longtext: 4QLA-pdb

content
HEADER    HYDROLASE                               11-JUN-14   4QLA
TITLE     CRYSTAL STRUCTURE OF JUVENILE HORMONE EPOXIDE HYDROLASE FROM THE
TITLE    2 SILKWORM BOMBYX MORI
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: JUVENILE HORMONE EPOXIDE HYDROLASE;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: UNP RESIDUES 23-461;
COMPND   5 SYNONYM: BOMMO-JHEH;
COMPND   6 EC: 3.3.2.9;
COMPND   7 ENGINEERED: YES;
COMPND   8 MUTATION: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: BOMBYX MORI;
SOURCE   3 ORGANISM_COMMON: SILK MOTH, SILKWORM;
SOURCE   4 ORGANISM_TAXID: 7091;
SOURCE   5 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   6 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   7 EXPRESSION_SYSTEM_STRAIN: BL21 (DE3);
SOURCE   8 EXPRESSION_SYSTEM_VECTOR_TYPE: PLASMID;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PET29A
KEYWDS    ALPHA/BETA HYDROLASE, EPOXIDE HYDROLASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    K.ZHOU,N.JIA,C.HU,Y.L.JIANG,J.P.YANG,Y.CHEN,S.LI,C.Z.ZHOU
REVDAT   1   03-SEP-14 4QLA    0
JRNL        AUTH   K.ZHOU,N.JIA,C.HU,Y.L.JIANG,J.P.YANG,Y.CHEN,S.LI,W.F.LI,
JRNL        AUTH 2 C.Z.ZHOU
JRNL        TITL   CRYSTAL STRUCTURE OF JUVENILE HORMONE EPOXIDE HYDROLASE FROM
JRNL        TITL 2 THE SILKWORM BOMBYX MORI
JRNL        REF    PROTEINS                                   2014
JRNL        REFN                   ESSN 1097-0134
JRNL        PMID   25143157
JRNL        DOI    10.1002/PROT.24676
REMARK   2
REMARK   2 RESOLUTION.    2.30 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.6.0117
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.30
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 39.39
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 98.9
REMARK   3   NUMBER OF REFLECTIONS             : 36297
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.206
REMARK   3   R VALUE            (WORKING SET) : 0.203
REMARK   3   FREE R VALUE                     : 0.251
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 1907
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.30
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.36
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2406
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 92.29
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2840
REMARK   3   BIN FREE R VALUE SET COUNT          : 131
REMARK   3   BIN FREE R VALUE                    : 0.3510
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 6491
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 32
REMARK   3   SOLVENT ATOMS            : 159
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 32.19
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 41.62
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 1.76000
REMARK   3    B22 (A**2) : -2.64000
REMARK   3    B33 (A**2) : 5.24000
REMARK   3    B12 (A**2) : -0.00000
REMARK   3    B13 (A**2) : 4.04000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.400
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.253
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.193
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 7.974
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.948
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.923
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  6712 ; 0.006 ; 0.020
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  9087 ; 1.067 ; 1.970
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   800 ; 5.439 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   288 ;37.805 ;23.785
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1153 ;16.767 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    29 ;12.561 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   970 ; 0.079 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  5036 ; 0.004 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN USED IF PRESENT IN
REMARK   3  THE INPUT
REMARK   4
REMARK   4 4QLA COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 25-JUN-14.
REMARK 100 THE RCSB ID CODE IS RCSB086197.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 26-SEP-10
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 8.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL17U
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97931
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : MAR SCANNER 345 MM PLATE
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 38204
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.300
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.2
REMARK 200  DATA REDUNDANCY                : 4.000
REMARK 200  R MERGE                    (I) : 0.10800
REMARK 200  R SYM                      (I) : 0.10800
REMARK 200   FOR THE DATA SET  : 11.7600
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.30
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.38
REMARK 200  COMPLETENESS FOR SHELL     (%) : 96.7
REMARK 200  DATA REDUNDANCY IN SHELL       : 3.50
REMARK 200  R MERGE FOR SHELL          (I) : 0.50000
REMARK 200  R SYM FOR SHELL            (I) : 0.50000
REMARK 200   FOR SHELL         : 2.880
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: MOLREP
REMARK 200 STARTING MODEL: 1QO7
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 42.32
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.13
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.1M TRIS-HCL, PH 8.0, 19%
REMARK 280  POLYETHYLENE GLYCOL 2000, VAPOR DIFFUSION, HANGING DROP,
REMARK 280  TEMPERATURE 288K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: C 1 2 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y,-Z
REMARK 290       3555   X+1/2,Y+1/2,Z
REMARK 290       4555   -X+1/2,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   3  1.000000  0.000000  0.000000       80.70200
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       25.73300
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   4 -1.000000  0.000000  0.000000       80.70200
REMARK 290   SMTRY2   4  0.000000  1.000000  0.000000       25.73300
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 5950 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 32380 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -22.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A    15
REMARK 465     HIS A    16
REMARK 465     HIS A    17
REMARK 465     HIS A    18
REMARK 465     HIS A    19
REMARK 465     HIS A    20
REMARK 465     HIS A    21
REMARK 465     GLY A    22
REMARK 465     VAL A    23
REMARK 465     LEU A    24
REMARK 465     LYS A    25
REMARK 465     SER A    26
REMARK 465     PRO A    27
REMARK 465     PRO A    28
REMARK 465     PRO A    29
REMARK 465     MET A    30
REMARK 465     PRO A    31
REMARK 465     LYS A    32
REMARK 465     VAL A   457
REMARK 465     LYS A   458
REMARK 465     THR A   459
REMARK 465     GLU A   460
REMARK 465     LEU A   461
REMARK 465     MET B    15
REMARK 465     HIS B    16
REMARK 465     HIS B    17
REMARK 465     HIS B    18
REMARK 465     HIS B    19
REMARK 465     HIS B    20
REMARK 465     HIS B    21
REMARK 465     GLY B    22
REMARK 465     VAL B    23
REMARK 465     LEU B    24
REMARK 465     LYS B    25
REMARK 465     SER B    26
REMARK 465     PRO B    27
REMARK 465     PRO B    28
REMARK 465     PRO B    29
REMARK 465     MET B    30
REMARK 465     PRO B    31
REMARK 465     LYS B    32
REMARK 465     LEU B   253
REMARK 465     THR B   254
REMARK 465     LEU B   255
REMARK 465     SER B   256
REMARK 465     PRO B   257
REMARK 465     ALA B   258
REMARK 465     ALA B   259
REMARK 465     THR B   260
REMARK 465     PHE B   261
REMARK 465     LEU B   262
REMARK 465     GLU B   263
REMARK 465     PHE B   264
REMARK 465     VAL B   265
REMARK 465     GLY B   266
REMARK 465     ALA B   267
REMARK 465     LEU B   268
REMARK 465     PHE B   269
REMARK 465     PRO B   270
REMARK 465     SER B   271
REMARK 465     LEU B   272
REMARK 465     ILE B   273
REMARK 465     VAL B   274
REMARK 465     GLU B   275
REMARK 465     PRO B   276
REMARK 465     GLU B   277
REMARK 465     LEU B   278
REMARK 465     ALA B   279
REMARK 465     ASN B   280
REMARK 465     ARG B   281
REMARK 465     LEU B   282
REMARK 465     TYR B   283
REMARK 465     PRO B   284
REMARK 465     LEU B   285
REMARK 465     SER B   286
REMARK 465     GLU B   287
REMARK 465     LYS B   288
REMARK 465     TYR B   289
REMARK 465     SER B   290
REMARK 465     ASP B   383
REMARK 465     LEU B   384
REMARK 465     LYS B   385
REMARK 465     LEU B   386
REMARK 465     ASP B   387
REMARK 465     GLU B   388
REMARK 465     VAL B   457
REMARK 465     LYS B   458
REMARK 465     THR B   459
REMARK 465     GLU B   460
REMARK 465     LEU B   461
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    LYS A  73       38.25    -89.21
REMARK 500    PRO A 155       46.90    -96.99
REMARK 500    SER A 157     -173.84   -172.64
REMARK 500    GLU A 160      -39.32    -39.56
REMARK 500    LYS A 173      -17.21     93.04
REMARK 500    ASP A 227     -121.62     51.64
REMARK 500    MET A 251       53.88   -145.79
REMARK 500    LEU A 255       48.84   -107.45
REMARK 500    SER A 271        2.32    -64.29
REMARK 500    GLU B  37      129.46    -36.83
REMARK 500    LYS B  47       32.48    -89.68
REMARK 500    PRO B 155       49.70    -98.17
REMARK 500    SER B 157     -177.92   -175.92
REMARK 500    ASP B 227     -134.65     58.73
REMARK 500    SER B 329      -71.72    -73.79
REMARK 500    ASN B 333      109.27   -166.69
REMARK 500    ARG B 347      -41.36   -130.53
REMARK 500    PHE B 377       36.00    -96.74
REMARK 500    GLN B 392       32.32    -99.44
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE 1PE A 501
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE 1PE B 501
DBREF  4QLA A   23   461  UNP    H9JWL6   H9JWL6_BOMMO    23    461
DBREF  4QLA B   23   461  UNP    H9JWL6   H9JWL6_BOMMO    23    461
SEQADV 4QLA MET A   15  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS A   16  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS A   17  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS A   18  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS A   19  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS A   20  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS A   21  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA GLY A   22  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA GLY A  142  UNP  H9JWL6    ASP   142 ENGINEERED MUTATION
SEQADV 4QLA LEU A  336  UNP  H9JWL6    PHE   336 ENGINEERED MUTATION
SEQADV 4QLA HIS A  402  UNP  H9JWL6    TYR   402 ENGINEERED MUTATION
SEQADV 4QLA MET A  414  UNP  H9JWL6    LEU   414 ENGINEERED MUTATION
SEQADV 4QLA ALA A  433  UNP  H9JWL6    SER   433 ENGINEERED MUTATION
SEQADV 4QLA MET B   15  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS B   16  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS B   17  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS B   18  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS B   19  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS B   20  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA HIS B   21  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA GLY B   22  UNP  H9JWL6              EXPRESSION TAG
SEQADV 4QLA GLY B  142  UNP  H9JWL6    ASP   142 ENGINEERED MUTATION
SEQADV 4QLA LEU B  336  UNP  H9JWL6    PHE   336 ENGINEERED MUTATION
SEQADV 4QLA HIS B  402  UNP  H9JWL6    TYR   402 ENGINEERED MUTATION
SEQADV 4QLA MET B  414  UNP  H9JWL6    LEU   414 ENGINEERED MUTATION
SEQADV 4QLA ALA B  433  UNP  H9JWL6    SER   433 ENGINEERED MUTATION
SEQRES   1 A  447  MET HIS HIS HIS HIS HIS HIS GLY VAL LEU LYS SER PRO
SEQRES   2 A  447  PRO PRO MET PRO LYS LEU ASP LEU GLU GLU TRP TRP GLY
SEQRES   3 A  447  PRO PRO GLU LEU LYS GLN LYS GLN ASP THR SER ILE LYS
SEQRES   4 A  447  PRO PHE GLU ILE THR PHE SER GLU THR MET VAL LYS GLU
SEQRES   5 A  447  LEU LYS GLU ARG ILE LYS LYS ARG ARG PRO PHE ALA PRO
SEQRES   6 A  447  PRO LEU GLU GLY VAL GLY PHE LYS TYR GLY PHE ASN SER
SEQRES   7 A  447  LYS GLN LEU ASP SER TRP LEU LYS TYR TRP ALA GLU GLU
SEQRES   8 A  447  TYR PRO PHE ALA GLU ARG GLN LYS PHE LEU ASN GLN TYR
SEQRES   9 A  447  PRO HIS PHE LYS THR ASN ILE GLN GLY LEU ASN ILE HIS
SEQRES  10 A  447  PHE MET ARG ILE THR PRO LYS VAL PRO LYS GLY VAL GLU
SEQRES  11 A  447  ILE VAL PRO LEU LEU LEU LEU HIS GLY TRP PRO GLY SER
SEQRES  12 A  447  VAL ARG GLU PHE TYR GLU ALA ILE PRO HIS LEU THR ALA
SEQRES  13 A  447  VAL SER LYS ASP ARG ASN PHE ALA LEU GLU ILE ILE ALA
SEQRES  14 A  447  PRO SER LEU PRO GLY TYR GLY PHE SER ASP ALA ALA VAL
SEQRES  15 A  447  ARG PRO GLY LEU ALA ALA ALA GLU VAL ALA VAL ILE PHE
SEQRES  16 A  447  LYS ASN LEU MET ALA ARG LEU GLY TYR LYS GLN TYR TYR
SEQRES  17 A  447  VAL GLN GLY GLY ASP TRP GLY ALA LEU ILE GLY SER ALA
SEQRES  18 A  447  MET ALA THR PHE PHE PRO LYS GLU ILE ILE GLY PHE HIS
SEQRES  19 A  447  SER ASN MET ALA LEU THR LEU SER PRO ALA ALA THR PHE
SEQRES  20 A  447  LEU GLU PHE VAL GLY ALA LEU PHE PRO SER LEU ILE VAL
SEQRES  21 A  447  GLU PRO GLU LEU ALA ASN ARG LEU TYR PRO LEU SER GLU
SEQRES  22 A  447  LYS TYR SER THR LEU LEU GLU GLU LEU GLY TYR MET HIS
SEQRES  23 A  447  ILE GLN ALA THR LYS PRO ASP THR VAL GLY ILE GLY LEU
SEQRES  24 A  447  THR ASP SER PRO ALA GLY LEU LEU ALA TYR ILE LEU GLU
SEQRES  25 A  447  LYS PHE SER THR TRP THR ASN PRO ASP LEU ARG SER LYS
SEQRES  26 A  447  GLU ASP GLY GLY LEU SER TYR ARG TRP THR LYS ASP GLN
SEQRES  27 A  447  LEU ILE ASP ASN LEU MET LEU TYR TRP SER THR LYS SER
SEQRES  28 A  447  ILE VAL THR SER MET ARG LEU TYR ALA GLU SER PHE SER
SEQRES  29 A  447  SER ARG HIS PHE ASP LEU LYS LEU ASP GLU ILE GLN VAL
SEQRES  30 A  447  GLN VAL PRO THR TRP VAL LEU GLN ALA LYS HIS GLU LEU
SEQRES  31 A  447  ALA TYR GLN PRO PRO CYS ILE LEU LYS MET LYS TYR PRO
SEQRES  32 A  447  LYS LEU VAL ASN ALA SER VAL ILE GLU ASP GLY GLY HIS
SEQRES  33 A  447  PHE LEU ALA PHE GLU LEU PRO GLU ILE PHE ALA LYS ASP
SEQRES  34 A  447  VAL LEU LYS ALA ILE GLY GLU PHE ARG LYS LEU LYS ASN
SEQRES  35 A  447  VAL LYS THR GLU LEU
SEQRES   1 B  447  MET HIS HIS HIS HIS HIS HIS GLY VAL LEU LYS SER PRO
SEQRES   2 B  447  PRO PRO MET PRO LYS LEU ASP LEU GLU GLU TRP TRP GLY
SEQRES   3 B  447  PRO PRO GLU LEU LYS GLN LYS GLN ASP THR SER ILE LYS
SEQRES   4 B  447  PRO PHE GLU ILE THR PHE SER GLU THR MET VAL LYS GLU
SEQRES   5 B  447  LEU LYS GLU ARG ILE LYS LYS ARG ARG PRO PHE ALA PRO
SEQRES   6 B  447  PRO LEU GLU GLY VAL GLY PHE LYS TYR GLY PHE ASN SER
SEQRES   7 B  447  LYS GLN LEU ASP SER TRP LEU LYS TYR TRP ALA GLU GLU
SEQRES   8 B  447  TYR PRO PHE ALA GLU ARG GLN LYS PHE LEU ASN GLN TYR
SEQRES   9 B  447  PRO HIS PHE LYS THR ASN ILE GLN GLY LEU ASN ILE HIS
SEQRES  10 B  447  PHE MET ARG ILE THR PRO LYS VAL PRO LYS GLY VAL GLU
SEQRES  11 B  447  ILE VAL PRO LEU LEU LEU LEU HIS GLY TRP PRO GLY SER
SEQRES  12 B  447  VAL ARG GLU PHE TYR GLU ALA ILE PRO HIS LEU THR ALA
SEQRES  13 B  447  VAL SER LYS ASP ARG ASN PHE ALA LEU GLU ILE ILE ALA
SEQRES  14 B  447  PRO SER LEU PRO GLY TYR GLY PHE SER ASP ALA ALA VAL
SEQRES  15 B  447  ARG PRO GLY LEU ALA ALA ALA GLU VAL ALA VAL ILE PHE
SEQRES  16 B  447  LYS ASN LEU MET ALA ARG LEU GLY TYR LYS GLN TYR TYR
SEQRES  17 B  447  VAL GLN GLY GLY ASP TRP GLY ALA LEU ILE GLY SER ALA
SEQRES  18 B  447  MET ALA THR PHE PHE PRO LYS GLU ILE ILE GLY PHE HIS
SEQRES  19 B  447  SER ASN MET ALA LEU THR LEU SER PRO ALA ALA THR PHE
SEQRES  20 B  447  LEU GLU PHE VAL GLY ALA LEU PHE PRO SER LEU ILE VAL
SEQRES  21 B  447  GLU PRO GLU LEU ALA ASN ARG LEU TYR PRO LEU SER GLU
SEQRES  22 B  447  LYS TYR SER THR LEU LEU GLU GLU LEU GLY TYR MET HIS
SEQRES  23 B  447  ILE GLN ALA THR LYS PRO ASP THR VAL GLY ILE GLY LEU
SEQRES  24 B  447  THR ASP SER PRO ALA GLY LEU LEU ALA TYR ILE LEU GLU
SEQRES  25 B  447  LYS PHE SER THR TRP THR ASN PRO ASP LEU ARG SER LYS
SEQRES  26 B  447  GLU ASP GLY GLY LEU SER TYR ARG TRP THR LYS ASP GLN
SEQRES  27 B  447  LEU ILE ASP ASN LEU MET LEU TYR TRP SER THR LYS SER
SEQRES  28 B  447  ILE VAL THR SER MET ARG LEU TYR ALA GLU SER PHE SER
SEQRES  29 B  447  SER ARG HIS PHE ASP LEU LYS LEU ASP GLU ILE GLN VAL
SEQRES  30 B  447  GLN VAL PRO THR TRP VAL LEU GLN ALA LYS HIS GLU LEU
SEQRES  31 B  447  ALA TYR GLN PRO PRO CYS ILE LEU LYS MET LYS TYR PRO
SEQRES  32 B  447  LYS LEU VAL ASN ALA SER VAL ILE GLU ASP GLY GLY HIS
SEQRES  33 B  447  PHE LEU ALA PHE GLU LEU PRO GLU ILE PHE ALA LYS ASP
SEQRES  34 B  447  VAL LEU LYS ALA ILE GLY GLU PHE ARG LYS LEU LYS ASN
SEQRES  35 B  447  VAL LYS THR GLU LEU
HET    1PE  A 501      16
HET    1PE  B 501      16
HETNAM     1PE PENTAETHYLENE GLYCOL
HETSYN     1PE PEG400
FORMUL   3  1PE    2(C10 H22 O6)
FORMUL   5  HOH   *159(H2 O)
HELIX    1   1 PRO A   41  LYS A   47  5                                   7
HELIX    2   2 SER A   60  LYS A   73  1                                  14
HELIX    3   3 ASN A   91  LYS A   93  5                                   3
HELIX    4   4 GLN A   94  GLU A  105  1                                  12
HELIX    5   5 PRO A  107  ASN A  116  1                                  10
HELIX    6   6 SER A  157  GLU A  163  5                                   7
HELIX    7   7 ALA A  164  THR A  169  1                                   6
HELIX    8   8 ALA A  201  GLY A  217  1                                  17
HELIX    9   9 ASP A  227  PHE A  240  1                                  14
HELIX   10  10 SER A  256  PHE A  269  1                                  14
HELIX   11  11 PRO A  270  ILE A  273  5                                   4
HELIX   12  12 LEU A  278  LEU A  282  5                                   5
HELIX   13  13 PRO A  284  LEU A  296  1                                  13
HELIX   14  14 LEU A  296  LYS A  305  1                                  10
HELIX   15  15 LYS A  305  SER A  316  1                                  12
HELIX   16  16 SER A  316  TRP A  331  1                                  16
HELIX   17  17 ASN A  333  LYS A  339  5                                   7
HELIX   18  18 THR A  349  SER A  362  1                                  14
HELIX   19  19 SER A  365  PHE A  377  1                                  13
HELIX   20  20 SER A  378  ASP A  383  1                                   6
HELIX   21  21 PRO A  408  TYR A  416  1                                   9
HELIX   22  22 PHE A  431  LEU A  436  1                                   6
HELIX   23  23 LEU A  436  LYS A  455  1                                  20
HELIX   24  24 PRO B   41  LYS B   45  5                                   5
HELIX   25  25 SER B   60  LYS B   73  1                                  14
HELIX   26  26 ASN B   91  LYS B   93  5                                   3
HELIX   27  27 GLN B   94  GLU B  105  1                                  12
HELIX   28  28 PRO B  107  ASN B  116  1                                  10
HELIX   29  29 SER B  157  TYR B  162  5                                   6
HELIX   30  30 ALA B  164  THR B  169  1                                   6
HELIX   31  31 ALA B  201  LEU B  216  1                                  16
HELIX   32  32 ASP B  227  PHE B  240  1                                  14
HELIX   33  33 LEU B  292  GLU B  295  5                                   4
HELIX   34  34 LEU B  296  ALA B  303  1                                   8
HELIX   35  35 LYS B  305  SER B  316  1                                  12
HELIX   36  36 SER B  316  TRP B  331  1                                  16
HELIX   37  37 ASN B  333  LYS B  339  5                                   7
HELIX   38  38 THR B  349  SER B  362  1                                  14
HELIX   39  39 SER B  365  PHE B  377  1                                  13
HELIX   40  40 PRO B  408  TYR B  416  1                                   9
HELIX   41  41 PHE B  431  LEU B  436  1                                   6
HELIX   42  42 LEU B  436  LYS B  455  1                                  20
SHEET    1   A 9 LYS A  53  PHE A  55  0
SHEET    2   A 9 HIS A 120  ILE A 125 -1  O  LYS A 122   N  LYS A  53
SHEET    3   A 9 LEU A 128  ILE A 135 -1  O  LEU A 128   N  ILE A 125
SHEET    4   A 9 PHE A 177  PRO A 184 -1  O  ALA A 183   N  MET A 133
SHEET    5   A 9 GLU A 144  LEU A 151  1  N  VAL A 146   O  ALA A 178
SHEET    6   A 9 TYR A 221  GLY A 225  1  O  TYR A 222   N  LEU A 149
SHEET    7   A 9 ILE A 244  SER A 249  1  O  HIS A 248   N  VAL A 223
SHEET    8   A 9 THR A 395  GLN A 399  1  O  LEU A 398   N  SER A 249
SHEET    9   A 9 LEU A 419  VAL A 424  1  O  VAL A 420   N  THR A 395
SHEET    1   B 9 LYS B  53  PHE B  55  0
SHEET    2   B 9 HIS B 120  ILE B 125 -1  O  HIS B 120   N  PHE B  55
SHEET    3   B 9 LEU B 128  ILE B 135 -1  O  LEU B 128   N  ILE B 125
SHEET    4   B 9 PHE B 177  PRO B 184 -1  O  ALA B 183   N  MET B 133
SHEET    5   B 9 GLU B 144  LEU B 151  1  N  LEU B 148   O  GLU B 180
SHEET    6   B 9 TYR B 221  GLY B 225  1  O  TYR B 222   N  LEU B 149
SHEET    7   B 9 ILE B 244  SER B 249  1  O  HIS B 248   N  VAL B 223
SHEET    8   B 9 THR B 395  ALA B 400  1  O  TRP B 396   N  SER B 249
SHEET    9   B 9 LEU B 419  ILE B 425  1  O  VAL B 420   N  THR B 395
CISPEP   1 TRP A  154    PRO A  155          0         1.47
CISPEP   2 TYR A  283    PRO A  284          0        -0.76
CISPEP   3 GLY A  342    GLY A  343          0       -16.80
CISPEP   4 TRP B  154    PRO B  155          0         4.46
CISPEP   5 GLY B  342    GLY B  343          0       -20.93
SITE     1 AC1  3 HIS A 300  PHE A 377  SER A 379
SITE     1 AC2  3 THR B 304  SER B 378  SER B 379
CRYST1  161.404   51.466  124.449  90.00 122.68  90.00 C 1 2 1       8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.006196  0.000000  0.003974        0.00000
SCALE2      0.000000  0.019430  0.000000        0.00000
SCALE3      0.000000  0.000000  0.009547        0.00000
TER    3420      ASN A 456
TER    6493      ASN B 456
MASTER      381    0    2   42   18    0    2    6 6682    2   32   70
END