longtext: 4UZ5-pdb

content
HEADER    HYDROLASE                               04-SEP-14   4UZ5
TITLE     STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM IV - 2.1A
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: NOTUM;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: RESIDUES 80-452;
COMPND   5 EC: 3.1.1.1;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES;
COMPND   8 OTHER_DETAILS: GLYCOSYLATED AT N96
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE   6 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE   8 EXPRESSION_SYSTEM_CELL_LINE: HEK293T;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PHLSEC
KEYWDS    HYDROLASE, ESTERASE, EXTRACELLULAR, ALPHA/BETA HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.ZEBISCH,E.Y.JONES
REVDAT   1   25-FEB-15 4UZ5    0
JRNL        AUTH   S.KAKUGAWA,P.F.LANGTON,M.ZEBISCH,S.A.HOWELL,T.-H.CHANG,
JRNL        AUTH 2 Y.LIU,T.FEIZI,G.BINEVA,N.O'REILLY,A.P.SNIJDERS,E.Y.JONES,
JRNL        AUTH 3 J.-P.VINCENT
JRNL        TITL   NOTUM DEACYLATES WNT PROTEINS TO SUPPRESS SIGNALLING
JRNL        TITL 2 ACTIVITY
JRNL        REF    NATURE                                     2015
JRNL        REFN                   ESSN 1476-4687
JRNL        DOI    10.1038/NATURE14259
REMARK   2
REMARK   2 RESOLUTION.    2.10 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0073
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,
REMARK   3                 STEINER,NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.10
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 97.87
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.96
REMARK   3   NUMBER OF REFLECTIONS             : 25980
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.18936
REMARK   3   R VALUE            (WORKING SET) : 0.18746
REMARK   3   FREE R VALUE                     : 0.23710
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 3.9
REMARK   3   FREE R VALUE TEST SET COUNT      : 1046
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.100
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.155
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 1868
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 99.85
REMARK   3   BIN R VALUE           (WORKING SET) : 0.272
REMARK   3   BIN FREE R VALUE SET COUNT          : 73
REMARK   3   BIN FREE R VALUE                    : 0.292
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2796
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 15
REMARK   3   SOLVENT ATOMS            : 84
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 39.9
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 43.492
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.00
REMARK   3    B22 (A**2) : 0.00
REMARK   3    B33 (A**2) : 0.00
REMARK   3    B12 (A**2) : 0.00
REMARK   3    B13 (A**2) : 0.00
REMARK   3    B23 (A**2) : 0.00
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.179
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.169
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.126
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 9.873
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.957
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.930
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  2902 ; 0.013 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  2663 ; 0.001 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  3956 ; 1.504 ; 1.943
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  6099 ; 0.817 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   350 ; 6.216 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   141 ;36.152 ;22.553
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   456 ;13.178 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    29 ;18.033 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   420 ; 0.083 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  3295 ; 0.007 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):   726 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  1397 ; 2.266 ; 3.231
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  1396 ; 2.257 ; 3.228
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  1742 ; 3.398 ; 4.828
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1505 ; 3.380 ; 3.732
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 1
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A     1        A   451
REMARK   3    ORIGIN FOR THE GROUP (A): -21.6694  18.0125 -13.3796
REMARK   3    T TENSOR
REMARK   3      T11:   0.0080 T22:   0.0608
REMARK   3      T33:   0.0454 T12:   0.0067
REMARK   3      T13:  -0.0110 T23:   0.0218
REMARK   3    L TENSOR
REMARK   3      L11:   1.1714 L22:   0.7453
REMARK   3      L33:   0.6822 L12:   0.4021
REMARK   3      L13:  -0.2091 L23:  -0.1657
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0424 S12:   0.0265 S13:   0.0182
REMARK   3      S21:  -0.0528 S22:  -0.0012 S23:   0.0869
REMARK   3      S31:   0.0396 S32:   0.1118 S33:   0.0437
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE RIDING
REMARK   3  POSITIONS.
REMARK   4
REMARK   4 4UZ5 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 04-SEP-14.
REMARK 100 THE PDBE ID CODE IS EBI-61681.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 10-MAY-14
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.0
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I04
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9793
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL (PILATUS)
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : AIMLESS
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 27077
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.10
REMARK 200  RESOLUTION RANGE LOW       (A) : 69.00
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : 12.8
REMARK 200  R MERGE                    (I) : 0.11
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 18.30
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: OTHER
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NONE
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 52
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.5
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 10 %W/V PEG4000, 0.01 M CACL2, 0.05 M
REMARK 280  NACACOD PH 6.0, 0.20 M KCL, 1 MM HEPARIN HEXAMER
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 41 3 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X+1/2,-Y,Z+1/2
REMARK 290       3555   -X,Y+1/2,-Z+1/2
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290       5555   Z,X,Y
REMARK 290       6555   Z+1/2,-X+1/2,-Y
REMARK 290       7555   -Z+1/2,-X,Y+1/2
REMARK 290       8555   -Z,X+1/2,-Y+1/2
REMARK 290       9555   Y,Z,X
REMARK 290      10555   -Y,Z+1/2,-X+1/2
REMARK 290      11555   Y+1/2,-Z+1/2,-X
REMARK 290      12555   -Y+1/2,-Z,X+1/2
REMARK 290      13555   Y+3/4,X+1/4,-Z+1/4
REMARK 290      14555   -Y+3/4,-X+3/4,-Z+3/4
REMARK 290      15555   Y+1/4,-X+1/4,Z+3/4
REMARK 290      16555   -Y+1/4,X+3/4,Z+1/4
REMARK 290      17555   X+3/4,Z+1/4,-Y+1/4
REMARK 290      18555   -X+1/4,Z+3/4,Y+1/4
REMARK 290      19555   -X+3/4,-Z+3/4,-Y+3/4
REMARK 290      20555   X+1/4,-Z+1/4,Y+3/4
REMARK 290      21555   Z+3/4,Y+1/4,-X+1/4
REMARK 290      22555   Z+1/4,-Y+1/4,X+3/4
REMARK 290      23555   -Z+1/4,Y+3/4,X+1/4
REMARK 290      24555   -Z+3/4,-Y+3/4,-X+3/4
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000       69.20800
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       69.20800
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       69.20800
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000       69.20800
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       69.20800
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       69.20800
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   5  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY2   5  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   5  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY1   6  0.000000  0.000000  1.000000       69.20800
REMARK 290   SMTRY2   6 -1.000000  0.000000  0.000000       69.20800
REMARK 290   SMTRY3   6  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY1   7  0.000000  0.000000 -1.000000       69.20800
REMARK 290   SMTRY2   7 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  1.000000  0.000000       69.20800
REMARK 290   SMTRY1   8  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY2   8  1.000000  0.000000  0.000000       69.20800
REMARK 290   SMTRY3   8  0.000000 -1.000000  0.000000       69.20800
REMARK 290   SMTRY1   9  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   9  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY3   9  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  10  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2  10  0.000000  0.000000  1.000000       69.20800
REMARK 290   SMTRY3  10 -1.000000  0.000000  0.000000       69.20800
REMARK 290   SMTRY1  11  0.000000  1.000000  0.000000       69.20800
REMARK 290   SMTRY2  11  0.000000  0.000000 -1.000000       69.20800
REMARK 290   SMTRY3  11 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY1  12  0.000000 -1.000000  0.000000       69.20800
REMARK 290   SMTRY2  12  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY3  12  1.000000  0.000000  0.000000       69.20800
REMARK 290   SMTRY1  13  0.000000  1.000000  0.000000      103.81200
REMARK 290   SMTRY2  13  1.000000  0.000000  0.000000       34.60400
REMARK 290   SMTRY3  13  0.000000  0.000000 -1.000000       34.60400
REMARK 290   SMTRY1  14  0.000000 -1.000000  0.000000      103.81200
REMARK 290   SMTRY2  14 -1.000000  0.000000  0.000000      103.81200
REMARK 290   SMTRY3  14  0.000000  0.000000 -1.000000      103.81200
REMARK 290   SMTRY1  15  0.000000  1.000000  0.000000       34.60400
REMARK 290   SMTRY2  15 -1.000000  0.000000  0.000000       34.60400
REMARK 290   SMTRY3  15  0.000000  0.000000  1.000000      103.81200
REMARK 290   SMTRY1  16  0.000000 -1.000000  0.000000       34.60400
REMARK 290   SMTRY2  16  1.000000  0.000000  0.000000      103.81200
REMARK 290   SMTRY3  16  0.000000  0.000000  1.000000       34.60400
REMARK 290   SMTRY1  17  1.000000  0.000000  0.000000      103.81200
REMARK 290   SMTRY2  17  0.000000  0.000000  1.000000       34.60400
REMARK 290   SMTRY3  17  0.000000 -1.000000  0.000000       34.60400
REMARK 290   SMTRY1  18 -1.000000  0.000000  0.000000       34.60400
REMARK 290   SMTRY2  18  0.000000  0.000000  1.000000      103.81200
REMARK 290   SMTRY3  18  0.000000  1.000000  0.000000       34.60400
REMARK 290   SMTRY1  19 -1.000000  0.000000  0.000000      103.81200
REMARK 290   SMTRY2  19  0.000000  0.000000 -1.000000      103.81200
REMARK 290   SMTRY3  19  0.000000 -1.000000  0.000000      103.81200
REMARK 290   SMTRY1  20  1.000000  0.000000  0.000000       34.60400
REMARK 290   SMTRY2  20  0.000000  0.000000 -1.000000       34.60400
REMARK 290   SMTRY3  20  0.000000  1.000000  0.000000      103.81200
REMARK 290   SMTRY1  21  0.000000  0.000000  1.000000      103.81200
REMARK 290   SMTRY2  21  0.000000  1.000000  0.000000       34.60400
REMARK 290   SMTRY3  21 -1.000000  0.000000  0.000000       34.60400
REMARK 290   SMTRY1  22  0.000000  0.000000  1.000000       34.60400
REMARK 290   SMTRY2  22  0.000000 -1.000000  0.000000       34.60400
REMARK 290   SMTRY3  22  1.000000  0.000000  0.000000      103.81200
REMARK 290   SMTRY1  23  0.000000  0.000000 -1.000000       34.60400
REMARK 290   SMTRY2  23  0.000000  1.000000  0.000000      103.81200
REMARK 290   SMTRY3  23  1.000000  0.000000  0.000000       34.60400
REMARK 290   SMTRY1  24  0.000000  0.000000 -1.000000      103.81200
REMARK 290   SMTRY2  24  0.000000 -1.000000  0.000000      103.81200
REMARK 290   SMTRY3  24 -1.000000  0.000000  0.000000      103.81200
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 375
REMARK 375 SPECIAL POSITION
REMARK 375 THE FOLLOWING ATOMS ARE FOUND TO BE WITHIN 0.15 ANGSTROMS
REMARK 375 OF A SYMMETRY RELATED ATOM AND ARE ASSUMED TO BE ON SPECIAL
REMARK 375 POSITIONS.
REMARK 375
REMARK 375 ATOM RES CSSEQI
REMARK 375      HOH A2025   LIES ON A SPECIAL POSITION.
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A    78
REMARK 465     THR A    79
REMARK 465     GLY A    80
REMARK 465     SER A    81
REMARK 465     ALA A    82
REMARK 465     GLN A    83
REMARK 465     GLN A    84
REMARK 465     LEU A    85
REMARK 465     ASN A    86
REMARK 465     SER A   193
REMARK 465     LYS A   194
REMARK 465     SER A   195
REMARK 465     GLU A   196
REMARK 465     LYS A   197
REMARK 465     ASP A   420
REMARK 465     SER A   421
REMARK 465     HIS A   422
REMARK 465     LYS A   423
REMARK 465     ALA A   424
REMARK 465     SER A   425
REMARK 465     LYS A   426
REMARK 465     THR A   427
REMARK 465     PRO A   428
REMARK 465     LEU A   429
REMARK 465     LYS A   430
REMARK 465     GLY A   452
REMARK 465     THR A   453
REMARK 465     LYS A   454
REMARK 465     HIS A   455
REMARK 465     HIS A   456
REMARK 465     HIS A   457
REMARK 465     HIS A   458
REMARK 465     HIS A   459
REMARK 465     HIS A   460
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: COVALENT BOND ANGLES
REMARK 500
REMARK 500 THE STEREOCHEMICAL PARAMETERS OF THE FOLLOWING RESIDUES
REMARK 500 HAVE VALUES WHICH DEVIATE FROM EXPECTED VALUES BY MORE
REMARK 500 THAN 6*RMSD (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 500 IDENTIFIER; SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT: (10X,I3,1X,A3,1X,A1,I4,A1,3(1X,A4,2X),12X,F5.1)
REMARK 500
REMARK 500 EXPECTED VALUES PROTEIN: ENGH AND HUBER, 1999
REMARK 500 EXPECTED VALUES NUCLEIC ACID: CLOWNEY ET AL 1996
REMARK 500
REMARK 500  M RES CSSEQI ATM1   ATM2   ATM3
REMARK 500    ARG A 213   NE  -  CZ  -  NH1 ANGL. DEV. =   4.4 DEGREES
REMARK 500    ARG A 213   NE  -  CZ  -  NH2 ANGL. DEV. =  -4.0 DEGREES
REMARK 500    ARG A 218   NE  -  CZ  -  NH1 ANGL. DEV. =   3.5 DEGREES
REMARK 500    ARG A 218   NE  -  CZ  -  NH2 ANGL. DEV. =  -3.9 DEGREES
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    TRP A 128     -152.54     59.56
REMARK 500    MET A 143       45.79   -148.96
REMARK 500    ASN A 169       80.61   -152.60
REMARK 500    SER A 232     -128.56     65.65
REMARK 500    CYS A 279       81.86    -68.28
REMARK 500    THR A 352     -156.35    -85.37
REMARK 500    SER A 388     -169.59   -160.90
REMARK 500    GLU A 390      157.72     69.39
REMARK 500    ILE A 391      -37.27   -159.07
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL A1453
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800  NAG A1452  BOUND TO ASN A  96
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4UYU   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  I IODIDE COMPLEX - 2.3A
REMARK 900 RELATED ID: 4UYW   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   HEPARIN FRAGMENT COMPLEX - 1.7A
REMARK 900 RELATED ID: 4UYZ   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM II
REMARK 900   - 2.8A
REMARK 900 RELATED ID: 4UZ1   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  III - 1.4A
REMARK 900 RELATED ID: 4UZ6   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM V
REMARK 900   - SOS COMPLEX - 1.9A
REMARK 900 RELATED ID: 4UZ7   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  VII - SOS COMPLEX - 2.2A
REMARK 900 RELATED ID: 4UZ9   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  VII - SOS COMPLEX - 2.2A
REMARK 900 RELATED ID: 4UZA   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  VIII - PHOSPHATE COMPLEX - 2.4A
REMARK 900 RELATED ID: 4UZJ   RELATED DB: PDB
REMARK 900  STRUCTURE OF A WNT SIGNAL REGULATOR FROM DROSOPHILA -
REMARK 900  CRYSTAL FORM I - 2.4A
REMARK 900 RELATED ID: 4UZK   RELATED DB: PDB
REMARK 900  STRUCTURE OF A WNT SIGNAL REGULATOR FROM DROSOPHILA -
REMARK 900  CRYSTAL FORM II - 1.9A
REMARK 900 RELATED ID: 4UZL   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   MYRISTOLEATE COMPLEX - 2.1A
REMARK 900 RELATED ID: 4UZQ   RELATED DB: PDB
REMARK 900  STRUCTURE OF A WNT SIGNAL REGULATOR COMPLEX - CRYSTAL
REMARK 900  FORM IX - 1.5A
DBREF  4UZ5 A   80   452  UNP    Q6P988   NOTUM_HUMAN     80    452
SEQADV 4UZ5 GLU A   78  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 THR A   79  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 THR A  453  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 LYS A  454  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 HIS A  455  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 HIS A  456  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 HIS A  457  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 HIS A  458  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 HIS A  459  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 HIS A  460  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ5 SER A  330  UNP  Q6P988    CYS   330 ENGINEERED MUTATION
SEQRES   1 A  383  GLU THR GLY SER ALA GLN GLN LEU ASN GLU ASP LEU ARG
SEQRES   2 A  383  LEU HIS LEU LEU LEU ASN THR SER VAL THR CYS ASN ASP
SEQRES   3 A  383  GLY SER PRO ALA GLY TYR TYR LEU LYS GLU SER ARG GLY
SEQRES   4 A  383  SER ARG ARG TRP LEU LEU PHE LEU GLU GLY GLY TRP TYR
SEQRES   5 A  383  CYS PHE ASN ARG GLU ASN CYS ASP SER ARG TYR ASP THR
SEQRES   6 A  383  MET ARG ARG LEU MET SER SER ARG ASP TRP PRO ARG THR
SEQRES   7 A  383  ARG THR GLY THR GLY ILE LEU SER SER GLN PRO GLU GLU
SEQRES   8 A  383  ASN PRO TYR TRP TRP ASN ALA ASN MET VAL PHE ILE PRO
SEQRES   9 A  383  TYR CYS SER SER ASP VAL TRP SER GLY ALA SER SER LYS
SEQRES  10 A  383  SER GLU LYS ASN GLU TYR ALA PHE MET GLY ALA LEU ILE
SEQRES  11 A  383  ILE GLN GLU VAL VAL ARG GLU LEU LEU GLY ARG GLY LEU
SEQRES  12 A  383  SER GLY ALA LYS VAL LEU LEU LEU ALA GLY SER SER ALA
SEQRES  13 A  383  GLY GLY THR GLY VAL LEU LEU ASN VAL ASP ARG VAL ALA
SEQRES  14 A  383  GLU GLN LEU GLU LYS LEU GLY TYR PRO ALA ILE GLN VAL
SEQRES  15 A  383  ARG GLY LEU ALA ASP SER GLY TRP PHE LEU ASP ASN LYS
SEQRES  16 A  383  GLN TYR ARG HIS THR ASP CYS VAL ASP THR ILE THR CYS
SEQRES  17 A  383  ALA PRO THR GLU ALA ILE ARG ARG GLY ILE ARG TYR TRP
SEQRES  18 A  383  ASN GLY VAL VAL PRO GLU ARG CYS ARG ARG GLN PHE GLN
SEQRES  19 A  383  GLU GLY GLU GLU TRP ASN CYS PHE PHE GLY TYR LYS VAL
SEQRES  20 A  383  TYR PRO THR LEU ARG SER PRO VAL PHE VAL VAL GLN TRP
SEQRES  21 A  383  LEU PHE ASP GLU ALA GLN LEU THR VAL ASP ASN VAL HIS
SEQRES  22 A  383  LEU THR GLY GLN PRO VAL GLN GLU GLY LEU ARG LEU TYR
SEQRES  23 A  383  ILE GLN ASN LEU GLY ARG GLU LEU ARG HIS THR LEU LYS
SEQRES  24 A  383  ASP VAL PRO ALA SER PHE ALA PRO ALA CYS LEU SER HIS
SEQRES  25 A  383  GLU ILE ILE ILE ARG SER HIS TRP THR ASP VAL GLN VAL
SEQRES  26 A  383  LYS GLY THR SER LEU PRO ARG ALA LEU HIS CYS TRP ASP
SEQRES  27 A  383  ARG SER LEU HIS ASP SER HIS LYS ALA SER LYS THR PRO
SEQRES  28 A  383  LEU LYS GLY CYS PRO VAL HIS LEU VAL ASP SER CYS PRO
SEQRES  29 A  383  TRP PRO HIS CYS ASN PRO SER CYS PRO THR GLY THR LYS
SEQRES  30 A  383  HIS HIS HIS HIS HIS HIS
HET    NAG  A1452      14
HET     CL  A1453       1
HETNAM      CL CHLORIDE ION
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   2   CL    CL 1-
FORMUL   3  NAG    C8 H15 N O6
FORMUL   4  HOH   *84(H2 O)
HELIX    1   1 ASN A  132  MET A  143  1                                  12
HELIX    2   2 ARG A  144  SER A  148  5                                   5
HELIX    3   3 THR A  159  SER A  163  5                                   5
HELIX    4   4 MET A  203  GLY A  217  1                                  15
HELIX    5   5 ARG A  218  ALA A  223  5                                   6
HELIX    6   6 SER A  232  LEU A  252  1                                  21
HELIX    7   7 ALA A  286  ASN A  299  1                                  14
HELIX    8   8 PRO A  303  ARG A  308  1                                   6
HELIX    9   9 GLU A  314  PHE A  319  5                                   6
HELIX   10  10 PHE A  320  TYR A  325  1                                   6
HELIX   11  11 PRO A  326  LEU A  328  5                                   3
HELIX   12  12 GLU A  341  ASP A  347  1                                   7
HELIX   13  13 GLN A  357  LYS A  376  1                                  20
HELIX   14  14 ARG A  394  ASP A  399  5                                   6
HELIX   15  15 LEU A  407  HIS A  419  1                                  13
SHEET    1  AA10 THR A 155  ARG A 156  0
SHEET    2  AA10 LEU A  89  LEU A  93 -1  O  LEU A  89   N  ARG A 156
SHEET    3  AA10 GLY A 108  LYS A 112 -1  O  TYR A 109   N  HIS A  92
SHEET    4  AA10 ASN A 176  ILE A 180 -1  O  MET A 177   N  LYS A 112
SHEET    5  AA10 ARG A 119  LEU A 124  1  O  ARG A 119   N  ASN A 176
SHEET    6  AA10 VAL A 225  SER A 231  1  O  VAL A 225   N  TRP A 120
SHEET    7  AA10 GLN A 258  ASP A 264  1  O  GLN A 258   N  LEU A 226
SHEET    8  AA10 VAL A 332  VAL A 335  1  O  PHE A 333   N  ALA A 263
SHEET    9  AA10 SER A 381  ALA A 383  1  N  PHE A 382   O  VAL A 334
SHEET   10  AA10 HIS A 435  VAL A 437  1  O  LEU A 436   N  ALA A 383
SHEET    1  AB 2 PHE A 339  ASP A 340  0
SHEET    2  AB 2 LEU A 387  SER A 388  1  N  SER A 388   O  PHE A 339
SHEET    1  AC 2 GLN A 401  VAL A 402  0
SHEET    2  AC 2 THR A 405  SER A 406 -1  O  THR A 405   N  VAL A 402
SSBOND   1 CYS A  101    CYS A  183                          1555   1555  2.16
SSBOND   2 CYS A  130    CYS A  136                          1555   1555  2.02
SSBOND   3 CYS A  279    CYS A  285                          1555   1555  2.07
SSBOND   4 CYS A  306    CYS A  318                          1555   1555  2.15
SSBOND   5 CYS A  386    CYS A  449                          1555   1555  2.04
SSBOND   6 CYS A  413    CYS A  432                          1555   1555  2.03
SSBOND   7 CYS A  440    CYS A  445                          1555   1555  2.07
LINK         ND2 ASN A  96                 C1  NAG A1452     1555   1555  1.45
SITE     1 AC1  3 ARG A  90  HIS A  92  GLN A 311
SITE     1 AC2  3 ASN A  96  VAL A  99  HOH A2084
CRYST1  138.416  138.416  138.416  90.00  90.00  90.00 P 41 3 2      8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.007225  0.000000  0.000000        0.00000
SCALE2      0.000000  0.007225  0.000000        0.00000
SCALE3      0.000000  0.000000  0.007225        0.00000
TER    2803      THR A 451
MASTER      479    0    2   15   14    0    2    6 2901    1   29   30
END