longtext: 4UYZ-pdb

content
HEADER    HYDROLASE                               03-SEP-14   4UYZ
TITLE     STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM II - 2.8A
CAVEAT     4UYZ    NAG B 1452 IS PLANAR AT C1
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PROTEIN NOTUM HOMOLOG;
COMPND   3 CHAIN: A, B, C, D;
COMPND   4 FRAGMENT: RESIDUES 38-496;
COMPND   5 SYNONYM: NOTUM;
COMPND   6 EC: 3.1.1.1;
COMPND   7 ENGINEERED: YES;
COMPND   8 OTHER_DETAILS: GLYCOSYLATED AT N96;
COMPND   9 MOL_ID: 2;
COMPND  10 MOLECULE: POLY ALA;
COMPND  11 CHAIN: E;
COMPND  12 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE   6 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE   8 EXPRESSION_SYSTEM_CELL_LINE: HEK293T;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PHLSEC;
SOURCE  10 MOL_ID: 2;
SOURCE  11 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE  12 ORGANISM_COMMON: HUMAN;
SOURCE  13 ORGANISM_TAXID: 9606;
SOURCE  14 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE  15 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE  16 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE  17 EXPRESSION_SYSTEM_CELL_LINE: HEK293T;
SOURCE  18 EXPRESSION_SYSTEM_PLASMID: PHLSEC
KEYWDS    HYDROLASE, WNT, ESTERASE, EXTRACELLULAR, ALPHA/BETA HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.ZEBISCH,E.Y.JONES
REVDAT   1   25-FEB-15 4UYZ    0
JRNL        AUTH   S.KAKUGAWA,P.F.LANGTON,M.ZEBISCH,S.A.HOWELL,T.-H.CHANG,
JRNL        AUTH 2 Y.LIU,T.FEIZI,G.BINEVA,N.O'REILLY,A.P.SNIJDERS,E.Y.JONES,
JRNL        AUTH 3 J.-P.VINCENT
JRNL        TITL   NOTUM DEACYLATES WNT PROTEINS TO SUPPRESS SIGNALLING
JRNL        TITL 2 ACTIVITY
JRNL        REF    NATURE                                     2015
JRNL        REFN                   ESSN 1476-4687
JRNL        DOI    10.1038/NATURE14259
REMARK   2
REMARK   2 RESOLUTION.    2.80 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0073
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,
REMARK   3                 STEINER,NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.80
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 96.94
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NONE
REMARK   3   COMPLETENESS FOR RANGE        (%) : 88.93
REMARK   3   NUMBER OF REFLECTIONS             : 37125
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.24899
REMARK   3   R VALUE            (WORKING SET) : 0.24761
REMARK   3   FREE R VALUE                     : 0.29294
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 3.0
REMARK   3   FREE R VALUE TEST SET COUNT      : 1143
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.800
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.873
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2549
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 83.68
REMARK   3   BIN R VALUE           (WORKING SET) : 0.369
REMARK   3   BIN FREE R VALUE SET COUNT          : 77
REMARK   3   BIN FREE R VALUE                    : 0.401
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 10976
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 15
REMARK   3   SOLVENT ATOMS            : 0
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 60.8
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 46.978
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 2.28
REMARK   3    B22 (A**2) : -3.92
REMARK   3    B33 (A**2) : 1.64
REMARK   3    B12 (A**2) : 0.00
REMARK   3    B13 (A**2) : -0.06
REMARK   3    B23 (A**2) : 0.00
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): NULL
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.469
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.443
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 52.567
REMARK   3
REMARK   3  CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.883
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.851
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A): 11304 ; 0.007 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES): 15399 ; 1.267 ; 1.941
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):  1379 ; 5.896 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   522 ;33.218 ;22.337
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):  1732 ;17.490 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):   110 ;19.066 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):  1654 ; 0.086 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  8726 ; 0.005 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  5567 ; 0.916 ; 2.174
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  6929 ; 1.649 ; 3.253
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  5737 ; 1.135 ; 2.270
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ANISOTROPIC THERMAL FACTOR RESTRAINTS.   COUNT   RMS    WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 4
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B     1        B   451
REMARK   3    ORIGIN FOR THE GROUP (A):  63.0104 -27.1863   9.2323
REMARK   3    T TENSOR
REMARK   3      T11:   0.0787 T22:   0.5262
REMARK   3      T33:   0.0422 T12:  -0.0168
REMARK   3      T13:   0.0107 T23:  -0.0519
REMARK   3    L TENSOR
REMARK   3      L11:   1.4455 L22:   0.5150
REMARK   3      L33:   1.9008 L12:   0.3566
REMARK   3      L13:   0.1645 L23:   0.7065
REMARK   3    S TENSOR
REMARK   3      S11:  -0.1113 S12:  -0.0530 S13:   0.0802
REMARK   3      S21:  -0.1689 S22:   0.0477 S23:  -0.0300
REMARK   3      S31:  -0.1483 S32:  -0.0206 S33:   0.0637
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A    87        A   452
REMARK   3    ORIGIN FOR THE GROUP (A):  29.6177 -10.4429  41.7262
REMARK   3    T TENSOR
REMARK   3      T11:   0.0456 T22:   0.6676
REMARK   3      T33:   0.0697 T12:  -0.0005
REMARK   3      T13:  -0.0361 T23:  -0.0899
REMARK   3    L TENSOR
REMARK   3      L11:   0.7177 L22:   1.2475
REMARK   3      L33:   1.5071 L12:  -0.2226
REMARK   3      L13:   0.0596 L23:   0.8126
REMARK   3    S TENSOR
REMARK   3      S11:   0.0057 S12:   0.0463 S13:  -0.1531
REMARK   3      S21:   0.0803 S22:  -0.0915 S23:  -0.0160
REMARK   3      S31:  -0.1331 S32:  -0.0290 S33:   0.0858
REMARK   3
REMARK   3   TLS GROUP : 3
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   C    87        C   450
REMARK   3    ORIGIN FOR THE GROUP (A):  64.4630  38.0989  28.8750
REMARK   3    T TENSOR
REMARK   3      T11:   0.0749 T22:   0.5431
REMARK   3      T33:   0.0411 T12:   0.0669
REMARK   3      T13:  -0.0274 T23:  -0.0373
REMARK   3    L TENSOR
REMARK   3      L11:   1.5376 L22:   0.8592
REMARK   3      L33:   1.4980 L12:  -0.1716
REMARK   3      L13:   0.4387 L23:   0.2988
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0977 S12:   0.0420 S13:  -0.1735
REMARK   3      S21:   0.2457 S22:   0.1755 S23:  -0.1132
REMARK   3      S31:   0.0105 S32:  -0.0526 S33:  -0.0778
REMARK   3
REMARK   3   TLS GROUP : 4
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   D    87        D   472
REMARK   3    ORIGIN FOR THE GROUP (A):  32.2754  20.2252  -2.1352
REMARK   3    T TENSOR
REMARK   3      T11:   0.0173 T22:   0.5940
REMARK   3      T33:   0.0402 T12:   0.0289
REMARK   3      T13:   0.0039 T23:  -0.0946
REMARK   3    L TENSOR
REMARK   3      L11:   0.6694 L22:   1.7276
REMARK   3      L33:   1.2005 L12:   0.2218
REMARK   3      L13:   0.4629 L23:   0.7654
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0259 S12:  -0.0518 S13:   0.1062
REMARK   3      S21:  -0.1061 S22:  -0.1133 S23:   0.0093
REMARK   3      S31:   0.0270 S32:  -0.0413 S33:   0.1392
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3   RIDING POSITIONS.
REMARK   4
REMARK   4 4UYZ COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 04-SEP-14.
REMARK 100 THE PDBE ID CODE IS EBI-61677.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 28-JAN-12
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I24
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 1.02
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : NULL
REMARK 200  DETECTOR MANUFACTURER          : NULL
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 38300
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.80
REMARK 200  RESOLUTION RANGE LOW       (A) : 58.00
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 89.1
REMARK 200  DATA REDUNDANCY                : 2.1
REMARK 200  R MERGE                    (I) : 0.16
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 6.60
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: OTHER
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NONE
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 41.26
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.09
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: NULL
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 1 21 1
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000  1.000000  0.000000       96.94000
REMARK 290   SMTRY3   2  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1, 2, 3, 4, 5
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 2
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 3
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: C
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 4
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: D
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 350
REMARK 350 BIOMOLECULE: 5
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: E
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A    35
REMARK 465     THR A    36
REMARK 465     GLY A    37
REMARK 465     ARG A    38
REMARK 465     THR A    39
REMARK 465     GLU A    40
REMARK 465     ALA A    41
REMARK 465     ALA A    42
REMARK 465     PRO A    43
REMARK 465     ALA A    44
REMARK 465     ALA A    45
REMARK 465     GLY A    46
REMARK 465     GLN A    47
REMARK 465     PRO A    48
REMARK 465     VAL A    49
REMARK 465     GLU A    50
REMARK 465     SER A    51
REMARK 465     PHE A    52
REMARK 465     PRO A    53
REMARK 465     LEU A    54
REMARK 465     ASP A    55
REMARK 465     PHE A    56
REMARK 465     THR A    57
REMARK 465     ALA A    58
REMARK 465     VAL A    59
REMARK 465     GLU A    60
REMARK 465     GLY A    61
REMARK 465     ASN A    62
REMARK 465     MET A    63
REMARK 465     ASP A    64
REMARK 465     SER A    65
REMARK 465     PHE A    66
REMARK 465     MET A    67
REMARK 465     ALA A    68
REMARK 465     GLN A    69
REMARK 465     VAL A    70
REMARK 465     LYS A    71
REMARK 465     SER A    72
REMARK 465     LEU A    73
REMARK 465     ALA A    74
REMARK 465     GLN A    75
REMARK 465     SER A    76
REMARK 465     LEU A    77
REMARK 465     TYR A    78
REMARK 465     PRO A    79
REMARK 465     CYS A    80
REMARK 465     SER A    81
REMARK 465     ALA A    82
REMARK 465     GLN A    83
REMARK 465     GLN A    84
REMARK 465     LEU A    85
REMARK 465     ASN A    86
REMARK 465     SER A   195
REMARK 465     GLU A   196
REMARK 465     LYS A   197
REMARK 465     ASN A   198
REMARK 465     THR A   352
REMARK 465     GLY A   353
REMARK 465     GLN A   354
REMARK 465     PRO A   355
REMARK 465     HIS A   419
REMARK 465     ASP A   420
REMARK 465     SER A   421
REMARK 465     HIS A   422
REMARK 465     LYS A   423
REMARK 465     ALA A   424
REMARK 465     SER A   425
REMARK 465     LYS A   426
REMARK 465     THR A   427
REMARK 465     PRO A   428
REMARK 465     LEU A   429
REMARK 465     LYS A   430
REMARK 465     GLY A   431
REMARK 465     ARG A   453
REMARK 465     ASP A   454
REMARK 465     GLN A   455
REMARK 465     PHE A   456
REMARK 465     THR A   457
REMARK 465     GLY A   458
REMARK 465     GLN A   459
REMARK 465     GLU A   460
REMARK 465     MET A   461
REMARK 465     ASN A   462
REMARK 465     VAL A   463
REMARK 465     ALA A   464
REMARK 465     GLN A   465
REMARK 465     PHE A   466
REMARK 465     LEU A   467
REMARK 465     MET A   468
REMARK 465     HIS A   469
REMARK 465     MET A   470
REMARK 465     GLY A   471
REMARK 465     PHE A   472
REMARK 465     ASP A   473
REMARK 465     MET A   474
REMARK 465     GLN A   475
REMARK 465     THR A   476
REMARK 465     VAL A   477
REMARK 465     ALA A   478
REMARK 465     GLN A   479
REMARK 465     PRO A   480
REMARK 465     GLN A   481
REMARK 465     GLY A   482
REMARK 465     LEU A   483
REMARK 465     GLU A   484
REMARK 465     PRO A   485
REMARK 465     SER A   486
REMARK 465     GLU A   487
REMARK 465     LEU A   488
REMARK 465     LEU A   489
REMARK 465     GLY A   490
REMARK 465     MET A   491
REMARK 465     LEU A   492
REMARK 465     SER A   493
REMARK 465     ASN A   494
REMARK 465     GLY A   495
REMARK 465     SER A   496
REMARK 465     GLY A   497
REMARK 465     THR A   498
REMARK 465     LYS A   499
REMARK 465     HIS A   500
REMARK 465     HIS A   501
REMARK 465     HIS A   502
REMARK 465     HIS A   503
REMARK 465     HIS A   504
REMARK 465     HIS A   505
REMARK 465     GLU B    35
REMARK 465     THR B    36
REMARK 465     GLY B    37
REMARK 465     ARG B    38
REMARK 465     THR B    39
REMARK 465     GLU B    40
REMARK 465     ALA B    41
REMARK 465     ALA B    42
REMARK 465     PRO B    43
REMARK 465     ALA B    44
REMARK 465     ALA B    45
REMARK 465     GLY B    46
REMARK 465     GLN B    47
REMARK 465     PRO B    48
REMARK 465     VAL B    49
REMARK 465     GLU B    50
REMARK 465     SER B    51
REMARK 465     PHE B    52
REMARK 465     PRO B    53
REMARK 465     LEU B    54
REMARK 465     ASP B    55
REMARK 465     PHE B    56
REMARK 465     THR B    57
REMARK 465     ALA B    58
REMARK 465     VAL B    59
REMARK 465     GLU B    60
REMARK 465     GLY B    61
REMARK 465     ASN B    62
REMARK 465     MET B    63
REMARK 465     ASP B    64
REMARK 465     SER B    65
REMARK 465     PHE B    66
REMARK 465     MET B    67
REMARK 465     ALA B    68
REMARK 465     GLN B    69
REMARK 465     VAL B    70
REMARK 465     LYS B    71
REMARK 465     SER B    72
REMARK 465     LEU B    73
REMARK 465     ALA B    74
REMARK 465     GLN B    75
REMARK 465     SER B    76
REMARK 465     LEU B    77
REMARK 465     TYR B    78
REMARK 465     PRO B    79
REMARK 465     CYS B    80
REMARK 465     SER B    81
REMARK 465     ALA B    82
REMARK 465     GLN B    83
REMARK 465     GLN B    84
REMARK 465     LEU B    85
REMARK 465     ASN B    86
REMARK 465     SER B   193
REMARK 465     LYS B   194
REMARK 465     SER B   195
REMARK 465     GLU B   196
REMARK 465     LYS B   197
REMARK 465     ASN B   198
REMARK 465     GLU B   199
REMARK 465     HIS B   350
REMARK 465     LEU B   351
REMARK 465     THR B   352
REMARK 465     GLY B   353
REMARK 465     GLN B   354
REMARK 465     PRO B   355
REMARK 465     VAL B   356
REMARK 465     GLN B   357
REMARK 465     GLU B   358
REMARK 465     GLY B   359
REMARK 465     LEU B   360
REMARK 465     ASP B   420
REMARK 465     SER B   421
REMARK 465     HIS B   422
REMARK 465     LYS B   423
REMARK 465     ALA B   424
REMARK 465     SER B   425
REMARK 465     LYS B   426
REMARK 465     THR B   427
REMARK 465     PRO B   428
REMARK 465     LEU B   429
REMARK 465     LYS B   430
REMARK 465     GLY B   431
REMARK 465     VAL B   452
REMARK 465     ARG B   453
REMARK 465     ASP B   454
REMARK 465     GLN B   455
REMARK 465     PHE B   456
REMARK 465     THR B   457
REMARK 465     GLY B   458
REMARK 465     GLN B   459
REMARK 465     GLU B   460
REMARK 465     MET B   461
REMARK 465     ASN B   462
REMARK 465     VAL B   463
REMARK 465     ALA B   464
REMARK 465     GLN B   465
REMARK 465     PHE B   466
REMARK 465     LEU B   467
REMARK 465     MET B   468
REMARK 465     HIS B   469
REMARK 465     MET B   470
REMARK 465     GLY B   471
REMARK 465     PHE B   472
REMARK 465     ASP B   473
REMARK 465     MET B   474
REMARK 465     GLN B   475
REMARK 465     THR B   476
REMARK 465     VAL B   477
REMARK 465     ALA B   478
REMARK 465     GLN B   479
REMARK 465     PRO B   480
REMARK 465     GLN B   481
REMARK 465     GLY B   482
REMARK 465     LEU B   483
REMARK 465     GLU B   484
REMARK 465     PRO B   485
REMARK 465     SER B   486
REMARK 465     GLU B   487
REMARK 465     LEU B   488
REMARK 465     LEU B   489
REMARK 465     GLY B   490
REMARK 465     MET B   491
REMARK 465     LEU B   492
REMARK 465     SER B   493
REMARK 465     ASN B   494
REMARK 465     GLY B   495
REMARK 465     SER B   496
REMARK 465     GLY B   497
REMARK 465     THR B   498
REMARK 465     LYS B   499
REMARK 465     HIS B   500
REMARK 465     HIS B   501
REMARK 465     HIS B   502
REMARK 465     HIS B   503
REMARK 465     HIS B   504
REMARK 465     HIS B   505
REMARK 465     GLU C    35
REMARK 465     THR C    36
REMARK 465     GLY C    37
REMARK 465     ARG C    38
REMARK 465     THR C    39
REMARK 465     GLU C    40
REMARK 465     ALA C    41
REMARK 465     ALA C    42
REMARK 465     PRO C    43
REMARK 465     ALA C    44
REMARK 465     ALA C    45
REMARK 465     GLY C    46
REMARK 465     GLN C    47
REMARK 465     PRO C    48
REMARK 465     VAL C    49
REMARK 465     GLU C    50
REMARK 465     SER C    51
REMARK 465     PHE C    52
REMARK 465     PRO C    53
REMARK 465     LEU C    54
REMARK 465     ASP C    55
REMARK 465     PHE C    56
REMARK 465     THR C    57
REMARK 465     ALA C    58
REMARK 465     VAL C    59
REMARK 465     GLU C    60
REMARK 465     GLY C    61
REMARK 465     ASN C    62
REMARK 465     MET C    63
REMARK 465     ASP C    64
REMARK 465     SER C    65
REMARK 465     PHE C    66
REMARK 465     MET C    67
REMARK 465     ALA C    68
REMARK 465     GLN C    69
REMARK 465     VAL C    70
REMARK 465     LYS C    71
REMARK 465     SER C    72
REMARK 465     LEU C    73
REMARK 465     ALA C    74
REMARK 465     GLN C    75
REMARK 465     SER C    76
REMARK 465     LEU C    77
REMARK 465     TYR C    78
REMARK 465     PRO C    79
REMARK 465     CYS C    80
REMARK 465     SER C    81
REMARK 465     ALA C    82
REMARK 465     GLN C    83
REMARK 465     GLN C    84
REMARK 465     LEU C    85
REMARK 465     ASN C    86
REMARK 465     LYS C   194
REMARK 465     SER C   195
REMARK 465     GLU C   196
REMARK 465     LYS C   197
REMARK 465     ASN C   198
REMARK 465     VAL C   349
REMARK 465     HIS C   350
REMARK 465     LEU C   351
REMARK 465     THR C   352
REMARK 465     GLY C   353
REMARK 465     GLN C   354
REMARK 465     PRO C   355
REMARK 465     VAL C   356
REMARK 465     GLN C   357
REMARK 465     HIS C   419
REMARK 465     ASP C   420
REMARK 465     SER C   421
REMARK 465     HIS C   422
REMARK 465     LYS C   423
REMARK 465     ALA C   424
REMARK 465     SER C   425
REMARK 465     LYS C   426
REMARK 465     THR C   427
REMARK 465     PRO C   428
REMARK 465     LEU C   429
REMARK 465     LYS C   430
REMARK 465     GLY C   431
REMARK 465     THR C   451
REMARK 465     VAL C   452
REMARK 465     ARG C   453
REMARK 465     ASP C   454
REMARK 465     GLN C   455
REMARK 465     PHE C   456
REMARK 465     THR C   457
REMARK 465     GLY C   458
REMARK 465     GLN C   459
REMARK 465     GLU C   460
REMARK 465     MET C   461
REMARK 465     ASN C   462
REMARK 465     VAL C   463
REMARK 465     ALA C   464
REMARK 465     GLN C   465
REMARK 465     PHE C   466
REMARK 465     LEU C   467
REMARK 465     MET C   468
REMARK 465     HIS C   469
REMARK 465     MET C   470
REMARK 465     GLY C   471
REMARK 465     PHE C   472
REMARK 465     ASP C   473
REMARK 465     MET C   474
REMARK 465     GLN C   475
REMARK 465     THR C   476
REMARK 465     VAL C   477
REMARK 465     ALA C   478
REMARK 465     GLN C   479
REMARK 465     PRO C   480
REMARK 465     GLN C   481
REMARK 465     GLY C   482
REMARK 465     LEU C   483
REMARK 465     GLU C   484
REMARK 465     PRO C   485
REMARK 465     SER C   486
REMARK 465     GLU C   487
REMARK 465     LEU C   488
REMARK 465     LEU C   489
REMARK 465     GLY C   490
REMARK 465     MET C   491
REMARK 465     LEU C   492
REMARK 465     SER C   493
REMARK 465     ASN C   494
REMARK 465     GLY C   495
REMARK 465     SER C   496
REMARK 465     GLY C   497
REMARK 465     THR C   498
REMARK 465     LYS C   499
REMARK 465     HIS C   500
REMARK 465     HIS C   501
REMARK 465     HIS C   502
REMARK 465     HIS C   503
REMARK 465     HIS C   504
REMARK 465     HIS C   505
REMARK 465     GLU D    35
REMARK 465     THR D    36
REMARK 465     GLY D    37
REMARK 465     ARG D    38
REMARK 465     THR D    39
REMARK 465     GLU D    40
REMARK 465     ALA D    41
REMARK 465     ALA D    42
REMARK 465     PRO D    43
REMARK 465     ALA D    44
REMARK 465     ALA D    45
REMARK 465     GLY D    46
REMARK 465     GLN D    47
REMARK 465     PRO D    48
REMARK 465     VAL D    49
REMARK 465     GLU D    50
REMARK 465     SER D    51
REMARK 465     PHE D    52
REMARK 465     PRO D    53
REMARK 465     LEU D    54
REMARK 465     ASP D    55
REMARK 465     PHE D    56
REMARK 465     THR D    57
REMARK 465     ALA D    58
REMARK 465     VAL D    59
REMARK 465     GLU D    60
REMARK 465     GLY D    61
REMARK 465     ASN D    62
REMARK 465     MET D    63
REMARK 465     ASP D    64
REMARK 465     SER D    65
REMARK 465     PHE D    66
REMARK 465     MET D    67
REMARK 465     ALA D    68
REMARK 465     GLN D    69
REMARK 465     VAL D    70
REMARK 465     LYS D    71
REMARK 465     SER D    72
REMARK 465     LEU D    73
REMARK 465     ALA D    74
REMARK 465     GLN D    75
REMARK 465     SER D    76
REMARK 465     LEU D    77
REMARK 465     TYR D    78
REMARK 465     PRO D    79
REMARK 465     CYS D    80
REMARK 465     SER D    81
REMARK 465     ALA D    82
REMARK 465     GLN D    83
REMARK 465     GLN D    84
REMARK 465     LEU D    85
REMARK 465     ASN D    86
REMARK 465     SER D   195
REMARK 465     GLU D   196
REMARK 465     LYS D   197
REMARK 465     HIS D   419
REMARK 465     ASP D   420
REMARK 465     SER D   421
REMARK 465     HIS D   422
REMARK 465     LYS D   423
REMARK 465     ALA D   424
REMARK 465     SER D   425
REMARK 465     LYS D   426
REMARK 465     THR D   427
REMARK 465     PRO D   428
REMARK 465     LEU D   429
REMARK 465     LYS D   430
REMARK 465     GLY D   431
REMARK 465     VAL D   452
REMARK 465     ASP D   473
REMARK 465     MET D   474
REMARK 465     GLN D   475
REMARK 465     THR D   476
REMARK 465     VAL D   477
REMARK 465     ALA D   478
REMARK 465     GLN D   479
REMARK 465     PRO D   480
REMARK 465     GLN D   481
REMARK 465     GLY D   482
REMARK 465     LEU D   483
REMARK 465     GLU D   484
REMARK 465     PRO D   485
REMARK 465     SER D   486
REMARK 465     GLU D   487
REMARK 465     LEU D   488
REMARK 465     LEU D   489
REMARK 465     GLY D   490
REMARK 465     MET D   491
REMARK 465     LEU D   492
REMARK 465     SER D   493
REMARK 465     ASN D   494
REMARK 465     GLY D   495
REMARK 465     SER D   496
REMARK 465     GLY D   497
REMARK 465     THR D   498
REMARK 465     LYS D   499
REMARK 465     HIS D   500
REMARK 465     HIS D   501
REMARK 465     HIS D   502
REMARK 465     HIS D   503
REMARK 465     HIS D   504
REMARK 465     HIS D   505
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     LYS A 112    CG   CD   CE   NZ
REMARK 470     ARG A 118    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS A 194    CG   CD   CE   NZ
REMARK 470     GLU A 199    CG   CD   OE1  OE2
REMARK 470     ASP A 281    CG   OD1  OD2
REMARK 470     GLN A 309    CG   CD   OE1  NE2
REMARK 470     TRP A 442    CG   CD1  CD2  NE1  CE2  CE3  CZ2  CZ3
REMARK 470     TRP A 442    CH2
REMARK 470     HIS A 444    CG   ND1  CD2  CE1  NE2
REMARK 470     GLU B  87    CG   CD   OE1  OE2
REMARK 470     ASP B  88    CG   OD1  OD2
REMARK 470     LYS B 112    CG   CD   CE   NZ
REMARK 470     GLU B 210    CG   CD   OE1  OE2
REMARK 470     ARG B 213    CG   CD   NE   CZ   NH1  NH2
REMARK 470     ASP B 281    CG   OD1  OD2
REMARK 470     ILE B 283    CG1  CG2  CD1
REMARK 470     ARG B 305    CG   CD   NE   CZ   NH1  NH2
REMARK 470     ARG B 361    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LEU B 362    CG   CD1  CD2
REMARK 470     ARG B 394    CG   CD   NE   CZ   NH1  NH2
REMARK 470     ASP B 415    CG   OD1  OD2
REMARK 470     HIS B 419    CG   ND1  CD2  CE1  NE2
REMARK 470     HIS B 444    CG   ND1  CD2  CE1  NE2
REMARK 470     GLU C  87    CG   CD   OE1  OE2
REMARK 470     GLU C 199    CG   CD   OE1  OE2
REMARK 470     ASP C 270    CG   OD1  OD2
REMARK 470     ASP C 281    CG   OD1  OD2
REMARK 470     GLU C 358    CG   CD   OE1  OE2
REMARK 470     GLN C 365    CG   CD   OE1  NE2
REMARK 470     THR C 405    OG1  CG2
REMARK 470     HIS C 444    CG   ND1  CD2  CE1  NE2
REMARK 470     ASP D  88    CG   OD1  OD2
REMARK 470     ASP D 281    CG   OD1  OD2
REMARK 470     GLU D 312    CG   CD   OE1  OE2
REMARK 470     GLN D 354    CG   CD   OE1  NE2
REMARK 470     ARG D 372    CG   CD   NE   CZ   NH1  NH2
REMARK 470     LYS D 403    CG   CD   CE   NZ
REMARK 470     LEU D 418    CG   CD1  CD2
REMARK 470     TRP D 442    CG   CD1  CD2  NE1  CE2  CE3  CZ2  CZ3
REMARK 470     TRP D 442    CH2
REMARK 470     THR D 451    OG1  CG2
REMARK 470     ARG D 453    CG   CD   NE   CZ   NH1  NH2
REMARK 470     ASP D 454    CG   OD1  OD2
REMARK 470     GLN D 455    CG   CD   OE1  NE2
REMARK 470     PHE D 456    CG   CD1  CD2  CE1  CE2  CZ
REMARK 470     THR D 457    OG1  CG2
REMARK 470     GLN D 459    CG   CD   OE1  NE2
REMARK 470     GLU D 460    CG   CD   OE1  OE2
REMARK 470     ASN D 462    CG   OD1  ND2
REMARK 470     GLN D 465    CG   CD   OE1  NE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS IN SAME ASYMMETRIC UNIT
REMARK 500
REMARK 500 THE FOLLOWING ATOMS ARE IN CLOSE CONTACT.
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI           DISTANCE
REMARK 500   OG   SER A   184     OD1  ASP A   186              1.95
REMARK 500   NH1  ARG B   139     OG   SER B   185              1.98
REMARK 500   OG   SER B   184     OD1  ASP B   186              2.10
REMARK 500   O    ASP B   186     OG   SER B   189              1.95
REMARK 500   OD2  ASP B   186     N    GLY B   190              2.16
REMARK 500   O    LYS B   272     OH   TYR B   363              1.98
REMARK 500   NH2  ARG B   416     SG   CYS B   432              2.18
REMARK 500   CG   ASN C   132     OD1  ASN C   135              1.91
REMARK 500   OD1  ASN C   132     OD1  ASN C   135              1.94
REMARK 500   ND2  ASN C   132     OD1  ASN C   135              1.82
REMARK 500   NH1  ARG C   139     OG   SER C   185              1.37
REMARK 500   O    ASP C   186     OG   SER C   189              1.94
REMARK 500   OE2  GLU C   210     NE   ARG C   213              2.15
REMARK 500   OE2  GLU C   210     CZ   ARG C   213              2.10
REMARK 500   OE2  GLU C   210     NH2  ARG C   213              1.81
REMARK 500   ND2  ASN C   271     OD1  ASP C   347              2.14
REMARK 500   ND2  ASN C   271     OD2  ASP C   347              1.91
REMARK 500   O    LYS C   272     OH   TYR C   363              2.18
REMARK 500   O    ASP C   281     OG1  THR C   284              2.04
REMARK 500   O    ASP D   186     OG   SER D   189              1.98
REMARK 500   OE2  GLU D   210     NE   ARG D   213              1.77
REMARK 500   OD1  ASN D   317     NZ   LYS D   323              1.55
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    CYS A 130   CA  -  CB  -  SG  ANGL. DEV. =  -8.1 DEGREES
REMARK 500    CYS A 183   CA  -  CB  -  SG  ANGL. DEV. =  -9.4 DEGREES
REMARK 500    CYS C 183   CA  -  CB  -  SG  ANGL. DEV. =  -7.3 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    TRP A 128     -148.36     58.56
REMARK 500    SER A 232     -125.04     63.68
REMARK 500    ALA A 385       58.33    -91.43
REMARK 500    GLU A 390      132.15     83.72
REMARK 500    ILE A 391      -33.85   -139.31
REMARK 500    VAL A 402      -77.38   -106.80
REMARK 500    LYS A 403       59.04   -103.11
REMARK 500    HIS A 435       79.29   -100.17
REMARK 500    SER A 439       68.93   -108.94
REMARK 500    PRO A 441        0.27    -67.92
REMARK 500    TRP B 128     -148.53     64.04
REMARK 500    ILE B 161       -5.17    -57.71
REMARK 500    ASN B 169       68.15   -163.26
REMARK 500    SER B 232     -128.07     59.39
REMARK 500    CYS B 318        3.70    -69.35
REMARK 500    GLU B 390      146.78     79.74
REMARK 500    ILE B 391      -49.57   -145.09
REMARK 500    ASN C  96       96.04    -68.12
REMARK 500    TRP C 128     -156.89     68.85
REMARK 500    SER C 232     -122.55     57.51
REMARK 500    GLU C 390      156.97     78.69
REMARK 500    ILE C 391      -58.29   -147.71
REMARK 500    HIS C 396        7.98    -67.84
REMARK 500    VAL C 402      -63.35    -99.40
REMARK 500    CYS C 440       87.68   -160.64
REMARK 500    PRO C 441       40.04    -71.63
REMARK 500    HIS C 444        5.75     58.76
REMARK 500    ASN D  96       80.21    -67.45
REMARK 500    TRP D 128     -157.10     65.12
REMARK 500    ALA D 191       18.94   -143.36
REMARK 500    SER D 232     -126.95     66.97
REMARK 500    LEU D 252       36.76    -75.27
REMARK 500    CYS D 318        3.35    -61.03
REMARK 500    GLU D 390      130.02     72.37
REMARK 500    ILE D 391      -46.13   -134.07
REMARK 500    VAL D 402      -70.94   -105.51
REMARK 500    LYS D 403       55.86   -111.96
REMARK 500    HIS D 435       79.83   -110.84
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL B1453
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800  NAG B1452  BOUND TO ASN B  96
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4UYU   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  I IODIDE COMPLEX - 2.3A
REMARK 900 RELATED ID: 4UYW   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   HEPARIN FRAGMENT COMPLEX - 1.7A
REMARK 900 RELATED ID: 4UZ1   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  III - 1.4A
REMARK 900 RELATED ID: 4UZ5   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM IV
REMARK 900   - 2.1A
REMARK 900 RELATED ID: 4UZ6   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM V
REMARK 900   - SOS COMPLEX - 1.9A
REMARK 900 RELATED ID: 4UZ7   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  VII - SOS COMPLEX - 2.2A
REMARK 900 RELATED ID: 4UZ9   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  VII - SOS COMPLEX - 2.2A
REMARK 900 RELATED ID: 4UZA   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  VIII - PHOSPHATE COMPLEX - 2.4A
REMARK 900 RELATED ID: 4UZJ   RELATED DB: PDB
REMARK 900  STRUCTURE OF A WNT SIGNAL REGULATOR FROM DROSOPHILA -
REMARK 900  CRYSTAL FORM I - 2.4A
REMARK 900 RELATED ID: 4UZK   RELATED DB: PDB
REMARK 900  STRUCTURE OF A WNT SIGNAL REGULATOR FROM DROSOPHILA -
REMARK 900  CRYSTAL FORM II - 1.9A
REMARK 900 RELATED ID: 4UZL   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   MYRISTOLEATE COMPLEX - 2.1A
REMARK 900 RELATED ID: 4UZQ   RELATED DB: PDB
REMARK 900  STRUCTURE OF A WNT SIGNAL REGULATOR COMPLEX - CRYSTAL
REMARK 900  FORM IX - 1.5A
DBREF  4UYZ A   38   496  UNP    Q6P988   NOTUM_HUMAN     38    496
DBREF  4UYZ B   38   496  UNP    Q6P988   NOTUM_HUMAN     38    496
DBREF  4UYZ C   38   496  UNP    Q6P988   NOTUM_HUMAN     38    496
DBREF  4UYZ D   38   496  UNP    Q6P988   NOTUM_HUMAN     38    496
DBREF  4UYZ E    7    16  PDB    4UYZ     4UYZ             7     16
SEQADV 4UYZ GLU A   35  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR A   36  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY A   37  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY A  497  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR A  498  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ LYS A  499  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS A  500  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS A  501  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS A  502  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS A  503  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS A  504  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS A  505  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLU B   35  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR B   36  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY B   37  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY B  497  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR B  498  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ LYS B  499  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS B  500  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS B  501  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS B  502  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS B  503  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS B  504  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS B  505  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLU C   35  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR C   36  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY C   37  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY C  497  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR C  498  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ LYS C  499  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS C  500  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS C  501  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS C  502  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS C  503  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS C  504  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS C  505  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLU D   35  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR D   36  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY D   37  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ GLY D  497  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ THR D  498  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ LYS D  499  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS D  500  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS D  501  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS D  502  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS D  503  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS D  504  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UYZ HIS D  505  UNP  Q6P988              EXPRESSION TAG
SEQRES   1 A  471  GLU THR GLY ARG THR GLU ALA ALA PRO ALA ALA GLY GLN
SEQRES   2 A  471  PRO VAL GLU SER PHE PRO LEU ASP PHE THR ALA VAL GLU
SEQRES   3 A  471  GLY ASN MET ASP SER PHE MET ALA GLN VAL LYS SER LEU
SEQRES   4 A  471  ALA GLN SER LEU TYR PRO CYS SER ALA GLN GLN LEU ASN
SEQRES   5 A  471  GLU ASP LEU ARG LEU HIS LEU LEU LEU ASN THR SER VAL
SEQRES   6 A  471  THR CYS ASN ASP GLY SER PRO ALA GLY TYR TYR LEU LYS
SEQRES   7 A  471  GLU SER ARG GLY SER ARG ARG TRP LEU LEU PHE LEU GLU
SEQRES   8 A  471  GLY GLY TRP TYR CYS PHE ASN ARG GLU ASN CYS ASP SER
SEQRES   9 A  471  ARG TYR ASP THR MET ARG ARG LEU MET SER SER ARG ASP
SEQRES  10 A  471  TRP PRO ARG THR ARG THR GLY THR GLY ILE LEU SER SER
SEQRES  11 A  471  GLN PRO GLU GLU ASN PRO TYR TRP TRP ASN ALA ASN MET
SEQRES  12 A  471  VAL PHE ILE PRO TYR CYS SER SER ASP VAL TRP SER GLY
SEQRES  13 A  471  ALA SER SER LYS SER GLU LYS ASN GLU TYR ALA PHE MET
SEQRES  14 A  471  GLY ALA LEU ILE ILE GLN GLU VAL VAL ARG GLU LEU LEU
SEQRES  15 A  471  GLY ARG GLY LEU SER GLY ALA LYS VAL LEU LEU LEU ALA
SEQRES  16 A  471  GLY SER SER ALA GLY GLY THR GLY VAL LEU LEU ASN VAL
SEQRES  17 A  471  ASP ARG VAL ALA GLU GLN LEU GLU LYS LEU GLY TYR PRO
SEQRES  18 A  471  ALA ILE GLN VAL ARG GLY LEU ALA ASP SER GLY TRP PHE
SEQRES  19 A  471  LEU ASP ASN LYS GLN TYR ARG HIS THR ASP CYS VAL ASP
SEQRES  20 A  471  THR ILE THR CYS ALA PRO THR GLU ALA ILE ARG ARG GLY
SEQRES  21 A  471  ILE ARG TYR TRP ASN GLY VAL VAL PRO GLU ARG CYS ARG
SEQRES  22 A  471  ARG GLN PHE GLN GLU GLY GLU GLU TRP ASN CYS PHE PHE
SEQRES  23 A  471  GLY TYR LYS VAL TYR PRO THR LEU ARG CYS PRO VAL PHE
SEQRES  24 A  471  VAL VAL GLN TRP LEU PHE ASP GLU ALA GLN LEU THR VAL
SEQRES  25 A  471  ASP ASN VAL HIS LEU THR GLY GLN PRO VAL GLN GLU GLY
SEQRES  26 A  471  LEU ARG LEU TYR ILE GLN ASN LEU GLY ARG GLU LEU ARG
SEQRES  27 A  471  HIS THR LEU LYS ASP VAL PRO ALA SER PHE ALA PRO ALA
SEQRES  28 A  471  CYS LEU SER HIS GLU ILE ILE ILE ARG SER HIS TRP THR
SEQRES  29 A  471  ASP VAL GLN VAL LYS GLY THR SER LEU PRO ARG ALA LEU
SEQRES  30 A  471  HIS CYS TRP ASP ARG SER LEU HIS ASP SER HIS LYS ALA
SEQRES  31 A  471  SER LYS THR PRO LEU LYS GLY CYS PRO VAL HIS LEU VAL
SEQRES  32 A  471  ASP SER CYS PRO TRP PRO HIS CYS ASN PRO SER CYS PRO
SEQRES  33 A  471  THR VAL ARG ASP GLN PHE THR GLY GLN GLU MET ASN VAL
SEQRES  34 A  471  ALA GLN PHE LEU MET HIS MET GLY PHE ASP MET GLN THR
SEQRES  35 A  471  VAL ALA GLN PRO GLN GLY LEU GLU PRO SER GLU LEU LEU
SEQRES  36 A  471  GLY MET LEU SER ASN GLY SER GLY THR LYS HIS HIS HIS
SEQRES  37 A  471  HIS HIS HIS
SEQRES   1 B  471  GLU THR GLY ARG THR GLU ALA ALA PRO ALA ALA GLY GLN
SEQRES   2 B  471  PRO VAL GLU SER PHE PRO LEU ASP PHE THR ALA VAL GLU
SEQRES   3 B  471  GLY ASN MET ASP SER PHE MET ALA GLN VAL LYS SER LEU
SEQRES   4 B  471  ALA GLN SER LEU TYR PRO CYS SER ALA GLN GLN LEU ASN
SEQRES   5 B  471  GLU ASP LEU ARG LEU HIS LEU LEU LEU ASN THR SER VAL
SEQRES   6 B  471  THR CYS ASN ASP GLY SER PRO ALA GLY TYR TYR LEU LYS
SEQRES   7 B  471  GLU SER ARG GLY SER ARG ARG TRP LEU LEU PHE LEU GLU
SEQRES   8 B  471  GLY GLY TRP TYR CYS PHE ASN ARG GLU ASN CYS ASP SER
SEQRES   9 B  471  ARG TYR ASP THR MET ARG ARG LEU MET SER SER ARG ASP
SEQRES  10 B  471  TRP PRO ARG THR ARG THR GLY THR GLY ILE LEU SER SER
SEQRES  11 B  471  GLN PRO GLU GLU ASN PRO TYR TRP TRP ASN ALA ASN MET
SEQRES  12 B  471  VAL PHE ILE PRO TYR CYS SER SER ASP VAL TRP SER GLY
SEQRES  13 B  471  ALA SER SER LYS SER GLU LYS ASN GLU TYR ALA PHE MET
SEQRES  14 B  471  GLY ALA LEU ILE ILE GLN GLU VAL VAL ARG GLU LEU LEU
SEQRES  15 B  471  GLY ARG GLY LEU SER GLY ALA LYS VAL LEU LEU LEU ALA
SEQRES  16 B  471  GLY SER SER ALA GLY GLY THR GLY VAL LEU LEU ASN VAL
SEQRES  17 B  471  ASP ARG VAL ALA GLU GLN LEU GLU LYS LEU GLY TYR PRO
SEQRES  18 B  471  ALA ILE GLN VAL ARG GLY LEU ALA ASP SER GLY TRP PHE
SEQRES  19 B  471  LEU ASP ASN LYS GLN TYR ARG HIS THR ASP CYS VAL ASP
SEQRES  20 B  471  THR ILE THR CYS ALA PRO THR GLU ALA ILE ARG ARG GLY
SEQRES  21 B  471  ILE ARG TYR TRP ASN GLY VAL VAL PRO GLU ARG CYS ARG
SEQRES  22 B  471  ARG GLN PHE GLN GLU GLY GLU GLU TRP ASN CYS PHE PHE
SEQRES  23 B  471  GLY TYR LYS VAL TYR PRO THR LEU ARG CYS PRO VAL PHE
SEQRES  24 B  471  VAL VAL GLN TRP LEU PHE ASP GLU ALA GLN LEU THR VAL
SEQRES  25 B  471  ASP ASN VAL HIS LEU THR GLY GLN PRO VAL GLN GLU GLY
SEQRES  26 B  471  LEU ARG LEU TYR ILE GLN ASN LEU GLY ARG GLU LEU ARG
SEQRES  27 B  471  HIS THR LEU LYS ASP VAL PRO ALA SER PHE ALA PRO ALA
SEQRES  28 B  471  CYS LEU SER HIS GLU ILE ILE ILE ARG SER HIS TRP THR
SEQRES  29 B  471  ASP VAL GLN VAL LYS GLY THR SER LEU PRO ARG ALA LEU
SEQRES  30 B  471  HIS CYS TRP ASP ARG SER LEU HIS ASP SER HIS LYS ALA
SEQRES  31 B  471  SER LYS THR PRO LEU LYS GLY CYS PRO VAL HIS LEU VAL
SEQRES  32 B  471  ASP SER CYS PRO TRP PRO HIS CYS ASN PRO SER CYS PRO
SEQRES  33 B  471  THR VAL ARG ASP GLN PHE THR GLY GLN GLU MET ASN VAL
SEQRES  34 B  471  ALA GLN PHE LEU MET HIS MET GLY PHE ASP MET GLN THR
SEQRES  35 B  471  VAL ALA GLN PRO GLN GLY LEU GLU PRO SER GLU LEU LEU
SEQRES  36 B  471  GLY MET LEU SER ASN GLY SER GLY THR LYS HIS HIS HIS
SEQRES  37 B  471  HIS HIS HIS
SEQRES   1 C  471  GLU THR GLY ARG THR GLU ALA ALA PRO ALA ALA GLY GLN
SEQRES   2 C  471  PRO VAL GLU SER PHE PRO LEU ASP PHE THR ALA VAL GLU
SEQRES   3 C  471  GLY ASN MET ASP SER PHE MET ALA GLN VAL LYS SER LEU
SEQRES   4 C  471  ALA GLN SER LEU TYR PRO CYS SER ALA GLN GLN LEU ASN
SEQRES   5 C  471  GLU ASP LEU ARG LEU HIS LEU LEU LEU ASN THR SER VAL
SEQRES   6 C  471  THR CYS ASN ASP GLY SER PRO ALA GLY TYR TYR LEU LYS
SEQRES   7 C  471  GLU SER ARG GLY SER ARG ARG TRP LEU LEU PHE LEU GLU
SEQRES   8 C  471  GLY GLY TRP TYR CYS PHE ASN ARG GLU ASN CYS ASP SER
SEQRES   9 C  471  ARG TYR ASP THR MET ARG ARG LEU MET SER SER ARG ASP
SEQRES  10 C  471  TRP PRO ARG THR ARG THR GLY THR GLY ILE LEU SER SER
SEQRES  11 C  471  GLN PRO GLU GLU ASN PRO TYR TRP TRP ASN ALA ASN MET
SEQRES  12 C  471  VAL PHE ILE PRO TYR CYS SER SER ASP VAL TRP SER GLY
SEQRES  13 C  471  ALA SER SER LYS SER GLU LYS ASN GLU TYR ALA PHE MET
SEQRES  14 C  471  GLY ALA LEU ILE ILE GLN GLU VAL VAL ARG GLU LEU LEU
SEQRES  15 C  471  GLY ARG GLY LEU SER GLY ALA LYS VAL LEU LEU LEU ALA
SEQRES  16 C  471  GLY SER SER ALA GLY GLY THR GLY VAL LEU LEU ASN VAL
SEQRES  17 C  471  ASP ARG VAL ALA GLU GLN LEU GLU LYS LEU GLY TYR PRO
SEQRES  18 C  471  ALA ILE GLN VAL ARG GLY LEU ALA ASP SER GLY TRP PHE
SEQRES  19 C  471  LEU ASP ASN LYS GLN TYR ARG HIS THR ASP CYS VAL ASP
SEQRES  20 C  471  THR ILE THR CYS ALA PRO THR GLU ALA ILE ARG ARG GLY
SEQRES  21 C  471  ILE ARG TYR TRP ASN GLY VAL VAL PRO GLU ARG CYS ARG
SEQRES  22 C  471  ARG GLN PHE GLN GLU GLY GLU GLU TRP ASN CYS PHE PHE
SEQRES  23 C  471  GLY TYR LYS VAL TYR PRO THR LEU ARG CYS PRO VAL PHE
SEQRES  24 C  471  VAL VAL GLN TRP LEU PHE ASP GLU ALA GLN LEU THR VAL
SEQRES  25 C  471  ASP ASN VAL HIS LEU THR GLY GLN PRO VAL GLN GLU GLY
SEQRES  26 C  471  LEU ARG LEU TYR ILE GLN ASN LEU GLY ARG GLU LEU ARG
SEQRES  27 C  471  HIS THR LEU LYS ASP VAL PRO ALA SER PHE ALA PRO ALA
SEQRES  28 C  471  CYS LEU SER HIS GLU ILE ILE ILE ARG SER HIS TRP THR
SEQRES  29 C  471  ASP VAL GLN VAL LYS GLY THR SER LEU PRO ARG ALA LEU
SEQRES  30 C  471  HIS CYS TRP ASP ARG SER LEU HIS ASP SER HIS LYS ALA
SEQRES  31 C  471  SER LYS THR PRO LEU LYS GLY CYS PRO VAL HIS LEU VAL
SEQRES  32 C  471  ASP SER CYS PRO TRP PRO HIS CYS ASN PRO SER CYS PRO
SEQRES  33 C  471  THR VAL ARG ASP GLN PHE THR GLY GLN GLU MET ASN VAL
SEQRES  34 C  471  ALA GLN PHE LEU MET HIS MET GLY PHE ASP MET GLN THR
SEQRES  35 C  471  VAL ALA GLN PRO GLN GLY LEU GLU PRO SER GLU LEU LEU
SEQRES  36 C  471  GLY MET LEU SER ASN GLY SER GLY THR LYS HIS HIS HIS
SEQRES  37 C  471  HIS HIS HIS
SEQRES   1 D  471  GLU THR GLY ARG THR GLU ALA ALA PRO ALA ALA GLY GLN
SEQRES   2 D  471  PRO VAL GLU SER PHE PRO LEU ASP PHE THR ALA VAL GLU
SEQRES   3 D  471  GLY ASN MET ASP SER PHE MET ALA GLN VAL LYS SER LEU
SEQRES   4 D  471  ALA GLN SER LEU TYR PRO CYS SER ALA GLN GLN LEU ASN
SEQRES   5 D  471  GLU ASP LEU ARG LEU HIS LEU LEU LEU ASN THR SER VAL
SEQRES   6 D  471  THR CYS ASN ASP GLY SER PRO ALA GLY TYR TYR LEU LYS
SEQRES   7 D  471  GLU SER ARG GLY SER ARG ARG TRP LEU LEU PHE LEU GLU
SEQRES   8 D  471  GLY GLY TRP TYR CYS PHE ASN ARG GLU ASN CYS ASP SER
SEQRES   9 D  471  ARG TYR ASP THR MET ARG ARG LEU MET SER SER ARG ASP
SEQRES  10 D  471  TRP PRO ARG THR ARG THR GLY THR GLY ILE LEU SER SER
SEQRES  11 D  471  GLN PRO GLU GLU ASN PRO TYR TRP TRP ASN ALA ASN MET
SEQRES  12 D  471  VAL PHE ILE PRO TYR CYS SER SER ASP VAL TRP SER GLY
SEQRES  13 D  471  ALA SER SER LYS SER GLU LYS ASN GLU TYR ALA PHE MET
SEQRES  14 D  471  GLY ALA LEU ILE ILE GLN GLU VAL VAL ARG GLU LEU LEU
SEQRES  15 D  471  GLY ARG GLY LEU SER GLY ALA LYS VAL LEU LEU LEU ALA
SEQRES  16 D  471  GLY SER SER ALA GLY GLY THR GLY VAL LEU LEU ASN VAL
SEQRES  17 D  471  ASP ARG VAL ALA GLU GLN LEU GLU LYS LEU GLY TYR PRO
SEQRES  18 D  471  ALA ILE GLN VAL ARG GLY LEU ALA ASP SER GLY TRP PHE
SEQRES  19 D  471  LEU ASP ASN LYS GLN TYR ARG HIS THR ASP CYS VAL ASP
SEQRES  20 D  471  THR ILE THR CYS ALA PRO THR GLU ALA ILE ARG ARG GLY
SEQRES  21 D  471  ILE ARG TYR TRP ASN GLY VAL VAL PRO GLU ARG CYS ARG
SEQRES  22 D  471  ARG GLN PHE GLN GLU GLY GLU GLU TRP ASN CYS PHE PHE
SEQRES  23 D  471  GLY TYR LYS VAL TYR PRO THR LEU ARG CYS PRO VAL PHE
SEQRES  24 D  471  VAL VAL GLN TRP LEU PHE ASP GLU ALA GLN LEU THR VAL
SEQRES  25 D  471  ASP ASN VAL HIS LEU THR GLY GLN PRO VAL GLN GLU GLY
SEQRES  26 D  471  LEU ARG LEU TYR ILE GLN ASN LEU GLY ARG GLU LEU ARG
SEQRES  27 D  471  HIS THR LEU LYS ASP VAL PRO ALA SER PHE ALA PRO ALA
SEQRES  28 D  471  CYS LEU SER HIS GLU ILE ILE ILE ARG SER HIS TRP THR
SEQRES  29 D  471  ASP VAL GLN VAL LYS GLY THR SER LEU PRO ARG ALA LEU
SEQRES  30 D  471  HIS CYS TRP ASP ARG SER LEU HIS ASP SER HIS LYS ALA
SEQRES  31 D  471  SER LYS THR PRO LEU LYS GLY CYS PRO VAL HIS LEU VAL
SEQRES  32 D  471  ASP SER CYS PRO TRP PRO HIS CYS ASN PRO SER CYS PRO
SEQRES  33 D  471  THR VAL ARG ASP GLN PHE THR GLY GLN GLU MET ASN VAL
SEQRES  34 D  471  ALA GLN PHE LEU MET HIS MET GLY PHE ASP MET GLN THR
SEQRES  35 D  471  VAL ALA GLN PRO GLN GLY LEU GLU PRO SER GLU LEU LEU
SEQRES  36 D  471  GLY MET LEU SER ASN GLY SER GLY THR LYS HIS HIS HIS
SEQRES  37 D  471  HIS HIS HIS
SEQRES   1 E   10  ALA ALA ALA ALA ALA ALA ALA ALA ALA ALA
HET    NAG  B1452      14
HET     CL  B1453       1
HETNAM      CL CHLORIDE ION
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   6   CL    CL 1-
FORMUL   7  NAG    C8 H15 N O6
HELIX    1   1 ASN A  132  MET A  143  1                                  12
HELIX    2   2 ARG A  144  MET A  147  5                                   4
HELIX    3   3 THR A  159  SER A  163  5                                   5
HELIX    4   4 MET A  203  GLY A  217  1                                  15
HELIX    5   5 ARG A  218  ALA A  223  5                                   6
HELIX    6   6 SER A  232  GLY A  253  1                                  22
HELIX    7   7 ALA A  286  ASN A  299  1                                  14
HELIX    8   8 PRO A  303  ARG A  308  1                                   6
HELIX    9   9 GLU A  314  PHE A  319  5                                   6
HELIX   10  10 PHE A  320  TYR A  325  1                                   6
HELIX   11  11 PRO A  326  LEU A  328  5                                   3
HELIX   12  12 ASP A  340  ASP A  347  1                                   8
HELIX   13  13 GLN A  357  LEU A  375  1                                  19
HELIX   14  14 ARG A  394  ASP A  399  5                                   6
HELIX   15  15 SER A  406  LEU A  418  1                                  13
HELIX   16  16 ASN B  132  MET B  143  1                                  12
HELIX   17  17 ARG B  144  MET B  147  5                                   4
HELIX   18  18 THR B  159  SER B  163  5                                   5
HELIX   19  19 MET B  203  ARG B  218  1                                  16
HELIX   20  20 GLY B  219  ALA B  223  5                                   5
HELIX   21  21 SER B  232  LEU B  252  1                                  21
HELIX   22  22 ALA B  286  ASN B  299  1                                  14
HELIX   23  23 PRO B  303  PHE B  310  1                                   8
HELIX   24  24 GLU B  314  PHE B  319  5                                   6
HELIX   25  25 PHE B  320  TYR B  325  1                                   6
HELIX   26  26 PRO B  326  LEU B  328  5                                   3
HELIX   27  27 ASP B  340  ASP B  347  1                                   8
HELIX   28  28 ARG B  361  LEU B  375  1                                  15
HELIX   29  29 ARG B  394  ASP B  399  5                                   6
HELIX   30  30 SER B  406  HIS B  419  1                                  14
HELIX   31  31 ASN C  132  MET C  143  1                                  12
HELIX   32  32 ARG C  144  MET C  147  5                                   4
HELIX   33  33 THR C  159  SER C  163  5                                   5
HELIX   34  34 MET C  203  GLY C  217  1                                  15
HELIX   35  35 SER C  232  GLY C  253  1                                  22
HELIX   36  36 ALA C  286  ASN C  299  1                                  14
HELIX   37  37 PRO C  303  ARG C  308  1                                   6
HELIX   38  38 GLU C  314  PHE C  319  5                                   6
HELIX   39  39 PHE C  320  TYR C  325  1                                   6
HELIX   40  40 PRO C  326  LEU C  328  5                                   3
HELIX   41  41 ASP C  340  ASN C  348  1                                   9
HELIX   42  42 GLU C  358  LYS C  376  1                                  19
HELIX   43  43 ARG C  394  ASP C  399  5                                   6
HELIX   44  44 SER C  406  LEU C  418  1                                  13
HELIX   45  45 ASN D  132  MET D  143  1                                  12
HELIX   46  46 ARG D  144  MET D  147  5                                   4
HELIX   47  47 THR D  159  SER D  163  5                                   5
HELIX   48  48 MET D  203  ARG D  218  1                                  16
HELIX   49  49 GLY D  219  ALA D  223  5                                   5
HELIX   50  50 SER D  232  LEU D  252  1                                  21
HELIX   51  51 ALA D  286  ASN D  299  1                                  14
HELIX   52  52 PRO D  303  ARG D  308  1                                   6
HELIX   53  53 GLU D  314  PHE D  319  5                                   6
HELIX   54  54 PHE D  320  TYR D  325  1                                   6
HELIX   55  55 GLU D  341  ASP D  347  1                                   7
HELIX   56  56 GLN D  357  LEU D  375  1                                  19
HELIX   57  57 ARG D  394  VAL D  400  5                                   7
HELIX   58  58 SER D  406  LEU D  418  1                                  13
HELIX   59  59 ASN D  462  MET D  470  1                                   9
HELIX   60  60 ALA E    9  ALA E   16  1                                   8
SHEET    1  AA10 THR A 155  ARG A 156  0
SHEET    2  AA10 LEU A  89  LEU A  93 -1  O  LEU A  89   N  ARG A 156
SHEET    3  AA10 GLY A 108  LYS A 112 -1  O  TYR A 109   N  HIS A  92
SHEET    4  AA10 ASN A 176  ILE A 180 -1  O  MET A 177   N  LYS A 112
SHEET    5  AA10 ARG A 119  LEU A 124  1  O  ARG A 119   N  ASN A 176
SHEET    6  AA10 VAL A 225  SER A 231  1  O  VAL A 225   N  TRP A 120
SHEET    7  AA10 GLN A 258  ASP A 264  1  O  GLN A 258   N  LEU A 226
SHEET    8  AA10 VAL A 332  VAL A 335  1  O  PHE A 333   N  ALA A 263
SHEET    9  AA10 SER A 381  ALA A 383  1  N  PHE A 382   O  VAL A 334
SHEET   10  AA10 HIS A 435  VAL A 437  1  O  LEU A 436   N  ALA A 383
SHEET    1  BA10 THR B 155  THR B 157  0
SHEET    2  BA10 ASP B  88  LEU B  93 -1  O  LEU B  89   N  ARG B 156
SHEET    3  BA10 GLY B 108  LYS B 112 -1  O  TYR B 109   N  HIS B  92
SHEET    4  BA10 ASN B 176  ILE B 180 -1  O  MET B 177   N  LYS B 112
SHEET    5  BA10 ARG B 119  LEU B 124  1  O  ARG B 119   N  ASN B 176
SHEET    6  BA10 VAL B 225  SER B 231  1  O  VAL B 225   N  TRP B 120
SHEET    7  BA10 GLN B 258  ASP B 264  1  O  GLN B 258   N  LEU B 226
SHEET    8  BA10 VAL B 332  VAL B 335  1  O  PHE B 333   N  ALA B 263
SHEET    9  BA10 SER B 381  ALA B 383  1  N  PHE B 382   O  VAL B 334
SHEET   10  BA10 HIS B 435  VAL B 437  1  O  LEU B 436   N  ALA B 383
SHEET    1  CA10 THR C 155  ARG C 156  0
SHEET    2  CA10 LEU C  89  LEU C  93 -1  O  LEU C  89   N  ARG C 156
SHEET    3  CA10 GLY C 108  LYS C 112 -1  O  TYR C 109   N  HIS C  92
SHEET    4  CA10 ASN C 176  ILE C 180 -1  O  MET C 177   N  LYS C 112
SHEET    5  CA10 ARG C 119  LEU C 124  1  O  ARG C 119   N  ASN C 176
SHEET    6  CA10 VAL C 225  SER C 231  1  O  VAL C 225   N  TRP C 120
SHEET    7  CA10 GLN C 258  ASP C 264  1  O  GLN C 258   N  LEU C 226
SHEET    8  CA10 VAL C 332  VAL C 335  1  O  PHE C 333   N  ALA C 263
SHEET    9  CA10 SER C 381  ALA C 383  1  N  PHE C 382   O  VAL C 334
SHEET   10  CA10 HIS C 435  VAL C 437  1  O  LEU C 436   N  ALA C 383
SHEET    1  DA10 THR D 155  ARG D 156  0
SHEET    2  DA10 LEU D  89  LEU D  93 -1  O  LEU D  89   N  ARG D 156
SHEET    3  DA10 GLY D 108  LYS D 112 -1  O  TYR D 109   N  HIS D  92
SHEET    4  DA10 ASN D 176  ILE D 180 -1  O  MET D 177   N  LYS D 112
SHEET    5  DA10 ARG D 119  LEU D 124  1  O  ARG D 119   N  ASN D 176
SHEET    6  DA10 VAL D 225  SER D 231  1  O  VAL D 225   N  TRP D 120
SHEET    7  DA10 GLN D 258  ASP D 264  1  O  GLN D 258   N  LEU D 226
SHEET    8  DA10 VAL D 332  VAL D 335  1  O  PHE D 333   N  ALA D 263
SHEET    9  DA10 SER D 381  ALA D 383  1  N  PHE D 382   O  VAL D 334
SHEET   10  DA10 HIS D 435  VAL D 437  1  O  LEU D 436   N  ALA D 383
SHEET    1  DB 2 PHE D 339  ASP D 340  0
SHEET    2  DB 2 LEU D 387  SER D 388
SSBOND   1 CYS A  101    CYS A  183                          1555   1555  2.05
SSBOND   2 CYS A  130    CYS A  136                          1555   1555  2.04
SSBOND   3 CYS A  279    CYS A  285                          1555   1555  2.03
SSBOND   4 CYS A  306    CYS A  318                          1555   1555  2.04
SSBOND   5 CYS A  386    CYS A  449                          1555   1555  2.04
SSBOND   6 CYS A  413    CYS A  432                          1555   1555  2.02
SSBOND   7 CYS A  440    CYS A  445                          1555   1555  2.04
SSBOND   8 CYS B  101    CYS B  183                          1555   1555  2.06
SSBOND   9 CYS B  130    CYS B  136                          1555   1555  2.04
SSBOND  10 CYS B  279    CYS B  285                          1555   1555  2.03
SSBOND  11 CYS B  306    CYS B  318                          1555   1555  2.05
SSBOND  12 CYS B  386    CYS B  449                          1555   1555  2.03
SSBOND  13 CYS B  413    CYS B  432                          1555   1555  2.03
SSBOND  14 CYS B  440    CYS B  445                          1555   1555  2.03
SSBOND  15 CYS C  101    CYS C  183                          1555   1555  2.03
SSBOND  16 CYS C  130    CYS C  136                          1555   1555  2.02
SSBOND  17 CYS C  279    CYS C  285                          1555   1555  2.02
SSBOND  18 CYS C  306    CYS C  318                          1555   1555  2.07
SSBOND  19 CYS C  386    CYS C  449                          1555   1555  2.03
SSBOND  20 CYS C  413    CYS C  432                          1555   1555  2.02
SSBOND  21 CYS C  440    CYS C  445                          1555   1555  2.03
SSBOND  22 CYS D  101    CYS D  183                          1555   1555  2.04
SSBOND  23 CYS D  130    CYS D  136                          1555   1555  2.05
SSBOND  24 CYS D  279    CYS D  285                          1555   1555  2.03
SSBOND  25 CYS D  306    CYS D  318                          1555   1555  2.05
SSBOND  26 CYS D  386    CYS D  449                          1555   1555  2.02
SSBOND  27 CYS D  413    CYS D  432                          1555   1555  2.03
LINK         ND2 ASN B  96                 C1  NAG B1452     1555   1555  1.46
SITE     1 AC1  1 ARG B 416
SITE     1 AC2  1 ASN B  96
CRYST1   60.829  193.880   75.717  90.00  91.88  90.00 P 1 21 1      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.016440  0.000000  0.000540        0.00000
SCALE2      0.000000  0.005158  0.000000        0.00000
SCALE3      0.000000  0.000000  0.013214        0.00000
MTRIX1   1 -1.000000 -0.003000 -0.015000       92.84500    1
MTRIX2   1  0.011000  0.586000 -0.810000       11.69000    1
MTRIX3   1  0.011000 -0.810000 -0.586000       24.68700    1
MTRIX1   2 -0.998000  0.045000 -0.043000       93.58000    1
MTRIX2   2 -0.061000 -0.587000  0.808000       -7.96800    1
MTRIX3   2  0.011000  0.809000  0.588000       -6.55000    1
MTRIX1   3  0.996000  0.061000 -0.058000       -3.82300    1
MTRIX2   3  0.061000 -0.998000  0.010000        8.51800    1
MTRIX3   3 -0.057000 -0.013000 -0.998000       40.67400    1
TER    2731      VAL A 452
TER    5364      THR B 451
TER    8041      PRO C 450
TER   10930      PHE D 472
TER   10981      ALA E  16
MASTER     1046    0    2   60   42    0    2   1510991    5   69  149
END