longtext: 4Q05-pdb

content
HEADER    HYDROLASE                               31-MAR-14   4Q05
TITLE     CRYSTAL STRUCTURE OF AN ESTERASE E25
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: ESTERASE E25;
COMPND   3 CHAIN: A, B;
COMPND   4 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: UNCULTURED BACTERIUM;
SOURCE   3 ORGANISM_TAXID: 77133;
SOURCE   4 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   5 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    ESTERASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    P.Y.LI,C.Y.LI,Y.Z.ZHANG
REVDAT   1   04-JUN-14 4Q05    0
JRNL        AUTH   P.Y.LI,P.JI,C.Y.LI,Y.ZHANG,G.L.WANG,X.Y.ZHANG,B.B.XIE,
JRNL        AUTH 2 Q.L.QIN,X.L.CHEN,B.C.ZHOU,Y.Z.ZHANG
JRNL        TITL   STRUCTURAL BASIS FOR DIMERIZATION AND CATALYSIS OF A NOVEL
JRNL        TITL 2 ESTERASE FROM THE GTSAG MOTIF SUBFAMILY OF BACTERIAL
JRNL        TITL 3 HORMONE-SENSITIVE LIPASE (HSL) FAMILY
JRNL        REF    J.BIOL.CHEM.                               2014
JRNL        REFN                   ESSN 1083-351X
JRNL        DOI    10.1074/JBC.M114.574913
REMARK   2
REMARK   2 RESOLUTION.    2.05 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE: 1.6.4_486)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-
REMARK   3               : KUNSTLEVE,LI-WEI HUNG,ROBERT IMMORMINO,
REMARK   3               : TOM IOERGER,AIRLIE MCCOY,ERIK MCKEE,NIGEL
REMARK   3               : MORIARTY,REETAL PAI,RANDY READ,JANE
REMARK   3               : RICHARDSON,DAVID RICHARDSON,TOD ROMO,JIM
REMARK   3               : SACCHETTINI,NICHOLAS SAUTER,JACOB SMITH,
REMARK   3               : LAURENT STORONI,TOM TERWILLIGER,PETER
REMARK   3               : ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : OVERALL
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.05
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 19.81
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.960
REMARK   3   COMPLETENESS FOR RANGE        (%) : 97.1
REMARK   3   NUMBER OF REFLECTIONS             : 64693
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.169
REMARK   3   R VALUE            (WORKING SET) : 0.168
REMARK   3   FREE R VALUE                     : 0.190
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.070
REMARK   3   FREE R VALUE TEST SET COUNT      : 3283
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 19.8075 -  5.7782    1.00     2737   163  0.1468 0.1546
REMARK   3     2  5.7782 -  4.6053    1.00     2730   150  0.1353 0.1502
REMARK   3     3  4.6053 -  4.0287    1.00     2771   134  0.1281 0.1324
REMARK   3     4  4.0287 -  3.6629    0.98     2692   148  0.1432 0.1695
REMARK   3     5  3.6629 -  3.4018    0.97     2658   138  0.1521 0.1677
REMARK   3     6  3.4018 -  3.2021    0.96     2622   154  0.1637 0.1991
REMARK   3     7  3.2021 -  3.0423    0.96     2675   136  0.1696 0.1823
REMARK   3     8  3.0423 -  2.9103    0.96     2627   143  0.1870 0.2181
REMARK   3     9  2.9103 -  2.7986    0.95     2629   135  0.1932 0.2016
REMARK   3    10  2.7986 -  2.7023    0.97     2641   158  0.1941 0.1961
REMARK   3    11  2.7023 -  2.6180    0.96     2627   149  0.1966 0.2256
REMARK   3    12  2.6180 -  2.5433    0.96     2651   150  0.1981 0.2428
REMARK   3    13  2.5433 -  2.4765    0.97     2631   138  0.1969 0.2229
REMARK   3    14  2.4765 -  2.4162    0.96     2708   127  0.2015 0.2153
REMARK   3    15  2.4162 -  2.3613    0.97     2618   129  0.1945 0.2005
REMARK   3    16  2.3613 -  2.3112    0.96     2687   139  0.1903 0.2738
REMARK   3    17  2.3112 -  2.2650    0.96     2649   131  0.2031 0.2470
REMARK   3    18  2.2650 -  2.2223    0.98     2703   146  0.1996 0.2970
REMARK   3    19  2.2223 -  2.1827    0.96     2653   146  0.2071 0.2476
REMARK   3    20  2.1827 -  2.1457    0.99     2696   151  0.2104 0.2577
REMARK   3    21  2.1457 -  2.1111    0.96     2653   139  0.2141 0.2786
REMARK   3    22  2.1111 -  2.0787    0.98     2676   139  0.2099 0.2623
REMARK   3    23  2.0787 -  2.0500    0.98     2676   140  0.2242 0.2424
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.20
REMARK   3   SHRINKAGE RADIUS   : 0.95
REMARK   3   K_SOL              : 0.33
REMARK   3   B_SOL              : 36.82
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.210
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 21.390
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -0.05110
REMARK   3    B22 (A**2) : -0.05110
REMARK   3    B33 (A**2) : 0.10230
REMARK   3    B12 (A**2) : -0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           5141
REMARK   3   ANGLE     :  1.037           7025
REMARK   3   CHIRALITY :  0.067            815
REMARK   3   PLANARITY :  0.005            920
REMARK   3   DIHEDRAL  : 11.142           1869
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4Q05 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 03-APR-14.
REMARK 100 THE RCSB ID CODE IS RCSB085437.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 10-NOV-13
REMARK 200  TEMPERATURE           (KELVIN) : 100.0
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : N
REMARK 200  RADIATION SOURCE               : FREE ELECTRON LASER
REMARK 200  BEAMLINE                       : NULL
REMARK 200  X-RAY GENERATOR MODEL          : SSRF BEAMLINE BL17U
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9793
REMARK 200  MONOCHROMATOR                  : DOUBLE CRYSTAL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 64925
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.050
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 0.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 97.4
REMARK 200  DATA REDUNDANCY                : NULL
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.05
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.12
REMARK 200  COMPLETENESS FOR SHELL     (%) : 98.3
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MLPHARE
REMARK 200 STARTING MODEL: SEMET DERIVATIVES
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 65.02
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 3.52
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.2M SODIUM ACETATE TRIHYDRATE, 0.1M
REMARK 280  SODIUM CACODYLATE TRIHYDRATE, 17% PEG8,000, PH 6.5, EVAPORATION,
REMARK 280  TEMPERATURE 293.0K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 31
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -Y,X-Y,Z+1/3
REMARK 290       3555   -X+Y,-X,Z+2/3
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -0.500000 -0.866025  0.000000        0.00000
REMARK 290   SMTRY2   2  0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       16.41567
REMARK 290   SMTRY1   3 -0.500000  0.866025  0.000000        0.00000
REMARK 290   SMTRY2   3 -0.866025 -0.500000  0.000000        0.00000
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       32.83133
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 5330 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 23680 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -71.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   -19
REMARK 465     GLY A   -18
REMARK 465     SER A   -17
REMARK 465     SER A   -16
REMARK 465     HIS A   -15
REMARK 465     HIS A   -14
REMARK 465     HIS A   -13
REMARK 465     HIS A   -12
REMARK 465     HIS A   -11
REMARK 465     HIS A   -10
REMARK 465     SER A    -9
REMARK 465     SER A    -8
REMARK 465     GLY A    -7
REMARK 465     LEU A    -6
REMARK 465     VAL A    -5
REMARK 465     PRO A    -4
REMARK 465     ARG A    -3
REMARK 465     GLY A    -2
REMARK 465     SER A    -1
REMARK 465     HIS A     0
REMARK 465     MET A     1
REMARK 465     THR A     2
REMARK 465     ASN A     3
REMARK 465     GLN A     4
REMARK 465     ASN A     5
REMARK 465     GLU A     6
REMARK 465     SER A     7
REMARK 465     ALA A     8
REMARK 465     ASN A     9
REMARK 465     SER A    10
REMARK 465     LYS A    11
REMARK 465     THR A    12
REMARK 465     GLN A    13
REMARK 465     MET B   -19
REMARK 465     GLY B   -18
REMARK 465     SER B   -17
REMARK 465     SER B   -16
REMARK 465     HIS B   -15
REMARK 465     HIS B   -14
REMARK 465     HIS B   -13
REMARK 465     HIS B   -12
REMARK 465     HIS B   -11
REMARK 465     HIS B   -10
REMARK 465     SER B    -9
REMARK 465     SER B    -8
REMARK 465     GLY B    -7
REMARK 465     LEU B    -6
REMARK 465     VAL B    -5
REMARK 465     PRO B    -4
REMARK 465     ARG B    -3
REMARK 465     GLY B    -2
REMARK 465     SER B    -1
REMARK 465     HIS B     0
REMARK 465     MET B     1
REMARK 465     THR B     2
REMARK 465     ASN B     3
REMARK 465     GLN B     4
REMARK 465     ASN B     5
REMARK 465     GLU B     6
REMARK 465     SER B     7
REMARK 465     ALA B     8
REMARK 465     ASN B     9
REMARK 465     SER B    10
REMARK 465     LYS B    11
REMARK 465     THR B    12
REMARK 465     GLN B    13
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   O    HOH A   780     O    HOH A   785              1.94
REMARK 500   O    HOH B   706     O    HOH B   712              1.95
REMARK 500   O    HOH A   772     O    HOH B   643              1.97
REMARK 500   O    HOH A   724     O    HOH A   733              2.00
REMARK 500   O    HOH A   744     O    HOH A   747              2.03
REMARK 500   OE1  GLN A   169     O    HOH A   741              2.06
REMARK 500   O    HOH B   657     O    HOH B   739              2.06
REMARK 500   O    HOH A   708     O    HOH B   643              2.09
REMARK 500   ND1  HIS B   170     O    HOH B   733              2.10
REMARK 500   O    HOH A   770     O    HOH A   775              2.10
REMARK 500   O    HOH A   683     O    HOH A   722              2.13
REMARK 500   O    HOH A   754     O    HOH A   756              2.14
REMARK 500   O    HOH A   681     O    HOH A   776              2.16
REMARK 500   O    HOH B   650     O    HOH B   698              2.17
REMARK 500   O    HOH B   770     O    HOH B   786              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    PHE A 121      -13.61     73.80
REMARK 500    PRO A 158       34.78    -96.28
REMARK 500    SER A 186     -122.36     45.56
REMARK 500    THR A 214       59.33     30.47
REMARK 500    VAL A 237      -58.08     63.16
REMARK 500    THR A 278     -163.44   -160.62
REMARK 500    PHE B 121      -10.00     68.04
REMARK 500    PRO B 158       37.55    -97.67
REMARK 500    SER B 186     -123.30     44.76
REMARK 500    THR B 214       49.09     32.92
REMARK 500    VAL B 237      -53.81     62.44
REMARK 500    THR B 278     -166.02   -160.21
REMARK 500
REMARK 500 REMARK: NULL
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA B 402  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 TYR B 149   O
REMARK 620 2 ASP B 148   OD1  98.3
REMARK 620 N                    1
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              NA B 403  NA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 LEU B 123   O
REMARK 620 2 GLY B 118   O    82.8
REMARK 620 3 HOH B 555   O   140.7  90.5
REMARK 620 N                    1     2
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA A 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA B 401
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA B 402
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE NA B 403
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 THE SEQUENCE OF THIS PROTEIN WAS NOT AVAILABLE AT THE UNIPROT
REMARK 999 KNOWLEDGEBASE DATABASE (UNIPROTKB) AT THE TIME OF DEPOSITION.
REMARK 999 AUTHORS STATE THAT THE GENEBANK ACCESSION NUMBER IS KJ624992 FOR
REMARK 999 THIS SEQUENCE.
DBREF  4Q05 A  -19   340  PDB    4Q05     4Q05           -19    340
DBREF  4Q05 B  -19   340  PDB    4Q05     4Q05           -19    340
SEQRES   1 A  360  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 A  360  LEU VAL PRO ARG GLY SER HIS MET THR ASN GLN ASN GLU
SEQRES   3 A  360  SER ALA ASN SER LYS THR GLN GLU ASN SER TRP GLN ILE
SEQRES   4 A  360  GLY PRO ARG THR LEU PRO ALA PRO SER GLY ALA SER ASP
SEQRES   5 A  360  VAL LEU TYR ASN ILE ILE SER LYS THR PRO THR PRO VAL
SEQRES   6 A  360  PRO THR ILE ASN LEU ASN LEU VAL PRO ARG THR GLU SER
SEQRES   7 A  360  GLU TRP ARG ALA ALA ILE THR GLN LEU ASP GLU GLY LYS
SEQRES   8 A  360  VAL ASP MET ALA ARG GLU ILE SER LYS GLN LEU SER VAL
SEQRES   9 A  360  SER VAL GLU HIS GLY VAL ILE GLU GLY VAL SER VAL TYR
SEQRES  10 A  360  TYR VAL THR PRO VAL GLU VAL ALA PRO ASP LEU GLU ASP
SEQRES  11 A  360  LYS LEU PHE VAL HIS THR HIS GLY GLY ALA PHE VAL LEU
SEQRES  12 A  360  ASN GLY GLY GLU ALA GLY THR ILE GLU ALA ILE VAL ILE
SEQRES  13 A  360  ALA THR LEU ALA LYS VAL ARG VAL LEU SER ILE ASP TYR
SEQRES  14 A  360  ARG MET PRO PRO SER HIS PRO ALA PRO ALA ALA ARG ASP
SEQRES  15 A  360  ASP VAL PHE THR VAL TYR GLN HIS LEU LEU LYS GLN GLY
SEQRES  16 A  360  SER ALA GLN LYS ILE ALA LEU GLY GLY SER SER GLY GLY
SEQRES  17 A  360  ALA ASN LEU THR MET GLY LEU VAL GLN HIS LEU ILE GLU
SEQRES  18 A  360  GLN GLU VAL ASP LEU PRO GLY ALA LEU PHE LEU GLY THR
SEQRES  19 A  360  PRO GLY ALA ASP MET SER LYS THR GLY ASP SER TYR TYR
SEQRES  20 A  360  ILE ASN ASP GLY ILE ASP ARG ASN LEU VAL THR TYR ASP
SEQRES  21 A  360  GLY PHE LEU GLU ALA ALA VAL ARG LEU TYR ALA ASN GLY
SEQRES  22 A  360  ARG ASP LEU LYS ASP PRO LEU VAL SER PRO LEU TYR GLY
SEQRES  23 A  360  ASP LEU HIS GLY PHE PRO PRO THR PHE LEU ILE THR GLY
SEQRES  24 A  360  THR ARG ASP LEU LEU LEU SER ALA THR VAL ARG THR HIS
SEQRES  25 A  360  ILE LYS LEU ARG GLN SER GLY VAL VAL ALA ASP LEU PHE
SEQRES  26 A  360  VAL TYR GLU GLY ILE ALA HIS GLY ASP TYR ALA VAL ASP
SEQRES  27 A  360  LEU THR ALA PRO GLU THR GLN HIS ALA PHE ALA GLU LEU
SEQRES  28 A  360  ASN ALA PHE LEU LEU GLN HIS LEU ARG
SEQRES   1 B  360  MET GLY SER SER HIS HIS HIS HIS HIS HIS SER SER GLY
SEQRES   2 B  360  LEU VAL PRO ARG GLY SER HIS MET THR ASN GLN ASN GLU
SEQRES   3 B  360  SER ALA ASN SER LYS THR GLN GLU ASN SER TRP GLN ILE
SEQRES   4 B  360  GLY PRO ARG THR LEU PRO ALA PRO SER GLY ALA SER ASP
SEQRES   5 B  360  VAL LEU TYR ASN ILE ILE SER LYS THR PRO THR PRO VAL
SEQRES   6 B  360  PRO THR ILE ASN LEU ASN LEU VAL PRO ARG THR GLU SER
SEQRES   7 B  360  GLU TRP ARG ALA ALA ILE THR GLN LEU ASP GLU GLY LYS
SEQRES   8 B  360  VAL ASP MET ALA ARG GLU ILE SER LYS GLN LEU SER VAL
SEQRES   9 B  360  SER VAL GLU HIS GLY VAL ILE GLU GLY VAL SER VAL TYR
SEQRES  10 B  360  TYR VAL THR PRO VAL GLU VAL ALA PRO ASP LEU GLU ASP
SEQRES  11 B  360  LYS LEU PHE VAL HIS THR HIS GLY GLY ALA PHE VAL LEU
SEQRES  12 B  360  ASN GLY GLY GLU ALA GLY THR ILE GLU ALA ILE VAL ILE
SEQRES  13 B  360  ALA THR LEU ALA LYS VAL ARG VAL LEU SER ILE ASP TYR
SEQRES  14 B  360  ARG MET PRO PRO SER HIS PRO ALA PRO ALA ALA ARG ASP
SEQRES  15 B  360  ASP VAL PHE THR VAL TYR GLN HIS LEU LEU LYS GLN GLY
SEQRES  16 B  360  SER ALA GLN LYS ILE ALA LEU GLY GLY SER SER GLY GLY
SEQRES  17 B  360  ALA ASN LEU THR MET GLY LEU VAL GLN HIS LEU ILE GLU
SEQRES  18 B  360  GLN GLU VAL ASP LEU PRO GLY ALA LEU PHE LEU GLY THR
SEQRES  19 B  360  PRO GLY ALA ASP MET SER LYS THR GLY ASP SER TYR TYR
SEQRES  20 B  360  ILE ASN ASP GLY ILE ASP ARG ASN LEU VAL THR TYR ASP
SEQRES  21 B  360  GLY PHE LEU GLU ALA ALA VAL ARG LEU TYR ALA ASN GLY
SEQRES  22 B  360  ARG ASP LEU LYS ASP PRO LEU VAL SER PRO LEU TYR GLY
SEQRES  23 B  360  ASP LEU HIS GLY PHE PRO PRO THR PHE LEU ILE THR GLY
SEQRES  24 B  360  THR ARG ASP LEU LEU LEU SER ALA THR VAL ARG THR HIS
SEQRES  25 B  360  ILE LYS LEU ARG GLN SER GLY VAL VAL ALA ASP LEU PHE
SEQRES  26 B  360  VAL TYR GLU GLY ILE ALA HIS GLY ASP TYR ALA VAL ASP
SEQRES  27 B  360  LEU THR ALA PRO GLU THR GLN HIS ALA PHE ALA GLU LEU
SEQRES  28 B  360  ASN ALA PHE LEU LEU GLN HIS LEU ARG
HET     NA  A 401       1
HET     NA  B 401       1
HET     NA  B 402       1
HET     NA  B 403       1
HETNAM      NA SODIUM ION
FORMUL   3   NA    4(NA 1+)
FORMUL   7  HOH   *577(H2 O)
HELIX    1   1 SER A   31  LYS A   40  1                                  10
HELIX    2   2 PRO A   46  LEU A   52  1                                   7
HELIX    3   3 THR A   56  GLU A   69  1                                  14
HELIX    4   4 GLY A   70  SER A   83  1                                  14
HELIX    5   5 ALA A  105  GLU A  109  5                                   5
HELIX    6   6 GLY A  125  GLY A  129  5                                   5
HELIX    7   7 THR A  130  LYS A  141  1                                  12
HELIX    8   8 PRO A  158  GLY A  175  1                                  18
HELIX    9   9 SER A  176  GLN A  178  5                                   3
HELIX   10  10 SER A  186  GLN A  202  1                                  17
HELIX   11  11 GLY A  223  ASN A  229  1                                   7
HELIX   12  12 GLY A  241  ASN A  252  1                                  12
HELIX   13  13 SER A  262  GLY A  266  5                                   5
HELIX   14  14 LEU A  284  SER A  298  1                                  15
HELIX   15  15 GLY A  313  VAL A  317  5                                   5
HELIX   16  16 ALA A  321  LEU A  339  1                                  19
HELIX   17  17 SER B   31  LYS B   40  1                                  10
HELIX   18  18 PRO B   46  LEU B   52  1                                   7
HELIX   19  19 THR B   56  GLU B   69  1                                  14
HELIX   20  20 GLY B   70  SER B   83  1                                  14
HELIX   21  21 ALA B  105  GLU B  109  5                                   5
HELIX   22  22 GLY B  125  ALA B  128  5                                   4
HELIX   23  23 GLY B  129  LYS B  141  1                                  13
HELIX   24  24 PRO B  158  GLY B  175  1                                  18
HELIX   25  25 SER B  176  GLN B  178  5                                   3
HELIX   26  26 SER B  186  GLN B  202  1                                  17
HELIX   27  27 GLY B  223  ASN B  229  1                                   7
HELIX   28  28 GLY B  241  ASN B  252  1                                  12
HELIX   29  29 SER B  262  GLY B  266  5                                   5
HELIX   30  30 LEU B  284  SER B  298  1                                  15
HELIX   31  31 GLY B  313  VAL B  317  5                                   5
HELIX   32  32 ALA B  321  LEU B  339  1                                  19
SHEET    1   A 2 TRP A  17  ILE A  19  0
SHEET    2   A 2 ARG B  22  LEU B  24 -1  O  ARG B  22   N  ILE A  19
SHEET    1   B 2 ARG A  22  LEU A  24  0
SHEET    2   B 2 TRP B  17  ILE B  19 -1  O  ILE B  19   N  ARG A  22
SHEET    1   C 8 SER A  85  ILE A  91  0
SHEET    2   C 8 VAL A  94  THR A 100 -1  O  TYR A  98   N  GLU A  87
SHEET    3   C 8 VAL A 144  ASP A 148 -1  O  VAL A 144   N  VAL A  99
SHEET    4   C 8 LEU A 112  THR A 116  1  N  HIS A 115   O  ILE A 147
SHEET    5   C 8 ILE A 180  SER A 185  1  O  ALA A 181   N  VAL A 114
SHEET    6   C 8 ALA A 209  PRO A 215  1  O  GLY A 213   N  GLY A 184
SHEET    7   C 8 THR A 274  GLY A 279  1  O  PHE A 275   N  LEU A 212
SHEET    8   C 8 ALA A 302  TYR A 307  1  O  ASP A 303   N  LEU A 276
SHEET    1   D 8 SER B  85  ILE B  91  0
SHEET    2   D 8 VAL B  94  THR B 100 -1  O  TYR B  98   N  GLU B  87
SHEET    3   D 8 VAL B 144  ASP B 148 -1  O  VAL B 144   N  VAL B  99
SHEET    4   D 8 LEU B 112  THR B 116  1  N  HIS B 115   O  ILE B 147
SHEET    5   D 8 ILE B 180  SER B 185  1  O  ALA B 181   N  VAL B 114
SHEET    6   D 8 ALA B 209  PRO B 215  1  O  GLY B 213   N  GLY B 184
SHEET    7   D 8 THR B 274  GLY B 279  1  O  PHE B 275   N  LEU B 212
SHEET    8   D 8 ALA B 302  TYR B 307  1  O  ASP B 303   N  LEU B 276
LINK         O   TYR B 149                NA    NA B 402     1555   1555  2.73
LINK         O   LEU B 123                NA    NA B 403     1555   1555  2.75
LINK         OD1 ASP B 148                NA    NA B 402     1555   1555  2.77
LINK         O   VAL B 306                NA    NA A 401     1555   1555  2.88
LINK         O   VAL A 306                NA    NA B 401     1555   1555  2.91
LINK         O   GLY B 118                NA    NA B 403     1555   1555  3.03
LINK        NA    NA B 403                 O   HOH B 555     1555   1555  2.79
CISPEP   1 PRO A  152    PRO A  153          0         4.07
CISPEP   2 ALA A  157    PRO A  158          0         3.20
CISPEP   3 PRO B  152    PRO B  153          0         5.26
CISPEP   4 ALA B  157    PRO B  158          0         3.86
SITE     1 AC1  5 VAL A 289  HIS A 292  ILE A 293  LEU A 304
SITE     2 AC1  5 VAL B 306
SITE     1 AC2  5 VAL A 306  VAL B 289  HIS B 292  ILE B 293
SITE     2 AC2  5 LEU B 304
SITE     1 AC3  4 HIS B 117  ASP B 148  TYR B 149  MET B 151
SITE     1 AC4  8 HIS B 117  GLY B 118  GLY B 119  PHE B 121
SITE     2 AC4  8 VAL B 122  LEU B 123  MET B 151  HOH B 555
CRYST1  138.796  138.796   49.247  90.00  90.00 120.00 P 31          6
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.007205  0.004160  0.000000        0.00000
SCALE2      0.000000  0.008319  0.000000        0.00000
SCALE3      0.000000  0.000000  0.020306        0.00000
TER    2509      ARG A 340
TER    5026      ARG B 340
MASTER      380    0    4   32   20    0    7    6 5569    2   11   56
END