longtext: 4UZ9-pdb

content
HEADER    HYDROLASE                               04-SEP-14   4UZ9
TITLE     STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM VII -
TITLE    2 SOS COMPLEX - 2.2A
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PROTEIN NOTUM HOMOLOG;
COMPND   3 CHAIN: A;
COMPND   4 FRAGMENT: RESIDUES 81-451;
COMPND   5 EC: 3.1.1.1;
COMPND   6 ENGINEERED: YES;
COMPND   7 MUTATION: YES;
COMPND   8 OTHER_DETAILS: GLYCOSYLATED AT N96
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE   6 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE   7 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE   8 EXPRESSION_SYSTEM_CELL_LINE: HEK293T;
SOURCE   9 EXPRESSION_SYSTEM_PLASMID: PHLSEC
KEYWDS    HYDROLASE, WNT, ESTERASE, EXTRACELLULAR, ALPHA/BETA HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.ZEBISCH,E.Y.JONES
REVDAT   1   25-FEB-15 4UZ9    0
JRNL        AUTH   S.KAKUGAWA,P.F.LANGTON,M.ZEBISCH,S.A.HOWELL,T.-H.CHANG,
JRNL        AUTH 2 Y.LIU,T.FEIZI,G.BINEVA,N.O'REILLY,A.P.SNIJDERS,E.Y.JONES,
JRNL        AUTH 3 J.-P.VINCENT
JRNL        TITL   NOTUM DEACYLATES WNT PROTEINS TO SUPPRESS SIGNALLING
JRNL        TITL 2 ACTIVITY
JRNL        REF    NATURE                                     2015
JRNL        REFN                   ESSN 1476-4687
JRNL        DOI    10.1038/NATURE14259
REMARK   2
REMARK   2 RESOLUTION.    2.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0073
REMARK   3   AUTHORS     : MURSHUDOV,SKUBAK,LEBEDEV,PANNU,
REMARK   3                 STEINER,NICHOLLS,WINN,LONG,VAGIN
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 65.58
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NONE
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.97
REMARK   3   NUMBER OF REFLECTIONS             : 24266
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.19196
REMARK   3   R VALUE            (WORKING SET) : 0.19007
REMARK   3   FREE R VALUE                     : 0.23653
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 4.1
REMARK   3   FREE R VALUE TEST SET COUNT      : 1030
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.200
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.257
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 1743
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.00
REMARK   3   BIN R VALUE           (WORKING SET) : 0.263
REMARK   3   BIN FREE R VALUE SET COUNT          : 77
REMARK   3   BIN FREE R VALUE                    : 0.305
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 2875
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 70
REMARK   3   SOLVENT ATOMS            : 115
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : 26.2
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 39.040
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 1.19
REMARK   3    B22 (A**2) : 1.19
REMARK   3    B33 (A**2) : -2.38
REMARK   3    B12 (A**2) : 0.00
REMARK   3    B13 (A**2) : 0.00
REMARK   3    B23 (A**2) : 0.00
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.214
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.187
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.138
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 11.304
REMARK   3
REMARK   3  CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.948
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.912
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  3033 ; 0.014 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  2753 ; 0.001 ; 0.020
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  4149 ; 1.674 ; 1.967
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  6315 ; 0.864 ; 3.000
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   360 ; 6.581 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   147 ;32.934 ;22.585
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   477 ;15.456 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    31 ;22.521 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   446 ; 0.096 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  3388 ; 0.007 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):   747 ; 0.001 ; 0.020
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  1428 ; 2.223 ; 2.572
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  1427 ; 2.224 ; 2.572
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  1783 ; 3.599 ; 3.841
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  1605 ; 3.441 ; 3.275
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ANISOTROPIC THERMAL FACTOR RESTRAINTS.   COUNT   RMS    WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 1
REMARK   3
REMARK   3   TLS GROUP :     1
REMARK   3    NUMBER OF COMPONENTS GROUP :    1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A    86        A   451
REMARK   3    ORIGIN FOR THE GROUP (A): -34.1646   1.1693  -5.0749
REMARK   3    T TENSOR
REMARK   3      T11:   0.0132 T22:   0.0286
REMARK   3      T33:   0.1584 T12:  -0.0120
REMARK   3      T13:   0.0157 T23:   0.0075
REMARK   3    L TENSOR
REMARK   3      L11:   0.5446 L22:   0.7960
REMARK   3      L33:   0.9861 L12:   0.2896
REMARK   3      L13:   0.0419 L23:  -0.0344
REMARK   3    S TENSOR
REMARK   3      S11:   0.0440 S12:  -0.0372 S13:  -0.0658
REMARK   3      S21:  -0.0021 S22:  -0.0728 S23:  -0.0636
REMARK   3      S31:   0.0186 S32:   0.0132 S33:   0.0288
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN ADDED IN THE
REMARK   3   RIDING POSITIONS.
REMARK   4
REMARK   4 4UZ9 COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY PDBE ON 05-SEP-14.
REMARK 100 THE PDBE ID CODE IS EBI-61696.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 15-APR-14
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.5
REMARK 200  NUMBER OF CRYSTALS USED        : 1
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I03
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.97
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : PIXEL (PILATUS)
REMARK 200  DETECTOR MANUFACTURER          : DECTRIS
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : NULL
REMARK 200  DATA SCALING SOFTWARE          : NULL
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 25372
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.20
REMARK 200  RESOLUTION RANGE LOW       (A) : 97.00
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : -3.0
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 100.0
REMARK 200  DATA REDUNDANCY                : 7.1
REMARK 200  R MERGE                    (I) : 0.16
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 13.40
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : NULL
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : NULL
REMARK 200  COMPLETENESS FOR SHELL     (%) : NULL
REMARK 200  DATA REDUNDANCY IN SHELL       : NULL
REMARK 200  R MERGE FOR SHELL          (I) : NULL
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : NULL
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: OTHER
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NONE
REMARK 200
REMARK 200 REMARK: NONE
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 55
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 2.7
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 200 MM AMMONIUM SULFATE 100 MM
REMARK 280  MES 6.5   35 % W/V PEP 629 20MM SOS
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 41 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z+1/2
REMARK 290       3555   -Y+1/2,X+1/2,Z+1/4
REMARK 290       4555   Y+1/2,-X+1/2,Z+3/4
REMARK 290       5555   -X+1/2,Y+1/2,-Z+1/4
REMARK 290       6555   X+1/2,-Y+1/2,-Z+3/4
REMARK 290       7555   Y,X,-Z
REMARK 290       8555   -Y,-X,-Z+1/2
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000       97.57400
REMARK 290   SMTRY1   3  0.000000 -1.000000  0.000000       34.81500
REMARK 290   SMTRY2   3  1.000000  0.000000  0.000000       34.81500
REMARK 290   SMTRY3   3  0.000000  0.000000  1.000000       48.78700
REMARK 290   SMTRY1   4  0.000000  1.000000  0.000000       34.81500
REMARK 290   SMTRY2   4 -1.000000  0.000000  0.000000       34.81500
REMARK 290   SMTRY3   4  0.000000  0.000000  1.000000      146.36100
REMARK 290   SMTRY1   5 -1.000000  0.000000  0.000000       34.81500
REMARK 290   SMTRY2   5  0.000000  1.000000  0.000000       34.81500
REMARK 290   SMTRY3   5  0.000000  0.000000 -1.000000       48.78700
REMARK 290   SMTRY1   6  1.000000  0.000000  0.000000       34.81500
REMARK 290   SMTRY2   6  0.000000 -1.000000  0.000000       34.81500
REMARK 290   SMTRY3   6  0.000000  0.000000 -1.000000      146.36100
REMARK 290   SMTRY1   7  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY2   7  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   7  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   8  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY2   8 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY3   8  0.000000  0.000000 -1.000000       97.57400
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: MONOMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: MONOMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A    78
REMARK 465     THR A    79
REMARK 465     GLY A    80
REMARK 465     SER A    81
REMARK 465     ALA A    82
REMARK 465     GLN A    83
REMARK 465     GLN A    84
REMARK 465     LEU A    85
REMARK 465     HIS A   422
REMARK 465     LYS A   423
REMARK 465     ALA A   424
REMARK 465     SER A   425
REMARK 465     LYS A   426
REMARK 465     THR A   427
REMARK 465     PRO A   428
REMARK 465     LEU A   429
REMARK 465     LYS A   430
REMARK 465     GLY A   431
REMARK 465     GLY A   452
REMARK 465     THR A   453
REMARK 465     LYS A   454
REMARK 465     HIS A   455
REMARK 465     HIS A   456
REMARK 465     HIS A   457
REMARK 465     HIS A   458
REMARK 465     HIS A   459
REMARK 465     HIS A   460
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    TRP A 128     -140.64     54.92
REMARK 500    MET A 143       53.55   -142.57
REMARK 500    SER A 232     -123.51     64.28
REMARK 500    SER A 388     -169.80   -160.41
REMARK 500    GLU A 390      160.49     70.55
REMARK 500    ILE A 391      -38.20   -158.01
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE SCR A1453
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR RESIDUE  CL A1454
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: BINDING SITE FOR MONO-SACCHARIDE
REMARK 800  NAG A1452  BOUND TO ASN A  96
REMARK 900
REMARK 900 RELATED ENTRIES
REMARK 900 RELATED ID: 4UYU   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   IODIDE COMPLEX - 2.3A
REMARK 900 RELATED ID: 4UYW   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   HEPARIN FRAGMENT COMPLEX - 1.7A
REMARK 900 RELATED ID: 4UYZ   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM II
REMARK 900   - 2.8A
REMARK 900 RELATED ID: 4UZ1   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  III - 1.4A
REMARK 900 RELATED ID: 4UZ5   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM IV
REMARK 900   - 2.1A
REMARK 900 RELATED ID: 4UZ6   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM V
REMARK 900   - SOS COMPLEX - 1.9A
REMARK 900 RELATED ID: 4UZ7   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACETYLASE NOTUM - CRYSTAL FORM
REMARK 900  VII - SOS COMPLEX - 2.2A
REMARK 900 RELATED ID: 4UZA   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  VIII - PHOSPHATE COMPLEX - 2.4A
REMARK 900 RELATED ID: 4UZJ   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM
REMARK 900  III - 1.4A
REMARK 900 RELATED ID: 4UZK   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM FROM DROSOPHILA -
REMARK 900  CRYSTAL FORM II - 1.9A
REMARK 900 RELATED ID: 4UZL   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I
REMARK 900   MYRISTOLEATE COMPLEX - 2.1A
REMARK 900 RELATED ID: 4UZQ   RELATED DB: PDB
REMARK 900  STRUCTURE OF THE WNT DEACYLASE NOTUM IN COMPLEX WITH
REMARK 900  O-PALMITOLEOYL SERINE - CRYSTAL FORM IX - 1.5A
REMARK 900 RELATED ID: 4WBH   RELATED DB: PDB
REMARK 999
REMARK 999 SEQUENCE
REMARK 999 C330S ENGINEERED MUTATION
DBREF  4UZ9 A   81   451  UNP    Q6P988   NOTUM_HUMAN     81    451
SEQADV 4UZ9 GLU A   78  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 THR A   79  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 GLY A   80  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 GLY A  452  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 THR A  453  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 LYS A  454  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 HIS A  455  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 HIS A  456  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 HIS A  457  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 HIS A  458  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 HIS A  459  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 HIS A  460  UNP  Q6P988              EXPRESSION TAG
SEQADV 4UZ9 SER A  330  UNP  Q6P988    CYS   330 ENGINEERED MUTATION
SEQRES   1 A  383  GLU THR GLY SER ALA GLN GLN LEU ASN GLU ASP LEU ARG
SEQRES   2 A  383  LEU HIS LEU LEU LEU ASN THR SER VAL THR CYS ASN ASP
SEQRES   3 A  383  GLY SER PRO ALA GLY TYR TYR LEU LYS GLU SER ARG GLY
SEQRES   4 A  383  SER ARG ARG TRP LEU LEU PHE LEU GLU GLY GLY TRP TYR
SEQRES   5 A  383  CYS PHE ASN ARG GLU ASN CYS ASP SER ARG TYR ASP THR
SEQRES   6 A  383  MET ARG ARG LEU MET SER SER ARG ASP TRP PRO ARG THR
SEQRES   7 A  383  ARG THR GLY THR GLY ILE LEU SER SER GLN PRO GLU GLU
SEQRES   8 A  383  ASN PRO TYR TRP TRP ASN ALA ASN MET VAL PHE ILE PRO
SEQRES   9 A  383  TYR CYS SER SER ASP VAL TRP SER GLY ALA SER SER LYS
SEQRES  10 A  383  SER GLU LYS ASN GLU TYR ALA PHE MET GLY ALA LEU ILE
SEQRES  11 A  383  ILE GLN GLU VAL VAL ARG GLU LEU LEU GLY ARG GLY LEU
SEQRES  12 A  383  SER GLY ALA LYS VAL LEU LEU LEU ALA GLY SER SER ALA
SEQRES  13 A  383  GLY GLY THR GLY VAL LEU LEU ASN VAL ASP ARG VAL ALA
SEQRES  14 A  383  GLU GLN LEU GLU LYS LEU GLY TYR PRO ALA ILE GLN VAL
SEQRES  15 A  383  ARG GLY LEU ALA ASP SER GLY TRP PHE LEU ASP ASN LYS
SEQRES  16 A  383  GLN TYR ARG HIS THR ASP CYS VAL ASP THR ILE THR CYS
SEQRES  17 A  383  ALA PRO THR GLU ALA ILE ARG ARG GLY ILE ARG TYR TRP
SEQRES  18 A  383  ASN GLY VAL VAL PRO GLU ARG CYS ARG ARG GLN PHE GLN
SEQRES  19 A  383  GLU GLY GLU GLU TRP ASN CYS PHE PHE GLY TYR LYS VAL
SEQRES  20 A  383  TYR PRO THR LEU ARG SER PRO VAL PHE VAL VAL GLN TRP
SEQRES  21 A  383  LEU PHE ASP GLU ALA GLN LEU THR VAL ASP ASN VAL HIS
SEQRES  22 A  383  LEU THR GLY GLN PRO VAL GLN GLU GLY LEU ARG LEU TYR
SEQRES  23 A  383  ILE GLN ASN LEU GLY ARG GLU LEU ARG HIS THR LEU LYS
SEQRES  24 A  383  ASP VAL PRO ALA SER PHE ALA PRO ALA CYS LEU SER HIS
SEQRES  25 A  383  GLU ILE ILE ILE ARG SER HIS TRP THR ASP VAL GLN VAL
SEQRES  26 A  383  LYS GLY THR SER LEU PRO ARG ALA LEU HIS CYS TRP ASP
SEQRES  27 A  383  ARG SER LEU HIS ASP SER HIS LYS ALA SER LYS THR PRO
SEQRES  28 A  383  LEU LYS GLY CYS PRO VAL HIS LEU VAL ASP SER CYS PRO
SEQRES  29 A  383  TRP PRO HIS CYS ASN PRO SER CYS PRO THR GLY THR LYS
SEQRES  30 A  383  HIS HIS HIS HIS HIS HIS
HET    NAG  A1452      14
HET    SCR  A1453      55
HET     CL  A1454       1
HETNAM      CL CHLORIDE ION
HETNAM     SCR SUCROSE OCTASULFATE
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
FORMUL   2   CL    CL 1-
FORMUL   3  SCR    C12 H22 O35 S8
FORMUL   4  NAG    C8 H15 N O6
FORMUL   5  HOH   *115(H2 O)
HELIX    1   1 ASN A  132  MET A  143  1                                  12
HELIX    2   2 ARG A  144  MET A  147  5                                   4
HELIX    3   3 THR A  159  SER A  163  5                                   5
HELIX    4   4 MET A  203  ARG A  218  1                                  16
HELIX    5   5 GLY A  219  ALA A  223  5                                   5
HELIX    6   6 SER A  232  GLY A  253  1                                  22
HELIX    7   7 ALA A  286  ASN A  299  1                                  14
HELIX    8   8 PRO A  303  ARG A  308  1                                   6
HELIX    9   9 GLU A  314  PHE A  319  5                                   6
HELIX   10  10 PHE A  320  TYR A  325  1                                   6
HELIX   11  11 PRO A  326  LEU A  328  5                                   3
HELIX   12  12 GLU A  341  ASP A  347  1                                   7
HELIX   13  13 GLN A  357  LYS A  376  1                                  20
HELIX   14  14 ARG A  394  ASP A  399  5                                   6
HELIX   15  15 LEU A  407  ASP A  420  1                                  14
SHEET    1  AA10 THR A 155  ARG A 156  0
SHEET    2  AA10 LEU A  89  LEU A  93 -1  O  LEU A  89   N  ARG A 156
SHEET    3  AA10 GLY A 108  LYS A 112 -1  O  TYR A 109   N  HIS A  92
SHEET    4  AA10 ASN A 176  ILE A 180 -1  O  MET A 177   N  LYS A 112
SHEET    5  AA10 ARG A 119  LEU A 124  1  O  ARG A 119   N  ASN A 176
SHEET    6  AA10 VAL A 225  SER A 231  1  O  VAL A 225   N  TRP A 120
SHEET    7  AA10 GLN A 258  ASP A 264  1  O  GLN A 258   N  LEU A 226
SHEET    8  AA10 VAL A 332  VAL A 335  1  O  PHE A 333   N  ALA A 263
SHEET    9  AA10 SER A 381  ALA A 383  1  N  PHE A 382   O  VAL A 334
SHEET   10  AA10 HIS A 435  VAL A 437  1  O  LEU A 436   N  ALA A 383
SHEET    1  AB 2 PHE A 339  ASP A 340  0
SHEET    2  AB 2 LEU A 387  SER A 388  0
SHEET    1  AC 2 GLN A 401  VAL A 402  0
SHEET    2  AC 2 THR A 405  SER A 406 -1  O  THR A 405   N  VAL A 402
SSBOND   1 CYS A  101    CYS A  183                          1555   1555  2.13
SSBOND   2 CYS A  130    CYS A  136                          1555   1555  1.99
SSBOND   3 CYS A  279    CYS A  285                          1555   1555  2.06
SSBOND   4 CYS A  306    CYS A  318                          1555   1555  2.11
SSBOND   5 CYS A  386    CYS A  449                          1555   1555  2.06
SSBOND   6 CYS A  413    CYS A  432                          1555   1555  2.04
SSBOND   7 CYS A  440    CYS A  445                          1555   1555  2.04
LINK         ND2 ASN A  96                 C1  NAG A1452     1555   1555  1.45
SITE     1 AC1 15 SER A 117  ARG A 118  ARG A 119  ASN A 174
SITE     2 AC1 15 LYS A 224  LYS A 272  HIS A 276  GLU A 289
SITE     3 AC1 15 ARG A 292  HIS A 412  ARG A 416  HIS A 419
SITE     4 AC1 15 HOH A2088  HOH A2114  HOH A2115
SITE     1 AC2  1 ARG A  90
SITE     1 AC3  4 ASN A  96  VAL A  99  SER A 193  PRO A 447
CRYST1   69.630   69.630  195.148  90.00  90.00  90.00 P 41 21 2     8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.014362  0.000000  0.000000        0.00000
SCALE2      0.000000  0.014362  0.000000        0.00000
SCALE3      0.000000  0.000000  0.005124        0.00000
TER    2876      THR A 451
MASTER      378    0    3   15   14    0    6    6 3060    1   84   30
END