longtext: 4KE7-pdb

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HEADER    HYDROLASE                               25-APR-13   4KE7
TITLE     CRYSTAL STRUCTURE OF MONOGLYCERIDE LIPASE FROM BACILLUS SP. H257 IN
TITLE    2 COMPLEX WITH AN 1-MYRISTOYL GLYCEROL ANALOGUE
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: THERMOSTABLE MONOACYLGLYCEROL LIPASE;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: MGLP;
COMPND   5 EC: 3.1.1.23;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: BACILLUS SP.;
SOURCE   3 ORGANISM_TAXID: 129908;
SOURCE   4 STRAIN: H-257;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 469008;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21(DE3);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PET28A(+)
KEYWDS    ALPHA/BETA HYDROLASE FOLD, MONOGLYCERIDE LIPASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    S.RENGACHARI,P.ASCHAUER,K.GRUBER,I.DREVENY,M.OBERER
REVDAT   1   18-SEP-13 4KE7    0
JRNL        AUTH   S.RENGACHARI,P.ASCHAUER,K.GRUBER,I.DREVENY,M.OBERER
JRNL        TITL   CONFORMATIONAL PLASTICITY AND LIGAND BINDING OF BACTERIAL
JRNL        TITL 2 MONOACYLGLYCEROL LIPASE
JRNL        REF    J.BIOL.CHEM.                               2013
JRNL        REFN                   ESSN 1083-351X
REMARK   2
REMARK   2 RESOLUTION.    1.70 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE: 1.8.2_1309)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VICENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-
REMARK   3               : KUNSTLEVE,LI-WEI HUNG,ROBERT IMMORMINO,
REMARK   3               : TOM IOERGER,AIRLIE MCCOY,ERIK MCKEE,NIGEL
REMARK   3               : MORIARTY,REETAL PAI,RANDY READ,JANE
REMARK   3               : RICHARDSON,DAVID RICHARDSON,TOD ROMO,JIM
REMARK   3               : SACCHETTINI,NICHOLAS SAUTER,JACOB SMITH,
REMARK   3               : LAURENT STORONI,TOM TERWILLIGER,PETER
REMARK   3               : ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.70
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 23.55
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 0.330
REMARK   3   COMPLETENESS FOR RANGE        (%) : 97.5
REMARK   3   NUMBER OF REFLECTIONS             : 47066
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.175
REMARK   3   R VALUE            (WORKING SET) : 0.173
REMARK   3   FREE R VALUE                     : 0.208
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.050
REMARK   3   FREE R VALUE TEST SET COUNT      : 2375
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 23.5574 -  4.3593    0.93     2581   143  0.1638 0.1906
REMARK   3     2  4.3593 -  3.4637    0.97     2652   125  0.1457 0.1525
REMARK   3     3  3.4637 -  3.0269    0.98     2635   164  0.1556 0.1831
REMARK   3     4  3.0269 -  2.7506    0.98     2648   132  0.1655 0.1911
REMARK   3     5  2.7506 -  2.5537    0.99     2663   154  0.1635 0.2013
REMARK   3     6  2.5537 -  2.4033    0.99     2656   157  0.1674 0.2189
REMARK   3     7  2.4033 -  2.2830    0.99     2681   139  0.1657 0.1997
REMARK   3     8  2.2830 -  2.1837    0.93     2495   134  0.2389 0.3009
REMARK   3     9  2.1837 -  2.0997    1.00     2667   143  0.1587 0.1992
REMARK   3    10  2.0997 -  2.0273    0.99     2687   132  0.1589 0.1886
REMARK   3    11  2.0273 -  1.9639    0.99     2680   127  0.1672 0.2083
REMARK   3    12  1.9639 -  1.9078    0.98     2627   125  0.2565 0.3024
REMARK   3    13  1.9078 -  1.8576    0.99     2685   153  0.1707 0.2293
REMARK   3    14  1.8576 -  1.8123    1.00     2663   138  0.1707 0.2102
REMARK   3    15  1.8123 -  1.7711    1.00     2681   137  0.1755 0.2382
REMARK   3    16  1.7711 -  1.7334    1.00     2665   142  0.1904 0.2275
REMARK   3    17  1.7334 -  1.6990    0.87     2325   130  0.2294 0.2941
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.190
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 22.160
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 13.90
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 2.34960
REMARK   3    B22 (A**2) : -1.23700
REMARK   3    B33 (A**2) : -0.48320
REMARK   3    B12 (A**2) : -0.00000
REMARK   3    B13 (A**2) : -0.60550
REMARK   3    B23 (A**2) : -0.00000
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           3959
REMARK   3   ANGLE     :  1.109           5390
REMARK   3   CHIRALITY :  0.070            604
REMARK   3   PLANARITY :  0.006            688
REMARK   3   DIHEDRAL  : 12.057           1431
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 14
REMARK   3   TLS GROUP : 1
REMARK   3    SELECTION: chain 'A' and (resseq 2:69)
REMARK   3    ORIGIN FOR THE GROUP (A):   7.3223  -5.2310  25.8152
REMARK   3    T TENSOR
REMARK   3      T11:   0.0728 T22:   0.0615
REMARK   3      T33:   0.0661 T12:   0.0043
REMARK   3      T13:   0.0100 T23:   0.0049
REMARK   3    L TENSOR
REMARK   3      L11:   2.2699 L22:   1.9442
REMARK   3      L33:   0.9228 L12:   1.2861
REMARK   3      L13:  -0.7869 L23:  -0.5707
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1006 S12:   0.0555 S13:  -0.1902
REMARK   3      S21:  -0.1754 S22:   0.0183 S23:  -0.2346
REMARK   3      S31:   0.1332 S32:   0.0815 S33:   0.0777
REMARK   3   TLS GROUP : 2
REMARK   3    SELECTION: chain 'A' and (resseq 70:131)
REMARK   3    ORIGIN FOR THE GROUP (A):  -4.0375   0.4062  24.7946
REMARK   3    T TENSOR
REMARK   3      T11:   0.0605 T22:   0.0701
REMARK   3      T33:   0.0452 T12:  -0.0021
REMARK   3      T13:  -0.0151 T23:   0.0049
REMARK   3    L TENSOR
REMARK   3      L11:   1.0415 L22:   1.5961
REMARK   3      L33:   1.5838 L12:  -0.3518
REMARK   3      L13:  -0.5045 L23:   0.2340
REMARK   3    S TENSOR
REMARK   3      S11:   0.0354 S12:   0.2062 S13:   0.0104
REMARK   3      S21:  -0.1740 S22:  -0.0090 S23:   0.1207
REMARK   3      S31:   0.0413 S32:  -0.1222 S33:  -0.0206
REMARK   3   TLS GROUP : 3
REMARK   3    SELECTION: chain 'A' and (resseq 132:160)
REMARK   3    ORIGIN FOR THE GROUP (A):  15.4699   8.1375  32.5430
REMARK   3    T TENSOR
REMARK   3      T11:   0.1516 T22:   0.1116
REMARK   3      T33:   0.1251 T12:  -0.0343
REMARK   3      T13:   0.0007 T23:   0.0025
REMARK   3    L TENSOR
REMARK   3      L11:   1.7487 L22:   0.6930
REMARK   3      L33:   3.5024 L12:   0.8817
REMARK   3      L13:  -2.1278 L23:  -0.5553
REMARK   3    S TENSOR
REMARK   3      S11:   0.1258 S12:   0.0129 S13:   0.1433
REMARK   3      S21:  -0.0045 S22:   0.0666 S23:  -0.0070
REMARK   3      S31:  -0.4257 S32:   0.1830 S33:  -0.1190
REMARK   3   TLS GROUP : 4
REMARK   3    SELECTION: chain 'A' and (resseq 161:179)
REMARK   3    ORIGIN FOR THE GROUP (A):   3.6085  12.8567  22.1151
REMARK   3    T TENSOR
REMARK   3      T11:   0.1199 T22:   0.0779
REMARK   3      T33:   0.0865 T12:  -0.0150
REMARK   3      T13:   0.0065 T23:   0.0320
REMARK   3    L TENSOR
REMARK   3      L11:   5.3680 L22:   1.3531
REMARK   3      L33:   1.3182 L12:  -0.5315
REMARK   3      L13:  -1.3115 L23:  -0.0178
REMARK   3    S TENSOR
REMARK   3      S11:   0.1004 S12:   0.2161 S13:   0.4926
REMARK   3      S21:  -0.2217 S22:  -0.0288 S23:  -0.0599
REMARK   3      S31:  -0.1898 S32:   0.0257 S33:  -0.0221
REMARK   3   TLS GROUP : 5
REMARK   3    SELECTION: chain 'A' and (resseq 180:231)
REMARK   3    ORIGIN FOR THE GROUP (A):  -5.8109   6.7135  37.4684
REMARK   3    T TENSOR
REMARK   3      T11:   0.0746 T22:   0.0499
REMARK   3      T33:   0.0308 T12:  -0.0032
REMARK   3      T13:   0.0076 T23:  -0.0058
REMARK   3    L TENSOR
REMARK   3      L11:   3.1354 L22:   1.2881
REMARK   3      L33:   0.9888 L12:  -0.2136
REMARK   3      L13:   0.5960 L23:  -0.3016
REMARK   3    S TENSOR
REMARK   3      S11:   0.0213 S12:  -0.1804 S13:   0.1002
REMARK   3      S21:   0.1424 S22:  -0.0410 S23:   0.0816
REMARK   3      S31:  -0.0458 S32:  -0.0914 S33:   0.0240
REMARK   3   TLS GROUP : 6
REMARK   3    SELECTION: chain 'A' and (resseq 232:250)
REMARK   3    ORIGIN FOR THE GROUP (A):  -5.5485  -6.2125  41.4300
REMARK   3    T TENSOR
REMARK   3      T11:   0.1413 T22:   0.0748
REMARK   3      T33:   0.0685 T12:  -0.0211
REMARK   3      T13:   0.0220 T23:   0.0120
REMARK   3    L TENSOR
REMARK   3      L11:   9.4584 L22:   2.4616
REMARK   3      L33:   2.5137 L12:   3.8249
REMARK   3      L13:   4.1310 L23:   1.0374
REMARK   3    S TENSOR
REMARK   3      S11:   0.2304 S12:  -0.5074 S13:   0.0227
REMARK   3      S21:   0.2129 S22:  -0.2546 S23:   0.1759
REMARK   3      S31:   0.0256 S32:  -0.2043 S33:   0.0289
REMARK   3   TLS GROUP : 7
REMARK   3    SELECTION: chain 'B' and (resseq 2:13)
REMARK   3    ORIGIN FOR THE GROUP (A):  30.2700  -3.7421  56.0220
REMARK   3    T TENSOR
REMARK   3      T11:   0.1061 T22:   0.1052
REMARK   3      T33:   0.1792 T12:   0.0288
REMARK   3      T13:   0.0483 T23:  -0.0298
REMARK   3    L TENSOR
REMARK   3      L11:   4.9357 L22:   4.9775
REMARK   3      L33:   7.0141 L12:   3.7923
REMARK   3      L13:  -4.6452 L23:  -2.8913
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1769 S12:   0.2441 S13:  -0.1972
REMARK   3      S21:  -0.1193 S22:   0.1317 S23:  -0.1839
REMARK   3      S31:   0.1103 S32:   0.1104 S33:  -0.0646
REMARK   3   TLS GROUP : 8
REMARK   3    SELECTION: chain 'B' and (resseq 14:69)
REMARK   3    ORIGIN FOR THE GROUP (A):  19.7787  -4.8482  62.3438
REMARK   3    T TENSOR
REMARK   3      T11:   0.0618 T22:   0.0631
REMARK   3      T33:   0.0436 T12:   0.0080
REMARK   3      T13:   0.0064 T23:  -0.0040
REMARK   3    L TENSOR
REMARK   3      L11:   2.2807 L22:   2.1923
REMARK   3      L33:   1.0535 L12:   1.1031
REMARK   3      L13:  -0.3960 L23:  -0.4305
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0430 S12:   0.1903 S13:  -0.1427
REMARK   3      S21:  -0.0958 S22:   0.0019 S23:  -0.1818
REMARK   3      S31:   0.1144 S32:   0.0430 S33:   0.0436
REMARK   3   TLS GROUP : 9
REMARK   3    SELECTION: chain 'B' and (resseq 70:89)
REMARK   3    ORIGIN FOR THE GROUP (A):  11.3929  -4.4922  54.7917
REMARK   3    T TENSOR
REMARK   3      T11:   0.0774 T22:   0.1167
REMARK   3      T33:   0.0391 T12:   0.0101
REMARK   3      T13:  -0.0181 T23:  -0.0156
REMARK   3    L TENSOR
REMARK   3      L11:   2.5701 L22:   8.9545
REMARK   3      L33:   2.3658 L12:   2.7641
REMARK   3      L13:   0.0642 L23:  -0.6169
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0681 S12:   0.2118 S13:  -0.0049
REMARK   3      S21:  -0.3332 S22:  -0.0132 S23:   0.2779
REMARK   3      S31:   0.0455 S32:  -0.1383 S33:   0.0251
REMARK   3   TLS GROUP : 10
REMARK   3    SELECTION: chain 'B' and (resseq 90:143)
REMARK   3    ORIGIN FOR THE GROUP (A):  12.4410   5.6675  63.2271
REMARK   3    T TENSOR
REMARK   3      T11:   0.0598 T22:   0.0655
REMARK   3      T33:   0.0633 T12:   0.0033
REMARK   3      T13:  -0.0160 T23:   0.0177
REMARK   3    L TENSOR
REMARK   3      L11:   0.8838 L22:   1.0996
REMARK   3      L33:   1.6013 L12:  -0.7575
REMARK   3      L13:  -0.6898 L23:  -0.2829
REMARK   3    S TENSOR
REMARK   3      S11:   0.0601 S12:   0.1744 S13:   0.1924
REMARK   3      S21:  -0.0333 S22:  -0.0063 S23:  -0.0021
REMARK   3      S31:  -0.0809 S32:  -0.0640 S33:  -0.0264
REMARK   3   TLS GROUP : 11
REMARK   3    SELECTION: chain 'B' and (resseq 144:160)
REMARK   3    ORIGIN FOR THE GROUP (A):  29.2605   3.4300  72.9829
REMARK   3    T TENSOR
REMARK   3      T11:   0.1447 T22:   0.1952
REMARK   3      T33:   0.1648 T12:  -0.0396
REMARK   3      T13:  -0.0617 T23:  -0.0138
REMARK   3    L TENSOR
REMARK   3      L11:   1.8962 L22:  -0.0015
REMARK   3      L33:   6.1604 L12:  -0.1584
REMARK   3      L13:  -3.4295 L23:   0.2778
REMARK   3    S TENSOR
REMARK   3      S11:   0.2098 S12:  -0.1129 S13:  -0.0479
REMARK   3      S21:   0.2034 S22:   0.0684 S23:  -0.3278
REMARK   3      S31:  -0.0155 S32:   0.4706 S33:  -0.2243
REMARK   3   TLS GROUP : 12
REMARK   3    SELECTION: chain 'B' and (resseq 161:183)
REMARK   3    ORIGIN FOR THE GROUP (A):  14.4652  12.8493  58.8413
REMARK   3    T TENSOR
REMARK   3      T11:   0.1394 T22:   0.0693
REMARK   3      T33:   0.0842 T12:   0.0123
REMARK   3      T13:   0.0036 T23:   0.0314
REMARK   3    L TENSOR
REMARK   3      L11:   6.2105 L22:   1.6703
REMARK   3      L33:   1.6057 L12:   0.3782
REMARK   3      L13:  -1.9723 L23:  -0.4448
REMARK   3    S TENSOR
REMARK   3      S11:   0.1177 S12:   0.3550 S13:   0.5069
REMARK   3      S21:  -0.1642 S22:   0.0206 S23:  -0.0010
REMARK   3      S31:  -0.1974 S32:  -0.1068 S33:  -0.1093
REMARK   3   TLS GROUP : 13
REMARK   3    SELECTION: chain 'B' and (resseq 184:231)
REMARK   3    ORIGIN FOR THE GROUP (A):   9.6367   6.4441  74.8941
REMARK   3    T TENSOR
REMARK   3      T11:   0.0861 T22:   0.0352
REMARK   3      T33:   0.0542 T12:  -0.0043
REMARK   3      T13:   0.0093 T23:  -0.0103
REMARK   3    L TENSOR
REMARK   3      L11:   2.4165 L22:   1.0199
REMARK   3      L33:   1.4604 L12:  -0.3985
REMARK   3      L13:   0.5605 L23:  -0.4895
REMARK   3    S TENSOR
REMARK   3      S11:   0.0345 S12:  -0.1101 S13:   0.0505
REMARK   3      S21:   0.1493 S22:  -0.0002 S23:   0.1269
REMARK   3      S31:  -0.0881 S32:  -0.0527 S33:  -0.0359
REMARK   3   TLS GROUP : 14
REMARK   3    SELECTION: chain 'B' and (resseq 232:250)
REMARK   3    ORIGIN FOR THE GROUP (A):   8.7607  -6.4541  76.6927
REMARK   3    T TENSOR
REMARK   3      T11:   0.1062 T22:   0.1056
REMARK   3      T33:   0.0910 T12:  -0.0099
REMARK   3      T13:   0.0352 T23:  -0.0242
REMARK   3    L TENSOR
REMARK   3      L11:   2.1152 L22:   2.7767
REMARK   3      L33:   4.2695 L12:   3.0143
REMARK   3      L13:   5.2514 L23:   0.6308
REMARK   3    S TENSOR
REMARK   3      S11:   0.0034 S12:  -0.5960 S13:   0.1833
REMARK   3      S21:   0.3006 S22:  -0.1059 S23:   0.3201
REMARK   3      S31:   0.1435 S32:  -0.3706 S33:   0.0019
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4KE7 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 01-MAY-13.
REMARK 100 THE RCSB ID CODE IS RCSB079226.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 26-SEP-11
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : EMBL/DESY, HAMBURG
REMARK 200  BEAMLINE                       : X13
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.81
REMARK 200  MONOCHROMATOR                  : SI 111, HORIZONTALLY FOCUSSING
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MAR CCD 165 MM
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : MOSFLM
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 47227
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.690
REMARK 200  RESOLUTION RANGE LOW       (A) : 71.307
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 96.9
REMARK 200  DATA REDUNDANCY                : 3.000
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : 0.06500
REMARK 200   FOR THE DATA SET  : 8.7000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.69
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.78
REMARK 200  COMPLETENESS FOR SHELL     (%) : 87.8
REMARK 200  DATA REDUNDANCY IN SHELL       : 2.80
REMARK 200  R MERGE FOR SHELL          (I) : 0.21500
REMARK 200  R SYM FOR SHELL            (I) : 0.21500
REMARK 200   FOR SHELL         : 1.800
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 34.56
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.88
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1 M MES/IMIDAZOLE PH 6.5, 12.5% W/V
REMARK 280  PEG 1000, 12.5% W/V PEG 3350, 12.5% V/V MPD AND 0.02 M ALCOHOLS
REMARK 280  (0.2 M 1,6-HEXANEDIOL, 0.2 M 1-BUTANOL, 0.2 M (RS)-1, 2-
REMARK 280  PROPANEDIOL, 0.2 M 2-PROPANOL, 0.2 M 1,4-BUTANEDIOL, 0.2 M 1,3-
REMARK 280  PROPANEDIOL), VAPOR DIFFUSION, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       35.62000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   -19
REMARK 465     GLY A   -18
REMARK 465     SER A   -17
REMARK 465     SER A   -16
REMARK 465     HIS A   -15
REMARK 465     HIS A   -14
REMARK 465     HIS A   -13
REMARK 465     HIS A   -12
REMARK 465     HIS A   -11
REMARK 465     HIS A   -10
REMARK 465     SER A    -9
REMARK 465     SER A    -8
REMARK 465     GLY A    -7
REMARK 465     LEU A    -6
REMARK 465     VAL A    -5
REMARK 465     PRO A    -4
REMARK 465     ARG A    -3
REMARK 465     GLY A    -2
REMARK 465     SER A    -1
REMARK 465     HIS A     0
REMARK 465     THR A   133
REMARK 465     GLY A   134
REMARK 465     GLY A   135
REMARK 465     GLY A   136
REMARK 465     GLU A   137
REMARK 465     MET B   -19
REMARK 465     GLY B   -18
REMARK 465     SER B   -17
REMARK 465     SER B   -16
REMARK 465     HIS B   -15
REMARK 465     HIS B   -14
REMARK 465     HIS B   -13
REMARK 465     HIS B   -12
REMARK 465     HIS B   -11
REMARK 465     HIS B   -10
REMARK 465     SER B    -9
REMARK 465     SER B    -8
REMARK 465     GLY B    -7
REMARK 465     LEU B    -6
REMARK 465     VAL B    -5
REMARK 465     PRO B    -4
REMARK 465     ARG B    -3
REMARK 465     GLY B    -2
REMARK 465     SER B    -1
REMARK 465     HIS B     0
REMARK 465     MET B     1
REMARK 465     MET B   132
REMARK 465     THR B   133
REMARK 465     GLY B   134
REMARK 465     GLY B   135
REMARK 465     GLY B   136
REMARK 465     GLU B   137
REMARK 465     LEU B   138
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ASN A  18     -117.52   -132.66
REMARK 500    THR A  30       -3.16     70.90
REMARK 500    THR A  60      -80.34   -125.22
REMARK 500    SER A  97     -122.99     65.06
REMARK 500    TYR A 225     -149.74    -98.70
REMARK 500    ASN B  18     -113.67   -133.33
REMARK 500    THR B  30       -3.19     74.13
REMARK 500    THR B  60      -84.75   -121.07
REMARK 500    SER B  97     -120.99     62.95
REMARK 500    TYR B 225     -149.27    -95.80
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE 1QX A 301
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MPD A 302
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE 1QX B 301
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE MPD B 302
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4KE6   RELATED DB: PDB
REMARK 900 RELATED ID: 4KE8   RELATED DB: PDB
REMARK 900 RELATED ID: 4KE9   RELATED DB: PDB
REMARK 900 RELATED ID: 4KEA   RELATED DB: PDB
DBREF  4KE7 A    1   250  UNP    P82597   MGLP_BAC25       1    250
DBREF  4KE7 B    1   250  UNP    P82597   MGLP_BAC25       1    250
SEQADV 4KE7 MET A  -19  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 GLY A  -18  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER A  -17  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER A  -16  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A  -15  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A  -14  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A  -13  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A  -12  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A  -11  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A  -10  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER A   -9  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER A   -8  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 GLY A   -7  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 LEU A   -6  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 VAL A   -5  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 PRO A   -4  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 ARG A   -3  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 GLY A   -2  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER A   -1  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS A    0  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 MET B  -19  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 GLY B  -18  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER B  -17  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER B  -16  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B  -15  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B  -14  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B  -13  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B  -12  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B  -11  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B  -10  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER B   -9  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER B   -8  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 GLY B   -7  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 LEU B   -6  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 VAL B   -5  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 PRO B   -4  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 ARG B   -3  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 GLY B   -2  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 SER B   -1  UNP  P82597              EXPRESSION TAG
SEQADV 4KE7 HIS B    0  UNP  P82597              EXPRESSION TAG
SEQRES   1 A  270  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 A  270  LEU VAL PRO ARG GLY SER HIS MET SER GLU GLN TYR PRO
SEQRES   3 A  270  VAL LEU SER GLY ALA GLU PRO PHE TYR ALA GLU ASN GLY
SEQRES   4 A  270  PRO VAL GLY VAL LEU LEU VAL HIS GLY PHE THR GLY THR
SEQRES   5 A  270  PRO HIS SER MET ARG PRO LEU ALA GLU ALA TYR ALA LYS
SEQRES   6 A  270  ALA GLY TYR THR VAL CYS LEU PRO ARG LEU LYS GLY HIS
SEQRES   7 A  270  GLY THR HIS TYR GLU ASP MET GLU ARG THR THR PHE HIS
SEQRES   8 A  270  ASP TRP VAL ALA SER VAL GLU GLU GLY TYR GLY TRP LEU
SEQRES   9 A  270  LYS GLN ARG CYS GLN THR ILE PHE VAL THR GLY LEU SER
SEQRES  10 A  270  MET GLY GLY THR LEU THR LEU TYR LEU ALA GLU HIS HIS
SEQRES  11 A  270  PRO ASP ILE CYS GLY ILE VAL PRO ILE ASN ALA ALA VAL
SEQRES  12 A  270  ASP ILE PRO ALA ILE ALA ALA GLY MET THR GLY GLY GLY
SEQRES  13 A  270  GLU LEU PRO ARG TYR LEU ASP SER ILE GLY SER ASP LEU
SEQRES  14 A  270  LYS ASN PRO ASP VAL LYS GLU LEU ALA TYR GLU LYS THR
SEQRES  15 A  270  PRO THR ALA SER LEU LEU GLN LEU ALA ARG LEU MET ALA
SEQRES  16 A  270  GLN THR LYS ALA LYS LEU ASP ARG ILE VAL CYS PRO ALA
SEQRES  17 A  270  LEU ILE PHE VAL SER ASP GLU ASP HIS VAL VAL PRO PRO
SEQRES  18 A  270  GLY ASN ALA ASP ILE ILE PHE GLN GLY ILE SER SER THR
SEQRES  19 A  270  GLU LYS GLU ILE VAL ARG LEU ARG ASN SER TYR HIS VAL
SEQRES  20 A  270  ALA THR LEU ASP TYR ASP GLN PRO MET ILE ILE GLU ARG
SEQRES  21 A  270  SER LEU GLU PHE PHE ALA LYS HIS ALA GLY
SEQRES   1 B  270  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 B  270  LEU VAL PRO ARG GLY SER HIS MET SER GLU GLN TYR PRO
SEQRES   3 B  270  VAL LEU SER GLY ALA GLU PRO PHE TYR ALA GLU ASN GLY
SEQRES   4 B  270  PRO VAL GLY VAL LEU LEU VAL HIS GLY PHE THR GLY THR
SEQRES   5 B  270  PRO HIS SER MET ARG PRO LEU ALA GLU ALA TYR ALA LYS
SEQRES   6 B  270  ALA GLY TYR THR VAL CYS LEU PRO ARG LEU LYS GLY HIS
SEQRES   7 B  270  GLY THR HIS TYR GLU ASP MET GLU ARG THR THR PHE HIS
SEQRES   8 B  270  ASP TRP VAL ALA SER VAL GLU GLU GLY TYR GLY TRP LEU
SEQRES   9 B  270  LYS GLN ARG CYS GLN THR ILE PHE VAL THR GLY LEU SER
SEQRES  10 B  270  MET GLY GLY THR LEU THR LEU TYR LEU ALA GLU HIS HIS
SEQRES  11 B  270  PRO ASP ILE CYS GLY ILE VAL PRO ILE ASN ALA ALA VAL
SEQRES  12 B  270  ASP ILE PRO ALA ILE ALA ALA GLY MET THR GLY GLY GLY
SEQRES  13 B  270  GLU LEU PRO ARG TYR LEU ASP SER ILE GLY SER ASP LEU
SEQRES  14 B  270  LYS ASN PRO ASP VAL LYS GLU LEU ALA TYR GLU LYS THR
SEQRES  15 B  270  PRO THR ALA SER LEU LEU GLN LEU ALA ARG LEU MET ALA
SEQRES  16 B  270  GLN THR LYS ALA LYS LEU ASP ARG ILE VAL CYS PRO ALA
SEQRES  17 B  270  LEU ILE PHE VAL SER ASP GLU ASP HIS VAL VAL PRO PRO
SEQRES  18 B  270  GLY ASN ALA ASP ILE ILE PHE GLN GLY ILE SER SER THR
SEQRES  19 B  270  GLU LYS GLU ILE VAL ARG LEU ARG ASN SER TYR HIS VAL
SEQRES  20 B  270  ALA THR LEU ASP TYR ASP GLN PRO MET ILE ILE GLU ARG
SEQRES  21 B  270  SER LEU GLU PHE PHE ALA LYS HIS ALA GLY
MODRES 4KE7 SER B   97  SER  1.58
MODRES 4KE7 SER A   97  SER  1.62
HET    1QX  A 301      21
HET    MPD  A 302       8
HET    1QX  B 301      21
HET    MPD  B 302       8
HETNAM     1QX DODECYL HYDROGEN (S)-(3-AZIDOPROPYL)PHOSPHONATE
HETNAM     MPD (4S)-2-METHYL-2,4-PENTANEDIOL
FORMUL   3  1QX    2(C15 H32 N3 O3 P)
FORMUL   4  MPD    2(C6 H14 O2)
FORMUL   7  HOH   *408(H2 O)
HELIX    1   1 THR A   32  SER A   35  5                                   4
HELIX    2   2 MET A   36  ALA A   46  1                                  11
HELIX    3   3 HIS A   61  ARG A   67  1                                   7
HELIX    4   4 THR A   69  GLN A   86  1                                  18
HELIX    5   5 SER A   97  HIS A  110  1                                  14
HELIX    6   6 ILE A  125  MET A  132  1                                   8
HELIX    7   7 THR A  164  LYS A  180  1                                  17
HELIX    8   8 LEU A  181  ILE A  184  5                                   4
HELIX    9   9 GLY A  202  ILE A  211  1                                  10
HELIX   10  10 VAL A  227  ASP A  231  5                                   5
HELIX   11  11 ASP A  233  ALA A  249  1                                  17
HELIX   12  12 THR B   32  SER B   35  5                                   4
HELIX   13  13 MET B   36  ALA B   46  1                                  11
HELIX   14  14 HIS B   61  ARG B   67  1                                   7
HELIX   15  15 THR B   69  GLN B   86  1                                  18
HELIX   16  16 SER B   97  HIS B  110  1                                  14
HELIX   17  17 ILE B  125  GLY B  131  1                                   7
HELIX   18  18 THR B  164  LYS B  180  1                                  17
HELIX   19  19 LEU B  181  ILE B  184  5                                   4
HELIX   20  20 GLY B  202  ILE B  211  1                                  10
HELIX   21  21 VAL B  227  ASP B  231  5                                   5
HELIX   22  22 ASP B  233  ALA B  249  1                                  17
SHEET    1   A 5 PHE A  14  ALA A  16  0
SHEET    2   A 5 THR A  49  LEU A  52 -1  O  LEU A  52   N  PHE A  14
SHEET    3   A 5 VAL A  21  VAL A  26  1  N  LEU A  25   O  CYS A  51
SHEET    4   A 5 THR A  90  LEU A  96  1  O  THR A  90   N  GLY A  22
SHEET    5   A 5 GLY A 115  ILE A 119  1  O  ILE A 119   N  GLY A  95
SHEET    1   B 2 TYR A 141  ASP A 143  0
SHEET    2   B 2 LYS A 161  PRO A 163 -1  O  THR A 162   N  LEU A 142
SHEET    1   C 2 ALA A 188  SER A 193  0
SHEET    2   C 2 LYS A 216  LEU A 221  1  O  LEU A 221   N  VAL A 192
SHEET    1   D 5 PHE B  14  ALA B  16  0
SHEET    2   D 5 THR B  49  LEU B  52 -1  O  LEU B  52   N  PHE B  14
SHEET    3   D 5 VAL B  21  VAL B  26  1  N  LEU B  25   O  CYS B  51
SHEET    4   D 5 THR B  90  LEU B  96  1  O  THR B  90   N  GLY B  22
SHEET    5   D 5 GLY B 115  ILE B 119  1  O  ILE B 119   N  GLY B  95
SHEET    1   E 2 TYR B 141  ASP B 143  0
SHEET    2   E 2 LYS B 161  PRO B 163 -1  O  THR B 162   N  LEU B 142
SHEET    1   F 2 ALA B 188  SER B 193  0
SHEET    2   F 2 LYS B 216  LEU B 221  1  O  LEU B 221   N  VAL B 192
LINK         OG  SER B  97                 P1  1QX B 301     1555   1555  1.59
LINK         OG  SER A  97                 P1  1QX A 301     1555   1555  1.64
SITE     1 AC1 11 PHE A  29  THR A  30  SER A  97  MET A  98
SITE     2 AC1 11 ILE A 125  ALA A 127  GLY A 131  MET A 132
SITE     3 AC1 11 LEU A 170  VAL A 198  HIS A 226
SITE     1 AC2  6 VAL A  21  ALA A  46  GLY A  47  TYR A  48
SITE     2 AC2  6 HOH A 414  HOH A 442
SITE     1 AC3 10 GLY B  28  PHE B  29  THR B  30  SER B  97
SITE     2 AC3 10 MET B  98  ILE B 125  LEU B 167  LEU B 170
SITE     3 AC3 10 HIS B 226  HOH B 601
SITE     1 AC4  1 ILE B 206
CRYST1   43.750   71.240   72.900  90.00 102.00  90.00 P 1 21 1      4
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.022857  0.000000  0.004858        0.00000
SCALE2      0.000000  0.014037  0.000000        0.00000
SCALE3      0.000000  0.000000  0.014024        0.00000
TER    1910      GLY A 250
TER    3804      GLY B 250
MASTER      530    0    4   22   18    0    9    6 4234    2   60   42
END