longtext: 4TPK-pdb

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HEADER    HYDROLASE                               07-JUN-14   4TPK
TITLE     HUMAN BUTYRYLCHOLINESTERASE IN COMPLEX WITH N-((1-(2,3-DIHYDRO-1H-
TITLE    2 INDEN-2-YL)PIPERIDIN-3-YL)METHYL)-N-(2-METHOXYETHYL)-2-NAPHTHAMIDE
CAVEAT     4TPK    NAG A 608 HAS WRONG CHIRALITY AT C1 NAG A 607 HAS WRONG
CAVEAT   2 4TPK    CHIRALITY AT C1 NAG B 610 HAS WRONG CHIRALITY AT C1
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: CHOLINESTERASE;
COMPND   3 CHAIN: A, B;
COMPND   4 SYNONYM: ACYLCHOLINE ACYLHYDROLASE,BUTYRYLCHOLINE ESTERASE,CHOLINE
COMPND   5 ESTERASE II,PSEUDOCHOLINESTERASE;
COMPND   6 EC: 3.1.1.8;
COMPND   7 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: BCHE, CHE1;
SOURCE   6 EXPRESSION_SYSTEM: CRICETULUS GRISEUS;
SOURCE   7 EXPRESSION_SYSTEM_COMMON: CHINESE HAMSTER;
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 10029
KEYWDS    BUTYRYLCHOLINESTERASE INHIBITION, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    N.COQUELLE,B.BRUS,J.P.COLLETIER,S.GOBEC
REVDAT   1   22-OCT-14 4TPK    0
JRNL        AUTH   B.BRUS,U.KOSAK,S.TURK,A.PISLAR,N.COQUELLE,J.KOS,J.STOJAN,
JRNL        AUTH 2 J.P.COLLETIER,S.GOBEC
JRNL        TITL   DISCOVERY, BIOLOGICAL EVALUATION, AND CRYSTAL STRUCTURE OF A
JRNL        TITL 2 NOVEL NANOMOLAR SELECTIVE BUTYRYLCHOLINESTERASE INHIBITOR.
JRNL        REF    J.MED.CHEM.                   V.  57  8167 2014
JRNL        REFN                   ISSN 0022-2623
JRNL        PMID   25226236
JRNL        DOI    10.1021/JM501195E
REMARK   2
REMARK   2 RESOLUTION.    2.70 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : PHENIX (PHENIX.REFINE: 1.8.4_1496)
REMARK   3   AUTHORS     : PAUL ADAMS,PAVEL AFONINE,VINCENT CHEN,IAN
REMARK   3               : DAVIS,KRESHNA GOPAL,RALF GROSSE-KUNSTLEVE,
REMARK   3               : LI-WEI HUNG,ROBERT IMMORMINO,TOM IOERGER,
REMARK   3               : AIRLIE MCCOY,ERIK MCKEE,NIGEL MORIARTY,
REMARK   3               : REETAL PAI,RANDY READ,JANE RICHARDSON,
REMARK   3               : DAVID RICHARDSON,TOD ROMO,JIM SACCHETTINI,
REMARK   3               : NICHOLAS SAUTER,JACOB SMITH,LAURENT
REMARK   3               : STORONI,TOM TERWILLIGER,PETER ZWART
REMARK   3
REMARK   3    REFINEMENT TARGET : ML
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.70
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 46.20
REMARK   3   MIN(FOBS/SIGMA_FOBS)              : 1.360
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.8
REMARK   3   NUMBER OF REFLECTIONS             : 40022
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.186
REMARK   3   R VALUE            (WORKING SET) : 0.184
REMARK   3   FREE R VALUE                     : 0.239
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 3.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 1202
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT (IN BINS).
REMARK   3   BIN  RESOLUTION RANGE  COMPL.    NWORK NFREE   RWORK  RFREE
REMARK   3     1 46.2025 -  5.6133    0.99     4538   140  0.1924 0.2347
REMARK   3     2  5.6133 -  4.4566    1.00     4356   133  0.1560 0.1871
REMARK   3     3  4.4566 -  3.8936    1.00     4346   136  0.1490 0.2021
REMARK   3     4  3.8936 -  3.5377    1.00     4305   136  0.1747 0.2366
REMARK   3     5  3.5377 -  3.2842    1.00     4279   132  0.1990 0.2548
REMARK   3     6  3.2842 -  3.0907    1.00     4233   135  0.2189 0.3258
REMARK   3     7  3.0907 -  2.9359    1.00     4278   126  0.2226 0.3001
REMARK   3     8  2.9359 -  2.8081    1.00     4257   133  0.2354 0.3050
REMARK   3     9  2.8081 -  2.7000    1.00     4228   131  0.2319 0.3154
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED        : FLAT BULK SOLVENT MODEL
REMARK   3   SOLVENT RADIUS     : 1.11
REMARK   3   SHRINKAGE RADIUS   : 0.90
REMARK   3   K_SOL              : NULL
REMARK   3   B_SOL              : NULL
REMARK   3
REMARK   3  ERROR ESTIMATES.
REMARK   3   COORDINATE ERROR (MAXIMUM-LIKELIHOOD BASED)     : 0.370
REMARK   3   PHASE ERROR (DEGREES, MAXIMUM-LIKELIHOOD BASED) : 25.380
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : NULL
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : NULL
REMARK   3    B22 (A**2) : NULL
REMARK   3    B33 (A**2) : NULL
REMARK   3    B12 (A**2) : NULL
REMARK   3    B13 (A**2) : NULL
REMARK   3    B23 (A**2) : NULL
REMARK   3
REMARK   3  TWINNING INFORMATION.
REMARK   3   FRACTION: NULL
REMARK   3   OPERATOR: NULL
REMARK   3
REMARK   3  DEVIATIONS FROM IDEAL VALUES.
REMARK   3                 RMSD          COUNT
REMARK   3   BOND      :  0.007           9079
REMARK   3   ANGLE     :  1.021          12288
REMARK   3   CHIRALITY :  0.038           1328
REMARK   3   PLANARITY :  0.005           1546
REMARK   3   DIHEDRAL  : 14.508           3344
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : NULL
REMARK   3
REMARK   3  NCS DETAILS
REMARK   3   NUMBER OF NCS GROUPS : NULL
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: NULL
REMARK   4
REMARK   4 4TPK COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 10-JUN-14.
REMARK 100 THE DEPOSITION ID IS D_1000201991.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 28-JUL-13
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 7.4
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : ESRF
REMARK 200  BEAMLINE                       : ID23-2
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.873
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS PILATUS 2M
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : XSCALE
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 40037
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.700
REMARK 200  RESOLUTION RANGE LOW       (A) : 46.200
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.9
REMARK 200  DATA REDUNDANCY                : 5.200
REMARK 200  R MERGE                    (I) : 0.08829
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 13.6200
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.70
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.80
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.9
REMARK 200  DATA REDUNDANCY IN SHELL       : 5.20
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 2.390
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: MOLECULAR REPLACEMENT
REMARK 200 SOFTWARE USED: PHASER
REMARK 200 STARTING MODEL: 1POM
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 47.79
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.36
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 0.2M AMMONIUM ACETATE, 24% PEG 3350,
REMARK 280  10 MM TRIS PH 7.4
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 21
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       38.30000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000      115.82500
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       40.10000
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000      115.82500
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       38.30000
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       40.10000
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     MET A   -27
REMARK 465     HIS A   -26
REMARK 465     SER A   -25
REMARK 465     LYS A   -24
REMARK 465     VAL A   -23
REMARK 465     THR A   -22
REMARK 465     ILE A   -21
REMARK 465     ILE A   -20
REMARK 465     CYS A   -19
REMARK 465     ILE A   -18
REMARK 465     ARG A   -17
REMARK 465     PHE A   -16
REMARK 465     LEU A   -15
REMARK 465     PHE A   -14
REMARK 465     TRP A   -13
REMARK 465     PHE A   -12
REMARK 465     LEU A   -11
REMARK 465     LEU A   -10
REMARK 465     LEU A    -9
REMARK 465     CYS A    -8
REMARK 465     MET A    -7
REMARK 465     LEU A    -6
REMARK 465     ILE A    -5
REMARK 465     GLY A    -4
REMARK 465     LYS A    -3
REMARK 465     SER A    -2
REMARK 465     HIS A    -1
REMARK 465     THR A     0
REMARK 465     GLU A     1
REMARK 465     ASP A     2
REMARK 465     GLU A   531
REMARK 465     MET A   532
REMARK 465     THR A   533
REMARK 465     GLY A   534
REMARK 465     ASN A   535
REMARK 465     ILE A   536
REMARK 465     ASP A   537
REMARK 465     GLU A   538
REMARK 465     ALA A   539
REMARK 465     GLU A   540
REMARK 465     TRP A   541
REMARK 465     GLU A   542
REMARK 465     TRP A   543
REMARK 465     LYS A   544
REMARK 465     ALA A   545
REMARK 465     GLY A   546
REMARK 465     PHE A   547
REMARK 465     HIS A   548
REMARK 465     ARG A   549
REMARK 465     TRP A   550
REMARK 465     ASN A   551
REMARK 465     ASN A   552
REMARK 465     TYR A   553
REMARK 465     MET A   554
REMARK 465     MET A   555
REMARK 465     ASP A   556
REMARK 465     TRP A   557
REMARK 465     LYS A   558
REMARK 465     ASN A   559
REMARK 465     GLN A   560
REMARK 465     PHE A   561
REMARK 465     ASN A   562
REMARK 465     ASP A   563
REMARK 465     TYR A   564
REMARK 465     THR A   565
REMARK 465     SER A   566
REMARK 465     LYS A   567
REMARK 465     LYS A   568
REMARK 465     GLU A   569
REMARK 465     SER A   570
REMARK 465     CYS A   571
REMARK 465     VAL A   572
REMARK 465     GLY A   573
REMARK 465     LEU A   574
REMARK 465     MET B   -27
REMARK 465     HIS B   -26
REMARK 465     SER B   -25
REMARK 465     LYS B   -24
REMARK 465     VAL B   -23
REMARK 465     THR B   -22
REMARK 465     ILE B   -21
REMARK 465     ILE B   -20
REMARK 465     CYS B   -19
REMARK 465     ILE B   -18
REMARK 465     ARG B   -17
REMARK 465     PHE B   -16
REMARK 465     LEU B   -15
REMARK 465     PHE B   -14
REMARK 465     TRP B   -13
REMARK 465     PHE B   -12
REMARK 465     LEU B   -11
REMARK 465     LEU B   -10
REMARK 465     LEU B    -9
REMARK 465     CYS B    -8
REMARK 465     MET B    -7
REMARK 465     LEU B    -6
REMARK 465     ILE B    -5
REMARK 465     GLY B    -4
REMARK 465     LYS B    -3
REMARK 465     SER B    -2
REMARK 465     HIS B    -1
REMARK 465     THR B     0
REMARK 465     GLU B     1
REMARK 465     ASP B     2
REMARK 465     ASP B     3
REMARK 465     GLN B   484
REMARK 465     ASN B   485
REMARK 465     GLU B   531
REMARK 465     MET B   532
REMARK 465     THR B   533
REMARK 465     GLY B   534
REMARK 465     ASN B   535
REMARK 465     ILE B   536
REMARK 465     ASP B   537
REMARK 465     GLU B   538
REMARK 465     ALA B   539
REMARK 465     GLU B   540
REMARK 465     TRP B   541
REMARK 465     GLU B   542
REMARK 465     TRP B   543
REMARK 465     LYS B   544
REMARK 465     ALA B   545
REMARK 465     GLY B   546
REMARK 465     PHE B   547
REMARK 465     HIS B   548
REMARK 465     ARG B   549
REMARK 465     TRP B   550
REMARK 465     ASN B   551
REMARK 465     ASN B   552
REMARK 465     TYR B   553
REMARK 465     MET B   554
REMARK 465     MET B   555
REMARK 465     ASP B   556
REMARK 465     TRP B   557
REMARK 465     LYS B   558
REMARK 465     ASN B   559
REMARK 465     GLN B   560
REMARK 465     PHE B   561
REMARK 465     ASN B   562
REMARK 465     ASP B   563
REMARK 465     TYR B   564
REMARK 465     THR B   565
REMARK 465     SER B   566
REMARK 465     LYS B   567
REMARK 465     LYS B   568
REMARK 465     GLU B   569
REMARK 465     SER B   570
REMARK 465     CYS B   571
REMARK 465     VAL B   572
REMARK 465     GLY B   573
REMARK 465     LEU B   574
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A 255    CG   CD   OE1  OE2
REMARK 470     TYR A 282    CG   CD1  CD2  CE1  CE2  CZ   OH
REMARK 470     GLU A 506    CG   CD   OE1  OE2
REMARK 470     LEU A 530    CA   C    O    CB   CG   CD1  CD2
REMARK 470     ILE B   4    CG1  CG2  CD1
REMARK 470     LYS B  51    CG   CD   CE   NZ
REMARK 470     SER B  53    OG
REMARK 470     TYR B 282    CG   CD1  CD2  CE1  CE2  CZ   OH
REMARK 470     ASN B 486    CG   OD1  ND2
REMARK 470     SER B 507    CB   OG
REMARK 470     LEU B 530    CA   C    O    CB   CG   CD1  CD2
REMARK 480
REMARK 480 ZERO OCCUPANCY ATOM
REMARK 480 THE FOLLOWING RESIDUES HAVE ATOMS MODELED WITH ZERO
REMARK 480 OCCUPANCY. THE LOCATION AND PROPERTIES OF THESE ATOMS
REMARK 480 MAY NOT BE RELIABLE. (M=MODEL NUMBER; RES=RESIDUE NAME;
REMARK 480 C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE):
REMARK 480   M RES C SSEQI ATOMS
REMARK 480     ILE B    4   O
REMARK 480     SER B   53   CB
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   ND2  ASN A   341     C1   NAG A   608              1.41
REMARK 500   O4   NAG A   608     C1   NAG A   609              1.41
REMARK 500   O6   NAG A   608     C1   FUC A   610              1.46
REMARK 500   ND2  ASN B   481     C1   NAG B   610              1.46
REMARK 500   ND2  ASN B    17     O5   NAG B   601              1.61
REMARK 500   ND2  ASN A   341     O5   NAG A   608              1.64
REMARK 500   ND2  ASN B   256     O5   NAG B   608              1.66
REMARK 500   O4   NAG A   604     O5   NAG A   605              1.74
REMARK 500   ND2  ASN B   106     O5   NAG B   603              1.79
REMARK 500   ND2  ASN A   481     O5   NAG A   611              1.92
REMARK 500   ND2  ASN A   241     O5   NAG A   606              1.94
REMARK 500   C6   NAG A   608     C1   FUC A   610              2.01
REMARK 500   ND2  ASN B    57     C2   NAG B   602              2.03
REMARK 500   CG   ASN B   256     C1   NAG B   608              2.04
REMARK 500   ND2  ASN A   486     O5   NAG A   613              2.07
REMARK 500   ND2  ASN A   106     O5   NAG A   604              2.13
REMARK 500   O4   NAG A   611     O5   NAG A   612              2.13
REMARK 500   CG   ASN B    17     C1   NAG B   601              2.14
REMARK 500   OE1  GLU A    80     O    HOH A   714              2.15
REMARK 500   CG   ASN A   241     C1   NAG A   606              2.16
REMARK 500   CG   ASN A   486     C1   NAG A   613              2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   OH   TYR A   237     O    ARG B   453     1565     2.19
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND LENGTHS
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,2(A3,1X,A1,I4,A1,1X,A4,3X),1X,F6.3)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   RES CSSEQI ATM2   DEVIATION
REMARK 500    PHE A 364   CZ    PHE A 364   CE2    -0.118
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    THR A  50      -98.63    -92.37
REMARK 500    ASP A  54     -167.49    104.36
REMARK 500    SER A  89      148.39   -170.27
REMARK 500    ASN A 106       48.86   -153.14
REMARK 500    ALA A 162       75.00   -159.38
REMARK 500    SER A 198     -117.91     65.89
REMARK 500    TYR A 282     -104.98    -71.00
REMARK 500    ASP A 297      -71.79   -127.16
REMARK 500    SER A 362      175.35    -58.11
REMARK 500    ARG A 381       80.73     46.85
REMARK 500    PHE A 398      -56.08   -126.98
REMARK 500    GLU A 482      -82.07   -105.53
REMARK 500    PHE B  43       -2.29     74.85
REMARK 500    ASP B  54     -138.09    -85.66
REMARK 500    SER B  89      148.50   -172.60
REMARK 500    ASN B 106       43.88   -145.10
REMARK 500    ALA B 162       78.74   -162.94
REMARK 500    SER B 198     -111.85     63.00
REMARK 500    TYR B 282       54.70    -90.93
REMARK 500    ASP B 297      -71.35   -136.92
REMARK 500    PHE B 398      -60.40   -127.26
REMARK 500    PRO B 480       44.77    -78.03
REMARK 500    GLU B 506      -73.41    -70.49
REMARK 500    PHE B 525      -52.40   -127.58
REMARK 500
REMARK 500 REMARK: NULL
REMARK 610
REMARK 610 MISSING HETEROATOM
REMARK 610 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 610 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 610 I=INSERTION CODE):
REMARK 610   M RES C SSEQI
REMARK 610     NAG A  608
REMARK 610     NAG A  609
REMARK 610     FUC A  610
REMARK 610     NAG B  607
REMARK 610     NAG B  610
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue 3F9 A 615
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL A 616
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 617
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO A 618
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue NDG B 607
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue 3F9 B 611
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL B 612
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL B 613
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL B 614
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue GOL B 615
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue EDO B 616
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG A 601 bound
REMARK 800  to ASN A 17
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  602 through FUL A 603 bound to ASN A 57
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  604 through NAG A 605 bound to ASN A 106
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  606 through FUC A 607 bound to ASN A 241
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NDG A
REMARK 800  608 through FUC A 610 bound to ASN A 341
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD8
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  611 through NAG A 612 bound to ASN A 481
REMARK 800
REMARK 800 SITE_IDENTIFIER: AD9
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG A
REMARK 800  613 through NAG A 614 bound to ASN A 486
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG B 601 bound
REMARK 800  to ASN B 17
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG B 602 bound
REMARK 800  to ASN B 57
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG B
REMARK 800  603 through NAG B 604 bound to ASN B 106
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Poly-Saccharide residues NAG B
REMARK 800  605 through FUL B 606 bound to ASN B 241
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG B 608 bound
REMARK 800  to ASN B 256
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG B 609 bound
REMARK 800  to ASN B 341
REMARK 800
REMARK 800 SITE_IDENTIFIER: AE7
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NDG B 610 bound
REMARK 800  to ASN B 481
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4BDS   RELATED DB: PDB
REMARK 900 4BDS IS THE SAME PROTEIN COMPLEXED WITH TACRINE
DBREF  4TPK A  -27   574  UNP    P06276   CHLE_HUMAN       1    602
DBREF  4TPK B  -27   574  UNP    P06276   CHLE_HUMAN       1    602
SEQRES   1 A  602  MET HIS SER LYS VAL THR ILE ILE CYS ILE ARG PHE LEU
SEQRES   2 A  602  PHE TRP PHE LEU LEU LEU CYS MET LEU ILE GLY LYS SER
SEQRES   3 A  602  HIS THR GLU ASP ASP ILE ILE ILE ALA THR LYS ASN GLY
SEQRES   4 A  602  LYS VAL ARG GLY MET ASN LEU THR VAL PHE GLY GLY THR
SEQRES   5 A  602  VAL THR ALA PHE LEU GLY ILE PRO TYR ALA GLN PRO PRO
SEQRES   6 A  602  LEU GLY ARG LEU ARG PHE LYS LYS PRO GLN SER LEU THR
SEQRES   7 A  602  LYS TRP SER ASP ILE TRP ASN ALA THR LYS TYR ALA ASN
SEQRES   8 A  602  SER CYS CYS GLN ASN ILE ASP GLN SER PHE PRO GLY PHE
SEQRES   9 A  602  HIS GLY SER GLU MET TRP ASN PRO ASN THR ASP LEU SER
SEQRES  10 A  602  GLU ASP CYS LEU TYR LEU ASN VAL TRP ILE PRO ALA PRO
SEQRES  11 A  602  LYS PRO LYS ASN ALA THR VAL LEU ILE TRP ILE TYR GLY
SEQRES  12 A  602  GLY GLY PHE GLN THR GLY THR SER SER LEU HIS VAL TYR
SEQRES  13 A  602  ASP GLY LYS PHE LEU ALA ARG VAL GLU ARG VAL ILE VAL
SEQRES  14 A  602  VAL SER MET ASN TYR ARG VAL GLY ALA LEU GLY PHE LEU
SEQRES  15 A  602  ALA LEU PRO GLY ASN PRO GLU ALA PRO GLY ASN MET GLY
SEQRES  16 A  602  LEU PHE ASP GLN GLN LEU ALA LEU GLN TRP VAL GLN LYS
SEQRES  17 A  602  ASN ILE ALA ALA PHE GLY GLY ASN PRO LYS SER VAL THR
SEQRES  18 A  602  LEU PHE GLY GLU SER ALA GLY ALA ALA SER VAL SER LEU
SEQRES  19 A  602  HIS LEU LEU SER PRO GLY SER HIS SER LEU PHE THR ARG
SEQRES  20 A  602  ALA ILE LEU GLN SER GLY SER PHE ASN ALA PRO TRP ALA
SEQRES  21 A  602  VAL THR SER LEU TYR GLU ALA ARG ASN ARG THR LEU ASN
SEQRES  22 A  602  LEU ALA LYS LEU THR GLY CYS SER ARG GLU ASN GLU THR
SEQRES  23 A  602  GLU ILE ILE LYS CYS LEU ARG ASN LYS ASP PRO GLN GLU
SEQRES  24 A  602  ILE LEU LEU ASN GLU ALA PHE VAL VAL PRO TYR GLY THR
SEQRES  25 A  602  PRO LEU SER VAL ASN PHE GLY PRO THR VAL ASP GLY ASP
SEQRES  26 A  602  PHE LEU THR ASP MET PRO ASP ILE LEU LEU GLU LEU GLY
SEQRES  27 A  602  GLN PHE LYS LYS THR GLN ILE LEU VAL GLY VAL ASN LYS
SEQRES  28 A  602  ASP GLU GLY THR ALA PHE LEU VAL TYR GLY ALA PRO GLY
SEQRES  29 A  602  PHE SER LYS ASP ASN ASN SER ILE ILE THR ARG LYS GLU
SEQRES  30 A  602  PHE GLN GLU GLY LEU LYS ILE PHE PHE PRO GLY VAL SER
SEQRES  31 A  602  GLU PHE GLY LYS GLU SER ILE LEU PHE HIS TYR THR ASP
SEQRES  32 A  602  TRP VAL ASP ASP GLN ARG PRO GLU ASN TYR ARG GLU ALA
SEQRES  33 A  602  LEU GLY ASP VAL VAL GLY ASP TYR ASN PHE ILE CYS PRO
SEQRES  34 A  602  ALA LEU GLU PHE THR LYS LYS PHE SER GLU TRP GLY ASN
SEQRES  35 A  602  ASN ALA PHE PHE TYR TYR PHE GLU HIS ARG SER SER LYS
SEQRES  36 A  602  LEU PRO TRP PRO GLU TRP MET GLY VAL MET HIS GLY TYR
SEQRES  37 A  602  GLU ILE GLU PHE VAL PHE GLY LEU PRO LEU GLU ARG ARG
SEQRES  38 A  602  ASP ASN TYR THR LYS ALA GLU GLU ILE LEU SER ARG SER
SEQRES  39 A  602  ILE VAL LYS ARG TRP ALA ASN PHE ALA LYS TYR GLY ASN
SEQRES  40 A  602  PRO ASN GLU THR GLN ASN ASN SER THR SER TRP PRO VAL
SEQRES  41 A  602  PHE LYS SER THR GLU GLN LYS TYR LEU THR LEU ASN THR
SEQRES  42 A  602  GLU SER THR ARG ILE MET THR LYS LEU ARG ALA GLN GLN
SEQRES  43 A  602  CYS ARG PHE TRP THR SER PHE PHE PRO LYS VAL LEU GLU
SEQRES  44 A  602  MET THR GLY ASN ILE ASP GLU ALA GLU TRP GLU TRP LYS
SEQRES  45 A  602  ALA GLY PHE HIS ARG TRP ASN ASN TYR MET MET ASP TRP
SEQRES  46 A  602  LYS ASN GLN PHE ASN ASP TYR THR SER LYS LYS GLU SER
SEQRES  47 A  602  CYS VAL GLY LEU
SEQRES   1 B  602  MET HIS SER LYS VAL THR ILE ILE CYS ILE ARG PHE LEU
SEQRES   2 B  602  PHE TRP PHE LEU LEU LEU CYS MET LEU ILE GLY LYS SER
SEQRES   3 B  602  HIS THR GLU ASP ASP ILE ILE ILE ALA THR LYS ASN GLY
SEQRES   4 B  602  LYS VAL ARG GLY MET ASN LEU THR VAL PHE GLY GLY THR
SEQRES   5 B  602  VAL THR ALA PHE LEU GLY ILE PRO TYR ALA GLN PRO PRO
SEQRES   6 B  602  LEU GLY ARG LEU ARG PHE LYS LYS PRO GLN SER LEU THR
SEQRES   7 B  602  LYS TRP SER ASP ILE TRP ASN ALA THR LYS TYR ALA ASN
SEQRES   8 B  602  SER CYS CYS GLN ASN ILE ASP GLN SER PHE PRO GLY PHE
SEQRES   9 B  602  HIS GLY SER GLU MET TRP ASN PRO ASN THR ASP LEU SER
SEQRES  10 B  602  GLU ASP CYS LEU TYR LEU ASN VAL TRP ILE PRO ALA PRO
SEQRES  11 B  602  LYS PRO LYS ASN ALA THR VAL LEU ILE TRP ILE TYR GLY
SEQRES  12 B  602  GLY GLY PHE GLN THR GLY THR SER SER LEU HIS VAL TYR
SEQRES  13 B  602  ASP GLY LYS PHE LEU ALA ARG VAL GLU ARG VAL ILE VAL
SEQRES  14 B  602  VAL SER MET ASN TYR ARG VAL GLY ALA LEU GLY PHE LEU
SEQRES  15 B  602  ALA LEU PRO GLY ASN PRO GLU ALA PRO GLY ASN MET GLY
SEQRES  16 B  602  LEU PHE ASP GLN GLN LEU ALA LEU GLN TRP VAL GLN LYS
SEQRES  17 B  602  ASN ILE ALA ALA PHE GLY GLY ASN PRO LYS SER VAL THR
SEQRES  18 B  602  LEU PHE GLY GLU SER ALA GLY ALA ALA SER VAL SER LEU
SEQRES  19 B  602  HIS LEU LEU SER PRO GLY SER HIS SER LEU PHE THR ARG
SEQRES  20 B  602  ALA ILE LEU GLN SER GLY SER PHE ASN ALA PRO TRP ALA
SEQRES  21 B  602  VAL THR SER LEU TYR GLU ALA ARG ASN ARG THR LEU ASN
SEQRES  22 B  602  LEU ALA LYS LEU THR GLY CYS SER ARG GLU ASN GLU THR
SEQRES  23 B  602  GLU ILE ILE LYS CYS LEU ARG ASN LYS ASP PRO GLN GLU
SEQRES  24 B  602  ILE LEU LEU ASN GLU ALA PHE VAL VAL PRO TYR GLY THR
SEQRES  25 B  602  PRO LEU SER VAL ASN PHE GLY PRO THR VAL ASP GLY ASP
SEQRES  26 B  602  PHE LEU THR ASP MET PRO ASP ILE LEU LEU GLU LEU GLY
SEQRES  27 B  602  GLN PHE LYS LYS THR GLN ILE LEU VAL GLY VAL ASN LYS
SEQRES  28 B  602  ASP GLU GLY THR ALA PHE LEU VAL TYR GLY ALA PRO GLY
SEQRES  29 B  602  PHE SER LYS ASP ASN ASN SER ILE ILE THR ARG LYS GLU
SEQRES  30 B  602  PHE GLN GLU GLY LEU LYS ILE PHE PHE PRO GLY VAL SER
SEQRES  31 B  602  GLU PHE GLY LYS GLU SER ILE LEU PHE HIS TYR THR ASP
SEQRES  32 B  602  TRP VAL ASP ASP GLN ARG PRO GLU ASN TYR ARG GLU ALA
SEQRES  33 B  602  LEU GLY ASP VAL VAL GLY ASP TYR ASN PHE ILE CYS PRO
SEQRES  34 B  602  ALA LEU GLU PHE THR LYS LYS PHE SER GLU TRP GLY ASN
SEQRES  35 B  602  ASN ALA PHE PHE TYR TYR PHE GLU HIS ARG SER SER LYS
SEQRES  36 B  602  LEU PRO TRP PRO GLU TRP MET GLY VAL MET HIS GLY TYR
SEQRES  37 B  602  GLU ILE GLU PHE VAL PHE GLY LEU PRO LEU GLU ARG ARG
SEQRES  38 B  602  ASP ASN TYR THR LYS ALA GLU GLU ILE LEU SER ARG SER
SEQRES  39 B  602  ILE VAL LYS ARG TRP ALA ASN PHE ALA LYS TYR GLY ASN
SEQRES  40 B  602  PRO ASN GLU THR GLN ASN ASN SER THR SER TRP PRO VAL
SEQRES  41 B  602  PHE LYS SER THR GLU GLN LYS TYR LEU THR LEU ASN THR
SEQRES  42 B  602  GLU SER THR ARG ILE MET THR LYS LEU ARG ALA GLN GLN
SEQRES  43 B  602  CYS ARG PHE TRP THR SER PHE PHE PRO LYS VAL LEU GLU
SEQRES  44 B  602  MET THR GLY ASN ILE ASP GLU ALA GLU TRP GLU TRP LYS
SEQRES  45 B  602  ALA GLY PHE HIS ARG TRP ASN ASN TYR MET MET ASP TRP
SEQRES  46 B  602  LYS ASN GLN PHE ASN ASP TYR THR SER LYS LYS GLU SER
SEQRES  47 B  602  CYS VAL GLY LEU
HET    NAG  A 601      14
HET    NAG  A 602      14
HET    FUL  A 603      10
HET    NAG  A 604      14
HET    NAG  A 605      14
HET    NAG  A 606      14
HET    FUC  A 607      10
HET    NAG  A 608      14
HET    NAG  A 609      14
HET    FUC  A 610      10
HET    NAG  A 611      14
HET    NAG  A 612      14
HET    NAG  A 613      14
HET    NAG  A 614      14
HET    3F9  A 615      33
HET    GOL  A 616       6
HET    EDO  A 617       4
HET    EDO  A 618       4
HET    NAG  B 601      14
HET    NAG  B 602      14
HET    NAG  B 603      14
HET    NAG  B 604      14
HET    NAG  B 605      14
HET    FUL  B 606      10
HET    NAG  B 607      14
HET    NAG  B 608      14
HET    NAG  B 609      14
HET    NAG  B 610      14
HET    3F9  B 611      33
HET    GOL  B 612       6
HET    GOL  B 613       6
HET    GOL  B 614       6
HET    GOL  B 615       6
HET    EDO  B 616       4
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     FUL BETA-L-FUCOSE
HETNAM     FUC ALPHA-L-FUCOSE
HETNAM     3F9 N-{[(3R)-1-(2,3-DIHYDRO-1H-INDEN-2-YL)PIPERIDIN-3-
HETNAM   2 3F9  YL]METHYL}-N-(2-METHOXYETHYL)NAPHTHALENE-2-CARBOXAMIDE
HETNAM     GOL GLYCEROL
HETNAM     EDO 1,2-ETHANEDIOL
HETSYN     FUL 6-DEOXY-BETA-L-GALACTOSE
HETSYN     GOL GLYCERIN; PROPANE-1,2,3-TRIOL
HETSYN     EDO ETHYLENE GLYCOL
FORMUL   3  NAG    20(C8 H15 N O6)
FORMUL   4  FUL    2(C6 H12 O5)
FORMUL   6  FUC    2(C6 H12 O5)
FORMUL  10  3F9    2(C29 H34 N2 O2)
FORMUL  11  GOL    5(C3 H8 O3)
FORMUL  12  EDO    3(C2 H6 O2)
FORMUL  28  HOH   *172(H2 O)
HELIX    1 AA1 LEU A   38  ARG A   42  5                                   5
HELIX    2 AA2 PHE A   76  MET A   81  1                                   6
HELIX    3 AA3 LEU A  125  ASP A  129  5                                   5
HELIX    4 AA4 GLY A  130  ARG A  138  1                                   9
HELIX    5 AA5 GLY A  149  LEU A  154  1                                   6
HELIX    6 AA6 ASN A  165  ILE A  182  1                                  18
HELIX    7 AA7 ALA A  183  PHE A  185  5                                   3
HELIX    8 AA8 SER A  198  SER A  210  1                                  13
HELIX    9 AA9 SER A  210  SER A  215  1                                   6
HELIX   10 AB1 SER A  235  THR A  250  1                                  16
HELIX   11 AB2 ASN A  256  ARG A  265  1                                  10
HELIX   12 AB3 ASP A  268  LEU A  274  1                                   7
HELIX   13 AB4 ASN A  275  VAL A  280  5                                   6
HELIX   14 AB5 MET A  302  LEU A  309  1                                   8
HELIX   15 AB6 GLY A  326  VAL A  331  1                                   6
HELIX   16 AB7 THR A  346  PHE A  358  1                                  13
HELIX   17 AB8 SER A  362  TYR A  373  1                                  12
HELIX   18 AB9 GLU A  383  PHE A  398  1                                  16
HELIX   19 AC1 PHE A  398  GLU A  411  1                                  14
HELIX   20 AC2 PRO A  431  GLY A  435  5                                   5
HELIX   21 AC3 GLU A  441  PHE A  446  1                                   6
HELIX   22 AC4 GLY A  447  ASN A  455  5                                   9
HELIX   23 AC5 THR A  457  GLY A  478  1                                  22
HELIX   24 AC6 ARG A  515  PHE A  525  1                                  11
HELIX   25 AC7 PHE A  526  VAL A  529  5                                   4
HELIX   26 AC8 LEU B   38  ARG B   42  5                                   5
HELIX   27 AC9 PHE B   76  MET B   81  1                                   6
HELIX   28 AD1 LEU B  125  ASP B  129  5                                   5
HELIX   29 AD2 GLY B  130  ARG B  138  1                                   9
HELIX   30 AD3 GLY B  149  LEU B  154  1                                   6
HELIX   31 AD4 ASN B  165  ILE B  182  1                                  18
HELIX   32 AD5 ALA B  183  PHE B  185  5                                   3
HELIX   33 AD6 SER B  198  SER B  210  1                                  13
HELIX   34 AD7 PRO B  211  PHE B  217  5                                   7
HELIX   35 AD8 SER B  235  THR B  250  1                                  16
HELIX   36 AD9 ASN B  256  ARG B  265  1                                  10
HELIX   37 AE1 ASP B  268  LEU B  274  1                                   7
HELIX   38 AE2 ASN B  275  VAL B  280  5                                   6
HELIX   39 AE3 MET B  302  GLY B  310  1                                   9
HELIX   40 AE4 GLY B  326  VAL B  331  1                                   6
HELIX   41 AE5 THR B  346  PHE B  358  1                                  13
HELIX   42 AE6 SER B  362  TYR B  373  1                                  12
HELIX   43 AE7 GLU B  383  PHE B  398  1                                  16
HELIX   44 AE8 PHE B  398  GLU B  411  1                                  14
HELIX   45 AE9 PRO B  431  GLY B  435  5                                   5
HELIX   46 AF1 GLU B  441  PHE B  446  1                                   6
HELIX   47 AF2 GLY B  447  ASN B  455  5                                   9
HELIX   48 AF3 THR B  457  GLY B  478  1                                  22
HELIX   49 AF4 ARG B  515  PHE B  525  1                                  11
HELIX   50 AF5 PHE B  526  VAL B  529  5                                   4
SHEET    1 AA1 3 ILE A   5  THR A   8  0
SHEET    2 AA1 3 GLY A  11  ARG A  14 -1  O  VAL A  13   N  ILE A   6
SHEET    3 AA1 3 ILE A  55  ASN A  57  1  O  TRP A  56   N  ARG A  14
SHEET    1 AA211 MET A  16  VAL A  20  0
SHEET    2 AA211 GLY A  23  PRO A  32 -1  O  ALA A  27   N  MET A  16
SHEET    3 AA211 TYR A  94  ALA A 101 -1  O  LEU A  95   N  ILE A  31
SHEET    4 AA211 ILE A 140  MET A 144 -1  O  SER A 143   N  ASN A  96
SHEET    5 AA211 ALA A 107  ILE A 113  1  N  LEU A 110   O  ILE A 140
SHEET    6 AA211 GLY A 187  GLU A 197  1  O  ASN A 188   N  ALA A 107
SHEET    7 AA211 ARG A 219  GLN A 223  1  O  ILE A 221   N  LEU A 194
SHEET    8 AA211 ILE A 317  ASN A 322  1  O  LEU A 318   N  LEU A 222
SHEET    9 AA211 ALA A 416  PHE A 421  1  O  PHE A 417   N  VAL A 319
SHEET   10 AA211 LYS A 499  LEU A 503  1  O  LEU A 503   N  TYR A 420
SHEET   11 AA211 ILE A 510  THR A 512 -1  O  MET A 511   N  TYR A 500
SHEET    1 AA3 3 ILE B   5  THR B   8  0
SHEET    2 AA3 3 GLY B  11  ARG B  14 -1  O  GLY B  11   N  THR B   8
SHEET    3 AA3 3 ILE B  55  ASN B  57  1  O  TRP B  56   N  LYS B  12
SHEET    1 AA411 MET B  16  VAL B  20  0
SHEET    2 AA411 GLY B  23  PRO B  32 -1  O  VAL B  25   N  LEU B  18
SHEET    3 AA411 TYR B  94  ALA B 101 -1  O  VAL B  97   N  PHE B  28
SHEET    4 AA411 ILE B 140  MET B 144 -1  O  SER B 143   N  ASN B  96
SHEET    5 AA411 ALA B 107  ILE B 113  1  N  TRP B 112   O  VAL B 142
SHEET    6 AA411 GLY B 187  GLU B 197  1  O  THR B 193   N  VAL B 109
SHEET    7 AA411 ARG B 219  GLN B 223  1  O  ILE B 221   N  LEU B 194
SHEET    8 AA411 ILE B 317  ASN B 322  1  O  LEU B 318   N  LEU B 222
SHEET    9 AA411 ALA B 416  PHE B 421  1  O  PHE B 417   N  VAL B 319
SHEET   10 AA411 LYS B 499  LEU B 503  1  O  LEU B 503   N  TYR B 420
SHEET   11 AA411 ILE B 510  THR B 512 -1  O  MET B 511   N  TYR B 500
SSBOND   1 CYS A   65    CYS A   92                          1555   1555  2.04
SSBOND   2 CYS A  252    CYS A  263                          1555   1555  2.04
SSBOND   3 CYS A  400    CYS A  519                          1555   1555  2.06
SSBOND   4 CYS B   65    CYS B   92                          1555   1555  2.05
SSBOND   5 CYS B  252    CYS B  263                          1555   1555  2.05
SSBOND   6 CYS B  400    CYS B  519                          1555   1555  2.06
LINK         ND2 ASN A  17                 C1  NAG A 601     1555   1555  1.49
LINK         ND2 ASN A  57                 C1  NAG A 602     1555   1555  1.47
LINK         ND2 ASN A 106                 C1  NAG A 604     1555   1555  1.42
LINK         ND2 ASN A 241                 C1  NAG A 606     1555   1555  1.45
LINK         ND2 ASN A 481                 C1  NAG A 611     1555   1555  1.41
LINK         ND2 ASN A 486                 C1  NAG A 613     1555   1555  1.45
LINK         ND2 ASN B  17                 C1  NAG B 601     1555   1555  1.48
LINK         ND2 ASN B  57                 C1  NAG B 602     1555   1555  1.47
LINK         ND2 ASN B 106                 C1  NAG B 603     1555   1555  1.44
LINK         ND2 ASN B 241                 C1  NAG B 605     1555   1555  1.44
LINK         ND2 ASN B 256                 C1  NAG B 608     1555   1555  1.46
LINK         ND2 ASN B 341                 C1  NAG B 609     1555   1555  1.49
LINK         O6  NAG A 602                 C1  FUL A 603     1555   1555  1.42
LINK         O4  NAG A 604                 C1  NAG A 605     1555   1555  1.46
LINK         O6  NAG A 606                 C1  FUC A 607     1555   1555  1.46
LINK         O4  NAG A 611                 C1  NAG A 612     1555   1555  1.47
LINK         O4  NAG A 613                 C1  NAG A 614     1555   1555  1.45
LINK         O4  NAG B 603                 C1  NAG B 604     1555   1555  1.49
LINK         O6  NAG B 605                 C1  FUL B 606     1555   1555  1.49
CISPEP   1 ALA A  101    PRO A  102          0         4.37
CISPEP   2 GLN A  380    ARG A  381          0        -4.58
CISPEP   3 ALA B  101    PRO B  102          0         4.37
SITE     1 AC1 13 ASN A  68  ILE A  69  ASP A  70  TRP A  82
SITE     2 AC1 13 GLY A 116  GLY A 117  GLU A 197  SER A 198
SITE     3 AC1 13 PHE A 329  TYR A 332  GLY A 439  HOH A 767
SITE     4 AC1 13 HOH A 781
SITE     1 AC2  3 ASN A 228  CYS A 400  PRO A 401
SITE     1 AC3  2 HIS A  77  GLU A 443
SITE     1 AC4  4 LEU A  18  TRP A  98  ASP A 129  LYS A 131
SITE     1 AC5  1 NAG B 605
SITE     1 AC6 10 ASP B  70  TRP B  82  GLY B 116  SER B 198
SITE     2 AC6 10 TRP B 231  LEU B 286  PHE B 329  TYR B 332
SITE     3 AC6 10 HOH B 753  HOH B 785
SITE     1 AC7  5 LEU B  18  TYR B  61  TRP B  98  ASP B 129
SITE     2 AC7  5 LYS B 131
SITE     1 AC8  6 HIS B 372  TYR B 373  THR B 374  ASP B 375
SITE     2 AC8  6 GLN B 518  HOH B 783
SITE     1 AC9  5 SER A 489  ILE B  69  GLN B  71  SER B  72
SITE     2 AC9  5 HOH B 740
SITE     1 AD1  3 HIS B  77  MET B  81  GLU B 443
SITE     1 AD2  2 ARG B  42  LYS B 267
SITE     1 AD3  1 ASN A  17
SITE     1 AD4  3 ASP A  54  ILE A  55  ASN A  57
SITE     1 AD5  4 LYS A 105  ASN A 106  ASN A 188  LYS A 190
SITE     1 AD6  3 ASN A 241  ASN A 245  PHE A 278
SITE     1 AD7  4 PRO A 335  GLY A 336  SER A 338  ASN A 341
SITE     1 AD8  6 TYR A 477  ASN A 481  GLU A 482  GLN A 484
SITE     2 AD8  6 LEU B  88  GLN B 270
SITE     1 AD9  1 ASN A 486
SITE     1 AE1  2 ASN B  17  THR B  24
SITE     1 AE2  4 ARG B  14  ASP B  54  ILE B  55  ASN B  57
SITE     1 AE3  4 ASN B 106  ASN B 188  LYS B 190  HOH B 718
SITE     1 AE4  5 ASN B 241  ASN B 245  LYS B 248  PHE B 278
SITE     2 AE4  5 HOH B 748
SITE     1 AE5  2 ASN B 256  HOH B 745
SITE     1 AE6  5 ASN B 341  ASN B 342  HOH B 746  HOH B 747
SITE     2 AE6  5 HOH B 789
SITE     1 AE7  3 ASN B 481  GLU B 482  HOH B 780
CRYST1   76.600   80.200  231.650  90.00  90.00  90.00 P 21 21 21    8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.013055  0.000000  0.000000        0.00000
SCALE2      0.000000  0.012469  0.000000        0.00000
SCALE3      0.000000  0.000000  0.004317        0.00000
TER    4209      LEU A 530
TER    8381      LEU B 530
MASTER      625    0   34   50   28    0   36    6 8938    2  452   94
END