longtext: 4K6K-pdb

content
HEADER    HYDROLASE                               16-APR-13   4K6K
TITLE     CRYSTAL STRUCTURE OF CALB MUTANT D223G FROM CANDIDA ANTARCTICA
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: LIPASE B;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: CALB;
COMPND   5 EC: 3.1.1.3;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: CANDIDA ANTARCTICA;
SOURCE   3 ORGANISM_COMMON: YEAST;
SOURCE   4 ORGANISM_TAXID: 34362;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: ROSETTA AODE3AI;
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PET22B
KEYWDS    LIPASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    J.AN,Y.XIE,Y.FENG,G.WU
REVDAT   1   29-JAN-14 4K6K    0
JRNL        AUTH   Y.XIE,J.AN,G.YANG,G.WU,Y.ZHANG,L.CUI,Y.FENG
JRNL        TITL   ENHANCED ENZYME KINETIC STABILITY BY INCREASING RIGIDITY
JRNL        TITL 2 WITHIN THE ACTIVE SITE
JRNL        REF    J.BIOL.CHEM.                               2014
JRNL        REFN                   ESSN 1083-351X
JRNL        DOI    10.1074/JBC.M113.536045
REMARK   2
REMARK   2 RESOLUTION.    1.60 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.6.0117
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.60
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 50.00
REMARK   3   DATA CUTOFF            (SIGMA(F)) : 0.000
REMARK   3   COMPLETENESS FOR RANGE        (%) : 94.1
REMARK   3   NUMBER OF REFLECTIONS             : 71355
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.236
REMARK   3   R VALUE            (WORKING SET) : 0.235
REMARK   3   FREE R VALUE                     : 0.266
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 3598
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 1.60
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 1.64
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 4664
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 94.17
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2990
REMARK   3   BIN FREE R VALUE SET COUNT          : 250
REMARK   3   BIN FREE R VALUE                    : 0.3470
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 4571
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 36
REMARK   3   SOLVENT ATOMS            : 246
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 21.23
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -3.37000
REMARK   3    B22 (A**2) : 1.82000
REMARK   3    B33 (A**2) : 1.56000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : -0.00000
REMARK   3    B23 (A**2) : -0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.124
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.118
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.082
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 4.674
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.927
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.913
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  4719 ; 0.008 ; 0.020
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  6461 ; 1.163 ; 1.975
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   619 ; 5.438 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   162 ;33.695 ;24.938
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   674 ;12.593 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    16 ;18.774 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   754 ; 0.072 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  3568 ; 0.005 ; 0.022
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     1        A   317
REMARK   3    ORIGIN FOR THE GROUP (A): -11.5778  17.2955 -16.5919
REMARK   3    T TENSOR
REMARK   3      T11:   0.0491 T22:   0.0238
REMARK   3      T33:   0.0233 T12:   0.0197
REMARK   3      T13:   0.0066 T23:   0.0189
REMARK   3    L TENSOR
REMARK   3      L11:   0.9574 L22:   0.4531
REMARK   3      L33:   0.3427 L12:  -0.1507
REMARK   3      L13:  -0.0556 L23:   0.1063
REMARK   3    S TENSOR
REMARK   3      S11:   0.0522 S12:   0.1324 S13:   0.1401
REMARK   3      S21:  -0.0186 S22:  -0.0329 S23:  -0.0359
REMARK   3      S31:  -0.0329 S32:  -0.0376 S33:  -0.0193
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B     1        B   317
REMARK   3    ORIGIN FOR THE GROUP (A): -11.5893  24.0463 -52.1194
REMARK   3    T TENSOR
REMARK   3      T11:   0.0762 T22:   0.0169
REMARK   3      T33:   0.0504 T12:  -0.0070
REMARK   3      T13:   0.0179 T23:   0.0031
REMARK   3    L TENSOR
REMARK   3      L11:   2.5745 L22:   0.8237
REMARK   3      L33:   0.4737 L12:   0.9886
REMARK   3      L13:  -0.2416 L23:  -0.2436
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1201 S12:   0.0511 S13:  -0.3228
REMARK   3      S21:  -0.1068 S22:   0.0774 S23:  -0.1159
REMARK   3      S31:   0.1051 S32:   0.0065 S33:   0.0428
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN USED IF PRESENT IN
REMARK   3  THE INPUT U VALUES RESIDUAL ONLY
REMARK   4
REMARK   4 4K6K COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 02-MAY-13.
REMARK 100 THE RCSB ID CODE IS RCSB078954.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 09-OCT-11
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL17U
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.000
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL
REMARK 200  DATA SCALING SOFTWARE          : HKL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 71410
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.600
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 94.2
REMARK 200  DATA REDUNDANCY                : 6.300
REMARK 200  R MERGE                    (I) : 0.11800
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 13.7000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.60
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.66
REMARK 200  COMPLETENESS FOR SHELL     (%) : 95.8
REMARK 200  DATA REDUNDANCY IN SHELL       : 6.20
REMARK 200  R MERGE FOR SHELL          (I) : 0.45300
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 40.56
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.07
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 25% PEG 3350, 0.2M NAAC, 0.1M TRIS-BIS
REMARK 280  PH6.5, VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 287K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       23.35400
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       69.43500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       43.54550
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       69.43500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       23.35400
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       43.54550
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1970 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 23100 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -20.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     ALA A     0
REMARK 465     ALA A   141
REMARK 465     GLY A   142
REMARK 465     PRO A   143
REMARK 465     LEU A   144
REMARK 465     ASP A   145
REMARK 465     ALA A   146
REMARK 465     LEU A   318
REMARK 465     GLU A   319
REMARK 465     HIS A   320
REMARK 465     HIS A   321
REMARK 465     HIS A   322
REMARK 465     HIS A   323
REMARK 465     HIS A   324
REMARK 465     HIS A   325
REMARK 465     ALA B     0
REMARK 465     ALA B   141
REMARK 465     GLY B   142
REMARK 465     PRO B   143
REMARK 465     LEU B   144
REMARK 465     ASP B   145
REMARK 465     ALA B   146
REMARK 465     GLU B   319
REMARK 465     HIS B   320
REMARK 465     HIS B   321
REMARK 465     HIS B   322
REMARK 465     HIS B   323
REMARK 465     HIS B   324
REMARK 465     HIS B   325
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     LEU B   1    N
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   O1   EDO A   406     C2   EDO A   408     4555     1.37
REMARK 500   C1   EDO A   406     C2   EDO A   408     4555     1.57
REMARK 500   O1   EDO A   406     O2   EDO A   408     4555     1.64
REMARK 500   O2   EDO A   406     O1   EDO A   408     4555     2.08
REMARK 500   O1   EDO A   406     C1   EDO A   408     4555     2.16
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    SER A  29       83.43   -151.09
REMARK 500    ASN A  51      -91.76   -149.03
REMARK 500    SER A 105     -132.55     58.44
REMARK 500    ASP A 134       70.29   -119.19
REMARK 500    VAL A 149       69.43   -116.02
REMARK 500    ASN A 206       -1.60     76.71
REMARK 500    ALA A 305       30.41   -141.94
REMARK 500    ASN B  51      -91.50   -145.49
REMARK 500    SER B 105     -126.14     59.95
REMARK 500    ASP B 134       63.71   -110.96
REMARK 500    ASN B 206       -3.09     76.13
REMARK 500    ALA B 305       31.28   -140.02
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 403
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 404
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 405
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 406
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 407
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO A 408
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE EDO B 401
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4K5Q   RELATED DB: PDB
REMARK 900 RELATED ID: 4K6G   RELATED DB: PDB
REMARK 900 RELATED ID: 4K6H   RELATED DB: PDB
DBREF  4K6K A    1   317  UNP    P41365   LIPB_CANAR      26    342
DBREF  4K6K B    1   317  UNP    P41365   LIPB_CANAR      26    342
SEQADV 4K6K ALA A    0  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K GLY A  223  UNP  P41365    ASP   248 ENGINEERED MUTATION
SEQADV 4K6K LEU A  318  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K GLU A  319  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS A  320  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS A  321  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS A  322  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS A  323  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS A  324  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS A  325  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K ALA B    0  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K GLY B  223  UNP  P41365    ASP   248 ENGINEERED MUTATION
SEQADV 4K6K LEU B  318  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K GLU B  319  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS B  320  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS B  321  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS B  322  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS B  323  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS B  324  UNP  P41365              EXPRESSION TAG
SEQADV 4K6K HIS B  325  UNP  P41365              EXPRESSION TAG
SEQRES   1 A  326  ALA LEU PRO SER GLY SER ASP PRO ALA PHE SER GLN PRO
SEQRES   2 A  326  LYS SER VAL LEU ASP ALA GLY LEU THR CYS GLN GLY ALA
SEQRES   3 A  326  SER PRO SER SER VAL SER LYS PRO ILE LEU LEU VAL PRO
SEQRES   4 A  326  GLY THR GLY THR THR GLY PRO GLN SER PHE ASP SER ASN
SEQRES   5 A  326  TRP ILE PRO LEU SER THR GLN LEU GLY TYR THR PRO CYS
SEQRES   6 A  326  TRP ILE SER PRO PRO PRO PHE MET LEU ASN ASP THR GLN
SEQRES   7 A  326  VAL ASN THR GLU TYR MET VAL ASN ALA ILE THR ALA LEU
SEQRES   8 A  326  TYR ALA GLY SER GLY ASN ASN LYS LEU PRO VAL LEU THR
SEQRES   9 A  326  TRP SER GLN GLY GLY LEU VAL ALA GLN TRP GLY LEU THR
SEQRES  10 A  326  PHE PHE PRO SER ILE ARG SER LYS VAL ASP ARG LEU MET
SEQRES  11 A  326  ALA PHE ALA PRO ASP TYR LYS GLY THR VAL LEU ALA GLY
SEQRES  12 A  326  PRO LEU ASP ALA LEU ALA VAL SER ALA PRO SER VAL TRP
SEQRES  13 A  326  GLN GLN THR THR GLY SER ALA LEU THR THR ALA LEU ARG
SEQRES  14 A  326  ASN ALA GLY GLY LEU THR GLN ILE VAL PRO THR THR ASN
SEQRES  15 A  326  LEU TYR SER ALA THR ASP GLU ILE VAL GLN PRO GLN VAL
SEQRES  16 A  326  SER ASN SER PRO LEU ASP SER SER TYR LEU PHE ASN GLY
SEQRES  17 A  326  LYS ASN VAL GLN ALA GLN ALA VAL CYS GLY PRO LEU PHE
SEQRES  18 A  326  VAL ILE GLY HIS ALA GLY SER LEU THR SER GLN PHE SER
SEQRES  19 A  326  TYR VAL VAL GLY ARG SER ALA LEU ARG SER THR THR GLY
SEQRES  20 A  326  GLN ALA ARG SER ALA ASP TYR GLY ILE THR ASP CYS ASN
SEQRES  21 A  326  PRO LEU PRO ALA ASN ASP LEU THR PRO GLU GLN LYS VAL
SEQRES  22 A  326  ALA ALA ALA ALA LEU LEU ALA PRO ALA ALA ALA ALA ILE
SEQRES  23 A  326  VAL ALA GLY PRO LYS GLN ASN CYS GLU PRO ASP LEU MET
SEQRES  24 A  326  PRO TYR ALA ARG PRO PHE ALA VAL GLY LYS ARG THR CYS
SEQRES  25 A  326  SER GLY ILE VAL THR PRO LEU GLU HIS HIS HIS HIS HIS
SEQRES  26 A  326  HIS
SEQRES   1 B  326  ALA LEU PRO SER GLY SER ASP PRO ALA PHE SER GLN PRO
SEQRES   2 B  326  LYS SER VAL LEU ASP ALA GLY LEU THR CYS GLN GLY ALA
SEQRES   3 B  326  SER PRO SER SER VAL SER LYS PRO ILE LEU LEU VAL PRO
SEQRES   4 B  326  GLY THR GLY THR THR GLY PRO GLN SER PHE ASP SER ASN
SEQRES   5 B  326  TRP ILE PRO LEU SER THR GLN LEU GLY TYR THR PRO CYS
SEQRES   6 B  326  TRP ILE SER PRO PRO PRO PHE MET LEU ASN ASP THR GLN
SEQRES   7 B  326  VAL ASN THR GLU TYR MET VAL ASN ALA ILE THR ALA LEU
SEQRES   8 B  326  TYR ALA GLY SER GLY ASN ASN LYS LEU PRO VAL LEU THR
SEQRES   9 B  326  TRP SER GLN GLY GLY LEU VAL ALA GLN TRP GLY LEU THR
SEQRES  10 B  326  PHE PHE PRO SER ILE ARG SER LYS VAL ASP ARG LEU MET
SEQRES  11 B  326  ALA PHE ALA PRO ASP TYR LYS GLY THR VAL LEU ALA GLY
SEQRES  12 B  326  PRO LEU ASP ALA LEU ALA VAL SER ALA PRO SER VAL TRP
SEQRES  13 B  326  GLN GLN THR THR GLY SER ALA LEU THR THR ALA LEU ARG
SEQRES  14 B  326  ASN ALA GLY GLY LEU THR GLN ILE VAL PRO THR THR ASN
SEQRES  15 B  326  LEU TYR SER ALA THR ASP GLU ILE VAL GLN PRO GLN VAL
SEQRES  16 B  326  SER ASN SER PRO LEU ASP SER SER TYR LEU PHE ASN GLY
SEQRES  17 B  326  LYS ASN VAL GLN ALA GLN ALA VAL CYS GLY PRO LEU PHE
SEQRES  18 B  326  VAL ILE GLY HIS ALA GLY SER LEU THR SER GLN PHE SER
SEQRES  19 B  326  TYR VAL VAL GLY ARG SER ALA LEU ARG SER THR THR GLY
SEQRES  20 B  326  GLN ALA ARG SER ALA ASP TYR GLY ILE THR ASP CYS ASN
SEQRES  21 B  326  PRO LEU PRO ALA ASN ASP LEU THR PRO GLU GLN LYS VAL
SEQRES  22 B  326  ALA ALA ALA ALA LEU LEU ALA PRO ALA ALA ALA ALA ILE
SEQRES  23 B  326  VAL ALA GLY PRO LYS GLN ASN CYS GLU PRO ASP LEU MET
SEQRES  24 B  326  PRO TYR ALA ARG PRO PHE ALA VAL GLY LYS ARG THR CYS
SEQRES  25 B  326  SER GLY ILE VAL THR PRO LEU GLU HIS HIS HIS HIS HIS
SEQRES  26 B  326  HIS
HET    EDO  A 401       4
HET    EDO  A 402       4
HET    EDO  A 403       4
HET    EDO  A 404       4
HET    EDO  A 405       4
HET    EDO  A 406       4
HET    EDO  A 407       4
HET    EDO  A 408       4
HET    EDO  B 401       4
HETNAM     EDO 1,2-ETHANEDIOL
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   3  EDO    9(C2 H6 O2)
FORMUL  12  HOH   *246(H2 O)
HELIX    1   1 PRO A   12  GLY A   19  1                                   8
HELIX    2   2 THR A   43  ASP A   49  1                                   7
HELIX    3   3 ASN A   51  LEU A   59  1                                   9
HELIX    4   4 ASP A   75  SER A   94  1                                  20
HELIX    5   5 SER A  105  PHE A  118  1                                  14
HELIX    6   6 PRO A  119  ARG A  122  5                                   4
HELIX    7   7 ALA A  151  GLN A  157  1                                   7
HELIX    8   8 SER A  161  ALA A  170  1                                  10
HELIX    9   9 ALA A  212  GLY A  217  1                                   6
HELIX   10  10 ALA A  225  SER A  230  1                                   6
HELIX   11  11 SER A  230  SER A  243  1                                  14
HELIX   12  12 ARG A  249  TYR A  253  5                                   5
HELIX   13  13 GLY A  254  CYS A  258  5                                   5
HELIX   14  14 THR A  267  ALA A  276  1                                  10
HELIX   15  15 LEU A  278  GLY A  288  1                                  11
HELIX   16  16 ALA A  301  ALA A  305  5                                   5
HELIX   17  17 PRO B   12  GLY B   19  1                                   8
HELIX   18  18 THR B   43  ASP B   49  1                                   7
HELIX   19  19 ASN B   51  LEU B   59  1                                   9
HELIX   20  20 ASP B   75  SER B   94  1                                  20
HELIX   21  21 SER B  105  PHE B  118  1                                  14
HELIX   22  22 PRO B  119  SER B  123  5                                   5
HELIX   23  23 ALA B  151  GLN B  157  1                                   7
HELIX   24  24 SER B  161  ALA B  170  1                                  10
HELIX   25  25 ALA B  212  GLY B  217  1                                   6
HELIX   26  26 ALA B  225  SER B  230  1                                   6
HELIX   27  27 SER B  230  SER B  243  1                                  14
HELIX   28  28 ARG B  249  TYR B  253  5                                   5
HELIX   29  29 GLY B  254  CYS B  258  5                                   5
HELIX   30  30 THR B  267  ALA B  276  1                                  10
HELIX   31  31 LEU B  277  ALA B  287  1                                  11
HELIX   32  32 ALA B  301  VAL B  306  5                                   6
SHEET    1   A 7 LEU A  20  CYS A  22  0
SHEET    2   A 7 THR A  62  ILE A  66 -1  O  TRP A  65   N  THR A  21
SHEET    3   A 7 PRO A  33  VAL A  37  1  N  ILE A  34   O  CYS A  64
SHEET    4   A 7 LEU A  99  TRP A 104  1  O  LEU A 102   N  LEU A  35
SHEET    5   A 7 VAL A 125  PHE A 131  1  O  MET A 129   N  VAL A 101
SHEET    6   A 7 THR A 179  TYR A 183  1  O  THR A 180   N  ALA A 130
SHEET    7   A 7 LYS A 208  GLN A 211  1  O  VAL A 210   N  ASN A 181
SHEET    1   B 2 ARG A 309  THR A 310  0
SHEET    2   B 2 GLY A 313  ILE A 314 -1  O  GLY A 313   N  THR A 310
SHEET    1   C 7 LEU B  20  CYS B  22  0
SHEET    2   C 7 THR B  62  ILE B  66 -1  O  TRP B  65   N  THR B  21
SHEET    3   C 7 PRO B  33  VAL B  37  1  N  LEU B  36   O  CYS B  64
SHEET    4   C 7 LEU B  99  TRP B 104  1  O  LEU B 102   N  LEU B  35
SHEET    5   C 7 VAL B 125  PHE B 131  1  O  MET B 129   N  VAL B 101
SHEET    6   C 7 THR B 179  TYR B 183  1  O  THR B 180   N  ALA B 130
SHEET    7   C 7 LYS B 208  GLN B 211  1  O  VAL B 210   N  ASN B 181
SHEET    1   D 2 ARG B 309  THR B 310  0
SHEET    2   D 2 GLY B 313  ILE B 314 -1  O  GLY B 313   N  THR B 310
SSBOND   1 CYS A   22    CYS A   64                          1555   1555  2.06
SSBOND   2 CYS A  216    CYS A  258                          1555   1555  2.03
SSBOND   3 CYS A  293    CYS A  311                          1555   1555  2.02
SSBOND   4 CYS B   22    CYS B   64                          1555   1555  2.04
SSBOND   5 CYS B  216    CYS B  258                          1555   1555  2.02
SSBOND   6 CYS B  293    CYS B  311                          1555   1555  2.03
CISPEP   1 PRO A   69    PRO A   70          0        -9.63
CISPEP   2 GLN A  191    PRO A  192          0         0.66
CISPEP   3 PRO B   69    PRO B   70          0       -10.30
CISPEP   4 GLN B  191    PRO B  192          0         5.75
SITE     1 AC1  6 ALA A  18  EDO A 402  EDO A 404  HOH A 577
SITE     2 AC1  6 HOH A 626  EDO B 401
SITE     1 AC2  3 EDO A 401  EDO A 407  EDO A 408
SITE     1 AC3  5 TYR A 203  GLY A 207  LYS A 208  ASN A 209
SITE     2 AC3  5 HOH A 578
SITE     1 AC4  6 TYR A 203  PHE A 205  EDO A 401  HOH A 554
SITE     2 AC4  6 THR B 316  EDO B 401
SITE     1 AC5  4 TYR A  91  ASN A  96  EDO A 408  HOH A 635
SITE     1 AC6  1 EDO A 408
SITE     1 AC7  2 ALA A  18  EDO A 402
SITE     1 AC8  3 EDO A 402  EDO A 405  EDO A 406
SITE     1 AC9  6 EDO A 401  EDO A 404  HOH A 589  PRO B 317
SITE     2 AC9  6 LEU B 318  HOH B 561
CRYST1   46.708   87.091  138.870  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.021410  0.000000  0.000000        0.00000
SCALE2      0.000000  0.011482  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007201        0.00000
TER    2283      PRO A 317
TER    4573      LEU B 318
MASTER      411    0    9   32   18    0   13    6 4853    2   48   52
END