longtext: 4R1D-pdb

content
HEADER    HYDROLASE/HYDROLASE INHIBITOR           05-AUG-14   4R1D
TITLE     THE CRYSTAL STRUCTURE OF TLE4-TLI4 COMPLEX
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: UNCHARACTERIZED PROTEIN;
COMPND   3 CHAIN: A;
COMPND   4 ENGINEERED: YES;
COMPND   5 MOL_ID: 2;
COMPND   6 MOLECULE: UNCHARACTERIZED PROTEIN;
COMPND   7 CHAIN: B;
COMPND   8 FRAGMENT: UNP RESIDUES 33-380;
COMPND   9 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: PSEUDOMONAS AERUGINOSA PAO1;
SOURCE   3 ORGANISM_TAXID: 208964;
SOURCE   4 STRAIN: ATCC 15692 / PAO1 / 1C / PRS 101 / LMG 12228;
SOURCE   5 GENE: PA1510;
SOURCE   6 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 562;
SOURCE   8 MOL_ID: 2;
SOURCE   9 ORGANISM_SCIENTIFIC: PSEUDOMONAS AERUGINOSA PAO1;
SOURCE  10 ORGANISM_TAXID: 208964;
SOURCE  11 STRAIN: ATCC 15692 / PAO1 / 1C / PRS 101 / LMG 12228;
SOURCE  12 GENE: PA1509;
SOURCE  13 EXPRESSION_SYSTEM: ESCHERICHIA COLI;
SOURCE  14 EXPRESSION_SYSTEM_TAXID: 562
KEYWDS    LIPASE, HYDROLASE-HYDROLASE INHIBITOR COMPLEX
EXPDTA    X-RAY DIFFRACTION
AUTHOR    D.LU
REVDAT   1   17-DEC-14 4R1D    0
JRNL        AUTH   D.LU,Y.ZHENG,N.LIAO,L.WEI,B.XU,X.LIU,J.LIU
JRNL        TITL   THE STRUCTURAL BASIS OF THE TLE4-TLI4 COMPLEX REVEALS THE
JRNL        TITL 2 SELF-PROTECTION MECHANISM OF H2-T6SS IN PSEUDOMONAS
JRNL        TITL 3 AERUGINOSA.
JRNL        REF    ACTA CRYSTALLOGR.,SECT.D      V.  70  3233 2014
JRNL        REFN                   ISSN 0907-4449
JRNL        PMID   25478841
JRNL        DOI    10.1107/S1399004714023967
REMARK   2
REMARK   2 RESOLUTION.    1.75 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE: 1.6.4_486)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-
REMARK   3               : KUNSTLEVE,LI-WEI HUNG,ROBERT IMMORMINO,
REMARK   3               : TOM IOERGER,AIRLIE MCCOY,ERIK MCKEE,NIGEL
REMARK   3               : MORIARTY,REETAL PAI,RANDY READ,JANE
REMARK   3               : RICHARDSON,DAVID RICHARDSON,TOD ROMO,JIM
REMARK   3               : SACCHETTINI,NICHOLAS SAUTER,JACOB SMITH,
REMARK   3               : LAURENT STORONI,TOM TERWILLIGER,PETER
REMARK   3               : ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 1.75
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 20.00
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 94.4
REMARK   3   NUMBER OF REFLECTIONS             : 86321
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.177
REMARK   3   R VALUE            (WORKING SET) : 0.175
REMARK   3   FREE R VALUE                     : 0.214
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.020
REMARK   3   FREE R VALUE TEST SET COUNT      : 4331
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 20.0043 -  5.4003    0.99     2899   161  0.1726 0.2001
REMARK   3     2  5.4003 -  4.3004    0.99     2884   159  0.1438 0.1677
REMARK   3     3  4.3004 -  3.7609    0.96     2766   147  0.1511 0.1790
REMARK   3     4  3.7609 -  3.4189    0.96     2778   158  0.1686 0.2107
REMARK   3     5  3.4189 -  3.1749    0.98     2836   131  0.1790 0.2130
REMARK   3     6  3.1749 -  2.9883    0.98     2854   136  0.1906 0.2206
REMARK   3     7  2.9883 -  2.8391    0.97     2793   156  0.1798 0.2307
REMARK   3     8  2.8391 -  2.7158    0.97     2839   155  0.1755 0.1959
REMARK   3     9  2.7158 -  2.6115    0.97     2774   153  0.1803 0.2184
REMARK   3    10  2.6115 -  2.5216    0.97     2816   181  0.1809 0.2399
REMARK   3    11  2.5216 -  2.4429    0.97     2828   134  0.1787 0.1909
REMARK   3    12  2.4429 -  2.3732    0.97     2803   135  0.1701 0.2236
REMARK   3    13  2.3732 -  2.3108    0.97     2779   119  0.1649 0.2002
REMARK   3    14  2.3108 -  2.2545    0.95     2789   153  0.1680 0.2128
REMARK   3    15  2.2545 -  2.2033    0.95     2760   144  0.1691 0.2153
REMARK   3    16  2.2033 -  2.1565    0.96     2752   146  0.1758 0.2372
REMARK   3    17  2.1565 -  2.1134    0.94     2750   135  0.1838 0.2455
REMARK   3    18  2.1134 -  2.0735    0.95     2754   130  0.1842 0.2659
REMARK   3    19  2.0735 -  2.0365    0.93     2698   162  0.1859 0.2441
REMARK   3    20  2.0365 -  2.0020    0.93     2654   149  0.1928 0.2435
REMARK   3    21  2.0020 -  1.9698    0.93     2661   154  0.1925 0.2456
REMARK   3    22  1.9698 -  1.9395    0.92     2664   145  0.1868 0.2565
REMARK   3    23  1.9395 -  1.9110    0.92     2651   123  0.1963 0.2517
REMARK   3    24  1.9110 -  1.8841    0.91     2633   148  0.1964 0.2244
REMARK   3    25  1.8841 -  1.8587    0.91     2620   134  0.1996 0.2355
REMARK   3    26  1.8587 -  1.8345    0.92     2691   136  0.2112 0.2803
REMARK   3    27  1.8345 -  1.8116    0.90     2550   153  0.2091 0.2457
REMARK   3    28  1.8116 -  1.7898    0.90     2634   131  0.2156 0.2377
REMARK   3    29  1.7898 -  1.7690    0.88     2544   127  0.2171 0.2644
REMARK   3    30  1.7690 -  1.7491    0.88     2536   136  0.2385 0.2994
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.20
REMARK   3   SHRINKAGE RADIUS   : 0.95
REMARK   3   K_SOL              : 0.33
REMARK   3   B_SOL              : 37.93
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.210
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 21.750
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : -0.43250
REMARK   3    B22 (A**2) : -0.19640
REMARK   3    B33 (A**2) : 0.62890
REMARK   3    B12 (A**2) : -0.00000
REMARK   3    B13 (A**2) : -0.21010
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           6707
REMARK   3   ANGLE     :  1.065           9120
REMARK   3   CHIRALITY :  0.076            947
REMARK   3   PLANARITY :  0.005           1209
REMARK   3   DIHEDRAL  : 14.103           2505
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4R1D COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBJ ON 08-AUG-14.
REMARK 100 THE RCSB ID CODE IS RCSB086776.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 14-SEP-13
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : SSRF
REMARK 200  BEAMLINE                       : BL17U
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9790
REMARK 200  MONOCHROMATOR                  : SI 111 CHANNEL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 315R
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : HKL-2000
REMARK 200  DATA SCALING SOFTWARE          : HKL-2000
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 86321
REMARK 200  RESOLUTION RANGE HIGH      (A) : 1.749
REMARK 200  RESOLUTION RANGE LOW       (A) : 50.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : 2.000
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 94.4
REMARK 200  DATA REDUNDANCY                : NULL
REMARK 200  R MERGE                    (I) : NULL
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : NULL
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 1.75
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 1.81
REMARK 200  COMPLETENESS FOR SHELL     (%) : 89.3
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SAD
REMARK 200 SOFTWARE USED: SOLVE
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 45.62
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.26
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 20% PEG 3350, 0.1M BIS-TRIS PH 6.5,
REMARK 280  VAPOR DIFFUSION, HANGING DROP, TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       67.58950
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 5560 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 31250 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -33.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A     1
REMARK 465     SER A     2
REMARK 465     SER A     3
REMARK 465     GLU A     4
REMARK 465     PRO A     5
REMARK 465     LEU A     6
REMARK 465     GLU A     7
REMARK 465     PRO A     8
REMARK 465     ASN A     9
REMARK 465     GLN A    10
REMARK 465     ASP A    11
REMARK 465     VAL A    12
REMARK 465     ILE A    13
REMARK 465     ILE A    14
REMARK 465     PRO A    15
REMARK 465     ARG A    16
REMARK 465     SER A    17
REMARK 465     ARG A    18
REMARK 465     ASP A    19
REMARK 465     SER A    20
REMARK 465     LEU A    21
REMARK 465     GLY A    22
REMARK 465     ARG A    23
REMARK 465     PRO A    24
REMARK 465     VAL A    25
REMARK 465     TYR A    26
REMARK 465     LYS A    27
REMARK 465     ALA A    28
REMARK 465     GLN A    29
REMARK 465     LEU A    30
REMARK 465     THR A    31
REMARK 465     ARG A    32
REMARK 465     THR A    33
REMARK 465     ASP A    34
REMARK 465     ASN A    35
REMARK 465     GLN A    36
REMARK 465     SER A    37
REMARK 465     GLU A    38
REMARK 465     LYS A    39
REMARK 465     VAL A    40
REMARK 465     ALA A    41
REMARK 465     LEU A    42
REMARK 465     ILE A    43
REMARK 465     ARG A    44
REMARK 465     HIS A   562
REMARK 465     PRO A   563
REMARK 465     ASN A   564
REMARK 465     ASP A   565
REMARK 465     LYS A   566
REMARK 465     GLY A   567
REMARK 465     GLY A   568
REMARK 465     THR A   569
REMARK 465     SER B   334
REMARK 465     SER B   335
REMARK 465     PRO B   336
REMARK 465     ARG B   337
REMARK 465     GLY B   338
REMARK 465     GLY B   339
REMARK 465     ASN B   340
REMARK 465     ALA B   341
REMARK 465     GLY B   342
REMARK 465     PRO B   343
REMARK 465     SER B   344
REMARK 465     PRO B   345
REMARK 465     ALA B   346
REMARK 465     PRO B   347
REMARK 465     LYS B   348
REMARK 465     PRO B   349
REMARK 465     ALA B   350
REMARK 465     THR B   351
REMARK 465     PRO B   352
REMARK 465     GLY B   353
REMARK 465     GLY B   354
REMARK 465     GLN B   355
REMARK 465     THR B   356
REMARK 465     LEU B   357
REMARK 465     GLY B   358
REMARK 465     ASP B   359
REMARK 465     HIS B   360
REMARK 465     TYR B   361
REMARK 465     VAL B   362
REMARK 465     TYR B   363
REMARK 465     GLU B   364
REMARK 465     GLU B   365
REMARK 465     PHE B   366
REMARK 465     LEU B   367
REMARK 465     SER B   368
REMARK 465     SER B   369
REMARK 465     LEU B   370
REMARK 465     LYS B   371
REMARK 465     PRO B   372
REMARK 465     LYS B   373
REMARK 465     ASP B   374
REMARK 465     SER B   375
REMARK 465     TRP B   376
REMARK 465     LEU B   377
REMARK 465     ASP B   378
REMARK 465     ASP B   379
REMARK 465     LEU B   380
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    ARG A  73       80.65   -154.06
REMARK 500    MET A  81       71.31    -67.12
REMARK 500    TRP A  94      -58.55   -136.79
REMARK 500    LEU A 106       60.55   -100.46
REMARK 500    TRP A 167      -53.21     77.57
REMARK 500    ASP A 170      -75.45   -150.35
REMARK 500    SER A 256     -133.54     57.77
REMARK 500    ILE A 369      -56.38   -125.26
REMARK 500    LYS A 393      -34.26    -38.23
REMARK 500    GLU A 426       67.35   -117.64
REMARK 500    GLU A 498      -36.89     97.43
REMARK 500    LYS B  35       30.16    -95.94
REMARK 500    GLU B 109        9.47     57.64
REMARK 500    ASP B 189       23.82   -140.32
REMARK 500    ASN B 302       65.59     38.16
REMARK 500
REMARK 500 REMARK: NULL
REMARK 525
REMARK 525 SOLVENT
REMARK 525
REMARK 525 THE SOLVENT MOLECULES HAVE CHAIN IDENTIFIERS THAT
REMARK 525 INDICATE THE POLYMER CHAIN WITH WHICH THEY ARE MOST
REMARK 525 CLOSELY ASSOCIATED. THE REMARK LISTS ALL THE SOLVENT
REMARK 525 MOLECULES WHICH ARE MORE THAN 5A AWAY FROM THE
REMARK 525 NEAREST POLYMER CHAIN (M = MODEL NUMBER;
REMARK 525 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE
REMARK 525 NUMBER; I=INSERTION CODE):
REMARK 525
REMARK 525  M RES CSSEQI
REMARK 525    HOH A1120        DISTANCE =  6.28 ANGSTROMS
REMARK 525    HOH A1275        DISTANCE =  6.77 ANGSTROMS
REMARK 620
REMARK 620 METAL COORDINATION
REMARK 620 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 620 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 620
REMARK 620 COORDINATION ANGLES FOR:  M RES CSSEQI METAL
REMARK 620                              CA A 601  CA
REMARK 620 N RES CSSEQI ATOM
REMARK 620 1 PHE A 202   O
REMARK 620 2 GLU A 152   OE2 104.2
REMARK 620 3 ASP A 203   OD1  74.0  84.1
REMARK 620 4 ASP A 156   OD1  78.0 125.9 143.5
REMARK 620 5 ASP A 156   OD2 124.9  91.6 161.1  51.3
REMARK 620 6 HOH A 796   O   153.8  74.0  79.8 124.8  81.2
REMARK 620 7 HOH A 811   O    93.6 151.6  79.7  79.1  96.2  80.2
REMARK 620 N                    1     2     3     4     5     6
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE CA A 601
DBREF  4R1D A    1   569  UNP    Q9I3K2   Q9I3K2_PSEAE     1    569
DBREF  4R1D B   33   380  UNP    Q9I3K3   Q9I3K3_PSEAE    33    380
SEQRES   1 A  569  MET SER SER GLU PRO LEU GLU PRO ASN GLN ASP VAL ILE
SEQRES   2 A  569  ILE PRO ARG SER ARG ASP SER LEU GLY ARG PRO VAL TYR
SEQRES   3 A  569  LYS ALA GLN LEU THR ARG THR ASP ASN GLN SER GLU LYS
SEQRES   4 A  569  VAL ALA LEU ILE ARG GLN THR ALA PRO LEU PRO VAL ILE
SEQRES   5 A  569  PHE ILE PRO GLY ILE MET GLY THR ASN LEU ARG ASN LYS
SEQRES   6 A  569  ALA ASP LYS SER GLU VAL TRP ARG PRO PRO ASN GLY LEU
SEQRES   7 A  569  TRP PRO MET ASP ASP LEU PHE ALA SER ILE GLY ALA LEU
SEQRES   8 A  569  TRP THR TRP ALA TRP ARG GLY PRO LYS ALA ARG GLN GLU
SEQRES   9 A  569  LEU LEU LYS ALA GLU GLN VAL GLU VAL ASP ASP GLN GLY
SEQRES  10 A  569  THR ILE ASP VAL GLY GLN SER GLY LEU SER GLU GLU ALA
SEQRES  11 A  569  ALA ARG LEU ARG GLY TRP GLY LYS VAL MET ARG SER ALA
SEQRES  12 A  569  TYR ASN PRO VAL MET GLY LEU MET GLU ARG ARG LEU ASP
SEQRES  13 A  569  ASN ILE VAL SER ARG ARG GLU LEU GLN ALA TRP TRP ASN
SEQRES  14 A  569  ASP GLU ALA LEU SER PRO PRO GLY ASP GLN GLY GLU GLU
SEQRES  15 A  569  GLN GLY LYS VAL GLY PRO ILE ASP GLU GLU GLU LEU LEU
SEQRES  16 A  569  ARG ALA SER ARG TYR GLN PHE ASP VAL TRP CYS ALA GLY
SEQRES  17 A  569  TYR ASN TRP LEU GLN SER ASN ARG GLN SER ALA LEU ASP
SEQRES  18 A  569  VAL ARG ASP TYR ILE GLU ASN THR VAL LEU PRO PHE TYR
SEQRES  19 A  569  GLN LYS GLU CYS GLY LEU ASP PRO GLU GLN MET ARG ARG
SEQRES  20 A  569  MET LYS VAL ILE LEU VAL THR HIS SER MET GLY GLY LEU
SEQRES  21 A  569  VAL ALA ARG ALA LEU THR GLN LEU HIS GLY TYR GLU ARG
SEQRES  22 A  569  VAL LEU GLY VAL VAL HIS GLY VAL GLN PRO ALA THR GLY
SEQRES  23 A  569  SER SER THR ILE TYR HIS HIS MET ARG CYS GLY TYR GLU
SEQRES  24 A  569  GLY ILE ALA GLN VAL VAL LEU GLY ARG ASN ALA GLY GLU
SEQRES  25 A  569  VAL THR ALA ILE VAL ALA ASN SER ALA GLY ALA LEU GLU
SEQRES  26 A  569  LEU ALA PRO SER ALA GLU TYR ARG GLU GLY ARG PRO TRP
SEQRES  27 A  569  LEU PHE LEU CYS ASP ALA GLN GLY GLN VAL LEU LYS ASP
SEQRES  28 A  569  ILE ASP GLY LYS PRO ARG ALA TYR PRO GLN ASN GLN ASP
SEQRES  29 A  569  PRO TYR GLU GLU ILE TYR LYS ASN THR THR TRP TYR GLY
SEQRES  30 A  569  LEU VAL PRO GLU GLN ASN SER GLN TYR LEU ASP MET SER
SEQRES  31 A  569  ASP LYS LYS GLU GLY LEU ARG VAL GLY PRO ARG ASP ASN
SEQRES  32 A  569  PHE GLU ASP LEU ILE ASP SER ILE ALA ASN PHE HIS GLY
SEQRES  33 A  569  GLU LEU SER ALA ALA GLY TYR HIS SER GLU THR TYR ALA
SEQRES  34 A  569  HIS TYR GLY ALA ASP ASP SER ARG HIS SER TRP ARG ASP
SEQRES  35 A  569  LEU ILE TRP LYS GLY ASP PRO THR PRO LEU GLU THR PRO
SEQRES  36 A  569  GLY ALA THR LEU ASN ASP ASP GLU ASN GLY THR TYR ASN
SEQRES  37 A  569  SER TRP PHE ARG ARG GLY LEU PRO THR ILE VAL GLN GLY
SEQRES  38 A  569  PRO LEU GLU THR GLY ASN PRO LEU ASP ALA SER GLY SER
SEQRES  39 A  569  GLY GLY ASP GLU THR VAL PRO THR ASP SER GLY GLN ALA
SEQRES  40 A  569  PRO ALA LEU ALA GLY VAL LYS ALA SER PHE ARG HIS GLY
SEQRES  41 A  569  SER LYS GLY LYS GLY GLN ALA ASN THR LYS ARG GLY TYR
SEQRES  42 A  569  GLU HIS GLN GLU SER TYR ASN ASP ALA ARG ALA GLN TRP
SEQRES  43 A  569  ALA ALA LEU TYR GLY VAL ILE LYS ILE THR GLN LEU ALA
SEQRES  44 A  569  ASP TRP HIS PRO ASN ASP LYS GLY GLY THR
SEQRES   1 B  348  MET ASP LYS THR GLY TRP ILE THR HIS CYS PHE GLY ARG
SEQRES   2 B  348  PHE LEU ILE ASP LEU PRO PRO ASP ALA VAL ILE ASN ALA
SEQRES   3 B  348  GLY TYR TYR LEU TRP GLY ASP ARG ILE GLU TYR LEU ASP
SEQRES   4 B  348  ASP LYS PRO THR GLU LEU ALA ALA ARG VAL ASP ARG LEU
SEQRES   5 B  348  GLU GLN GLU TRP ARG THR GLN ARG HIS LYS SER LYS GLY
SEQRES   6 B  348  ASN MET PHE LEU ARG LYS ILE ASP PHE GLY ASN GLU SER
SEQRES   7 B  348  VAL GLY LEU LEU SER TRP SER SER GLU VAL ALA SER LYS
SEQRES   8 B  348  THR TYR LEU LEU ASP THR TYR VAL THR SER LYS PRO THR
SEQRES   9 B  348  TRP HIS VAL TYR ARG TRP LYS GLY LYS VAL SER VAL ASP
SEQRES  10 B  348  ARG GLU GLN HIS ALA VAL GLU ILE SER ARG ALA LEU ALA
SEQRES  11 B  348  ARG ASN LEU ARG SER ARG ALA PRO LYS GLU ILE PRO SER
SEQRES  12 B  348  GLU PRO GLY PHE CYS ILE ASP HIS ALA TYR ILE ALA GLY
SEQRES  13 B  348  ASP SER PHE GLN VAL GLU ARG PHE GLY VAL GLY VAL THR
SEQRES  14 B  348  PHE PRO GLU HIS PRO GLY ALA ARG PHE GLU PHE ARG SER
SEQRES  15 B  348  SER THR GLY ALA GLU LEU ASN SER LEU LEU GLU ARG VAL
SEQRES  16 B  348  ASP GLY PHE VAL GLN ASN MET LEU SER THR PHE ALA GLY
SEQRES  17 B  348  MET GLU THR LEU ARG LYS GLY LYS HIS PRO VAL GLY SER
SEQRES  18 B  348  LEU PRO GLY GLU GLU TYR LEU VAL ALA GLY SER ASP LYS
SEQRES  19 B  348  GLY GLN ARG GLY TYR THR PHE MET TRP GLU VAL GLN GLY
SEQRES  20 B  348  LYS GLU GLU SER LEU THR GLU PRO ASN LEU THR ALA GLY
SEQRES  21 B  348  LEU ALA VAL LEU GLU ARG SER ASN GLU ASN GLY LYS PRO
SEQRES  22 B  348  PRO PRO PRO ALA PHE LYS SER ASP LYS GLU ALA LEU GLU
SEQRES  23 B  348  LEU TRP ASP THR ILE VAL ASP SER ILE ARG VAL ARG PRO
SEQRES  24 B  348  THR SER SER SER PRO ARG GLY GLY ASN ALA GLY PRO SER
SEQRES  25 B  348  PRO ALA PRO LYS PRO ALA THR PRO GLY GLY GLN THR LEU
SEQRES  26 B  348  GLY ASP HIS TYR VAL TYR GLU GLU PHE LEU SER SER LEU
SEQRES  27 B  348  LYS PRO LYS ASP SER TRP LEU ASP ASP LEU
HET     CA  A 601       1
HETNAM      CA CALCIUM ION
FORMUL   3   CA    CA 2+
FORMUL   4  HOH   *960(H2 O)
HELIX    1   1 ASP A   83  TRP A   94  1                                  12
HELIX    2   2 GLY A   98  LEU A  106  1                                   9
HELIX    3   3 LYS A  107  GLU A  109  5                                   3
HELIX    4   4 SER A  127  ARG A  134  1                                   8
HELIX    5   5 GLY A  135  VAL A  139  5                                   5
HELIX    6   6 MET A  140  ASP A  156  1                                  17
HELIX    7   7 ASP A  170  SER A  174  5                                   5
HELIX    8   8 PRO A  175  GLY A  180  5                                   6
HELIX    9   9 ASP A  190  SER A  198  1                                   9
HELIX   10  10 SER A  214  THR A  229  1                                  16
HELIX   11  11 THR A  229  GLY A  239  1                                  11
HELIX   12  12 ASP A  241  MET A  248  1                                   8
HELIX   13  13 MET A  257  LEU A  268  1                                  12
HELIX   14  14 SER A  288  GLY A  297  1                                  10
HELIX   15  15 ASN A  309  ALA A  318  1                                  10
HELIX   16  16 SER A  320  GLU A  325  1                                   6
HELIX   17  17 GLU A  331  ARG A  336  5                                   6
HELIX   18  18 ASP A  364  ILE A  369  1                                   6
HELIX   19  19 PRO A  380  LEU A  387  5                                   8
HELIX   20  20 LYS A  392  LEU A  396  5                                   5
HELIX   21  21 GLY A  399  GLY A  422  1                                  24
HELIX   22  22 PRO A  449  THR A  454  1                                   6
HELIX   23  23 ASN A  487  SER A  494  5                                   8
HELIX   24  24 PRO A  501  GLN A  506  1                                   6
HELIX   25  25 GLN A  506  ALA A  511  1                                   6
HELIX   26  26 GLU A  537  ASN A  540  5                                   4
HELIX   27  27 ASP A  541  GLN A  557  1                                  17
HELIX   28  28 LYS B   73  GLN B   91  1                                  19
HELIX   29  29 ARG B  150  ASN B  164  1                                  15
HELIX   30  30 SER B  222  PHE B  238  1                                  17
HELIX   31  31 SER B  312  ASP B  325  1                                  14
SHEET    1   A 6 GLN A 201  CYS A 206  0
SHEET    2   A 6 PRO A  48  ILE A  54  1  N  PHE A  53   O  TRP A 205
SHEET    3   A 6 VAL A 250  HIS A 255  1  O  VAL A 253   N  ILE A  54
SHEET    4   A 6 VAL A 274  GLY A 280  1  O  LEU A 275   N  VAL A 250
SHEET    5   A 6 THR A 427  GLY A 432  1  O  TYR A 428   N  VAL A 277
SHEET    6   A 6 ALA A 515  HIS A 519  1  O  ALA A 515   N  ALA A 429
SHEET    1   B 3 GLU A  70  TRP A  72  0
SHEET    2   B 3 LEU A  62  ASN A  64 -1  N  LEU A  62   O  VAL A  71
SHEET    3   B 3 VAL A 111  VAL A 113 -1  O  GLU A 112   N  ARG A  63
SHEET    1   C 2 VAL A 159  SER A 160  0
SHEET    2   C 2 GLU A 163  LEU A 164 -1  O  GLU A 163   N  SER A 160
SHEET    1   D 4 ALA A 358  TYR A 359  0
SHEET    2   D 4 LEU A 339  CYS A 342 -1  N  LEU A 339   O  TYR A 359
SHEET    3   D 4 LEU A 443  LYS A 446  1  O  LEU A 443   N  PHE A 340
SHEET    4   D 4 THR A 477  VAL A 479 -1  O  THR A 477   N  LYS A 446
SHEET    1   E 3 ILE B  39  PHE B  43  0
SHEET    2   E 3 PHE B  46  LEU B  50 -1  O  LEU B  50   N  ILE B  39
SHEET    3   E 3 ARG B 328  VAL B 329 -1  O  ARG B 328   N  LEU B  47
SHEET    1   F12 PHE B 100  ASP B 105  0
SHEET    2   F12 VAL B 111  TRP B 116 -1  O  LEU B 114   N  ARG B 102
SHEET    3   F12 THR B 124  THR B 132 -1  O  TYR B 130   N  VAL B 111
SHEET    4   F12 VAL B 139  SER B 147 -1  O  VAL B 146   N  TYR B 125
SHEET    5   F12 ASP B  65  LEU B  70 -1  N  GLU B  68   O  ARG B 141
SHEET    6   F12 VAL B  55  LEU B  62 -1  N  LEU B  62   O  ASP B  65
SHEET    7   F12 GLU B 194  THR B 201 -1  O  ARG B 195   N  TYR B  61
SHEET    8   F12 ALA B 208  SER B 215 -1  O  PHE B 210   N  VAL B 200
SHEET    9   F12 ASN B 288  VAL B 295 -1  O  GLY B 292   N  GLU B 211
SHEET   10   F12 GLN B 268  VAL B 277 -1  N  PHE B 273   O  LEU B 293
SHEET   11   F12 GLU B 257  ASP B 265 -1  N  GLY B 263   O  GLY B 270
SHEET   12   F12 MET B 241  LYS B 248 -1  N  ARG B 245   O  LEU B 260
SHEET    1   G 3 LEU B 165  SER B 167  0
SHEET    2   G 3 ALA B 184  ALA B 187 -1  O  TYR B 185   N  ARG B 166
SHEET    3   G 3 GLY B 178  ILE B 181 -1  N  PHE B 179   O  ILE B 186
LINK         O   PHE A 202                CA    CA A 601     1555   1555  2.39
LINK         OE2 GLU A 152                CA    CA A 601     1555   1555  2.42
LINK         OD1 ASP A 203                CA    CA A 601     1555   1555  2.47
LINK         OD1 ASP A 156                CA    CA A 601     1555   1555  2.52
LINK         OD2 ASP A 156                CA    CA A 601     1555   1555  2.59
LINK        CA    CA A 601                 O   HOH A 796     1555   1555  2.57
LINK        CA    CA A 601                 O   HOH A 811     1555   1555  2.64
CISPEP   1 GLN A   45    THR A   46          0         4.49
CISPEP   2 PRO A   74    PRO A   75          0         2.89
CISPEP   3 TRP A   79    PRO A   80          0        -8.82
CISPEP   4 TYR A  359    PRO A  360          0        -3.58
CISPEP   5 GLY A  481    PRO A  482          0         4.26
CISPEP   6 LYS B  134    PRO B  135          0        -0.41
SITE     1 AC1  6 GLU A 152  ASP A 156  PHE A 202  ASP A 203
SITE     2 AC1  6 HOH A 796  HOH A 811
CRYST1   59.681  135.179   62.606  90.00 113.49  90.00 P 1 21 1      2
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016756  0.000000  0.007283        0.00000
SCALE2      0.000000  0.007398  0.000000        0.00000
SCALE3      0.000000  0.000000  0.017416        0.00000
TER    4112      TRP A 561
TER    6532      SER B 333
MASTER      387    0    1   31   33    0    2    6 7422    2    9   71
END