320  THR B 323  0
SHEET    2   F 2 GLN B 326  LEU B 330 -1  O  GLN B 326   N  THR B 323
CISPEP   1 TRP A   91    PRO A   92          0        -3.96
CISPEP   2 ALA A  129    PRO A  130          0         1.09
CISPEP   3 VAL A  345    PRO A  346          0        -3.55
CISPEP   4 TRP B   91    PRO B   92          0        -3.57
CISPEP   5 ALA B  129    PRO B  130          0        -0.31
CISPEP   6 VAL B  345    PRO B  346          0        -8.51
SITE     1 AC1 10 GLY A  76  GLY A  77  TYR A  78  SER A 163
SITE     2 AC1 10 ALA A 164  ILE A 206  LYS A 343  GLU A 357
SITE     3 AC1 10 ASP A 421  HIS A 451
SITE     1 AC2  9 TRP A 218  GLN A 219  LYS A 262  ALA A 287
SITE     2 AC2  9 GLY A 288  ARG A 293  SER A 333  LEU A 336
SITE     3 AC2  9 HOH A 836
SITE     1 AC3 10 GLY B  76  GLY B  77  TYR B  78  SER B 163
SITE     2 AC3 10 ALA B 164  LYS B 343  GLU B 357  ASP B 421
SITE     3 AC3 10 HIS B 451  HOH B 802
SITE     1 AC4 11 ALA B  65  VAL B 119  SER B 122  GLY B 123
SITE     2 AC4 11 GLN B 124  ARG B 125  ASP B 153  ASN B 155
SITE     3 AC4 11 ARG B 156  HOH B 798  HOH B 832
SITE     1 AC5  6 LEU B 353  ALA B 368  LYS B 409  HIS B 410
SITE     2 AC5  6 PEG B 504  HOH B 713
SITE     1 AC6  6 ASP B 352  THR B 354  VAL B 381  LYS B 409
SITE     2 AC6  6 HIS B 410  PEG B 503
CRYST1   46.099   62.734   83.192  70.20  87.10  79.19 P 1           2
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.021692 -0.004143  0.000310        0.00000
SCALE2      0.000000  0.016228 -0.005782        0.00000
SCALE3      0.000000  0.000000  0.012777        0.00000
TER    3491      GLN A 469
TER    7117      GLN B 469
MASTER      457    0    6   38   26    0   16    6 7646    2   65   74
END