longtext: 4WBH-pdb

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HEADER    HYDROLASE                               03-SEP-14   4WBH
TITLE     STRUCTURE OF THE WNT DEACYLASE NOTUM - CRYSTAL FORM I APO - 2.2A
COMPND    MOL_ID: 1;
COMPND   2 MOLECULE: PROTEIN NOTUM HOMOLOG;
COMPND   3 CHAIN: A, B;
COMPND   4 FRAGMENT: UNP RESIDUES 38-496;
COMPND   5 EC: 3.-.-.-;
COMPND   6 ENGINEERED: YES
SOURCE    MOL_ID: 1;
SOURCE   2 ORGANISM_SCIENTIFIC: HOMO SAPIENS;
SOURCE   3 ORGANISM_COMMON: HUMAN;
SOURCE   4 ORGANISM_TAXID: 9606;
SOURCE   5 GENE: NOTUM, OK/SW-CL.30;
SOURCE   6 EXPRESSION_SYSTEM: HOMO SAPIENS;
SOURCE   7 EXPRESSION_SYSTEM_COMMON: HUMAN;
SOURCE   8 EXPRESSION_SYSTEM_TAXID: 9606;
SOURCE   9 EXPRESSION_SYSTEM_CELL_LINE: HEK293T;
SOURCE  10 EXPRESSION_SYSTEM_PLASMID: PHLSEC
KEYWDS    WNT, EXTRACELLULAR ESTERASE, ALPHA/BETA HYDROLASE, HYDROLASE
EXPDTA    X-RAY DIFFRACTION
AUTHOR    M.ZEBISCH,E.Y.JONES
REVDAT   1   25-FEB-15 4WBH    0
JRNL        AUTH   S.KAKUGAWA,P.F.LANGTON,M.ZEBISCH,S.A.HOWELL,T.-H.CHANG,
JRNL        AUTH 2 Y.LIU,T.FEIZI,G.BINEVA,N.O'REILLY,A.P.SNIJDERS,E.Y.JONES,
JRNL        AUTH 3 J.-P.VINCENT
JRNL        TITL   NOTUM DEACYLATES WNT PROTEINS TO SUPPRESS SIGNALLING
JRNL        TITL 2 ACTIVITY
JRNL        REF    NATURE                                     2015
JRNL        REFN                   ESSN 1476-4687
JRNL        DOI    10.1038/NATURE14259
REMARK   2
REMARK   2 RESOLUTION.    2.20 ANGSTROMS.
REMARK   3
REMARK   3 REFINEMENT.
REMARK   3   PROGRAM     : REFMAC 5.8.0073
REMARK   3   AUTHORS     : MURSHUDOV,VAGIN,DODSON
REMARK   3
REMARK   3    REFINEMENT TARGET : MAXIMUM LIKELIHOOD
REMARK   3
REMARK   3  DATA USED IN REFINEMENT.
REMARK   3   RESOLUTION RANGE HIGH (ANGSTROMS) : 2.20
REMARK   3   RESOLUTION RANGE LOW  (ANGSTROMS) : 87.72
REMARK   3   DATA CUTOFF            (SIGMA(F)) : NULL
REMARK   3   COMPLETENESS FOR RANGE        (%) : 99.6
REMARK   3   NUMBER OF REFLECTIONS             : 38158
REMARK   3
REMARK   3  FIT TO DATA USED IN REFINEMENT.
REMARK   3   CROSS-VALIDATION METHOD          : THROUGHOUT
REMARK   3   FREE R VALUE TEST SET SELECTION  : RANDOM
REMARK   3   R VALUE     (WORKING + TEST SET) : 0.217
REMARK   3   R VALUE            (WORKING SET) : 0.215
REMARK   3   FREE R VALUE                     : 0.262
REMARK   3   FREE R VALUE TEST SET SIZE   (%) : 5.000
REMARK   3   FREE R VALUE TEST SET COUNT      : 2013
REMARK   3
REMARK   3  FIT IN THE HIGHEST RESOLUTION BIN.
REMARK   3   TOTAL NUMBER OF BINS USED           : 20
REMARK   3   BIN RESOLUTION RANGE HIGH       (A) : 2.20
REMARK   3   BIN RESOLUTION RANGE LOW        (A) : 2.26
REMARK   3   REFLECTION IN BIN     (WORKING SET) : 2802
REMARK   3   BIN COMPLETENESS (WORKING+TEST) (%) : 100.0
REMARK   3   BIN R VALUE           (WORKING SET) : 0.2830
REMARK   3   BIN FREE R VALUE SET COUNT          : 140
REMARK   3   BIN FREE R VALUE                    : 0.3090
REMARK   3
REMARK   3  NUMBER OF NON-HYDROGEN ATOMS USED IN REFINEMENT.
REMARK   3   PROTEIN ATOMS            : 5574
REMARK   3   NUCLEIC ACID ATOMS       : 0
REMARK   3   HETEROGEN ATOMS          : 23
REMARK   3   SOLVENT ATOMS            : 59
REMARK   3
REMARK   3  B VALUES.
REMARK   3   FROM WILSON PLOT           (A**2) : NULL
REMARK   3   MEAN B VALUE      (OVERALL, A**2) : 43.71
REMARK   3   OVERALL ANISOTROPIC B VALUE.
REMARK   3    B11 (A**2) : 0.64000
REMARK   3    B22 (A**2) : -0.70000
REMARK   3    B33 (A**2) : 0.06000
REMARK   3    B12 (A**2) : 0.00000
REMARK   3    B13 (A**2) : 0.00000
REMARK   3    B23 (A**2) : 0.00000
REMARK   3
REMARK   3  ESTIMATED OVERALL COORDINATE ERROR.
REMARK   3   ESU BASED ON R VALUE                            (A): 0.291
REMARK   3   ESU BASED ON FREE R VALUE                       (A): 0.226
REMARK   3   ESU BASED ON MAXIMUM LIKELIHOOD                 (A): 0.186
REMARK   3   ESU FOR B VALUES BASED ON MAXIMUM LIKELIHOOD (A**2): 16.789
REMARK   3
REMARK   3 CORRELATION COEFFICIENTS.
REMARK   3   CORRELATION COEFFICIENT FO-FC      : 0.937
REMARK   3   CORRELATION COEFFICIENT FO-FC FREE : 0.907
REMARK   3
REMARK   3  RMS DEVIATIONS FROM IDEAL VALUES        COUNT    RMS    WEIGHT
REMARK   3   BOND LENGTHS REFINED ATOMS        (A):  5777 ; 0.007 ; 0.019
REMARK   3   BOND LENGTHS OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   BOND ANGLES REFINED ATOMS   (DEGREES):  7872 ; 1.170 ; 1.943
REMARK   3   BOND ANGLES OTHERS          (DEGREES):  NULL ;  NULL ;  NULL
REMARK   3   TORSION ANGLES, PERIOD 1    (DEGREES):   697 ; 6.222 ; 5.000
REMARK   3   TORSION ANGLES, PERIOD 2    (DEGREES):   276 ;33.993 ;22.391
REMARK   3   TORSION ANGLES, PERIOD 3    (DEGREES):   911 ;15.103 ;15.000
REMARK   3   TORSION ANGLES, PERIOD 4    (DEGREES):    59 ;17.235 ;15.000
REMARK   3   CHIRAL-CENTER RESTRAINTS       (A**3):   838 ; 0.086 ; 0.200
REMARK   3   GENERAL PLANES REFINED ATOMS      (A):  4467 ; 0.005 ; 0.021
REMARK   3   GENERAL PLANES OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED CONTACTS OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   NON-BONDED TORSION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) REFINED ATOMS      (A):  NULL ;  NULL ;  NULL
REMARK   3   H-BOND (X...Y) OTHERS             (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION REFINED ATOMS (A):  NULL ;  NULL ;  NULL
REMARK   3   POTENTIAL METAL-ION OTHERS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW REFINED ATOMS        (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY VDW OTHERS               (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND REFINED ATOMS     (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY H-BOND OTHERS            (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION REFINED ATOMS  (A):  NULL ;  NULL ;  NULL
REMARK   3   SYMMETRY METAL-ION OTHERS         (A):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  ISOTROPIC THERMAL FACTOR RESTRAINTS.     COUNT   RMS    WEIGHT
REMARK   3   MAIN-CHAIN BOND REFINED ATOMS  (A**2):  2794 ; 1.487 ; 3.048
REMARK   3   MAIN-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   MAIN-CHAIN ANGLE REFINED ATOMS (A**2):  3483 ; 2.559 ; 4.556
REMARK   3   MAIN-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN BOND REFINED ATOMS  (A**2):  2981 ; 1.846 ; 3.309
REMARK   3   SIDE-CHAIN BOND OTHER ATOMS    (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE REFINED ATOMS (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SIDE-CHAIN ANGLE OTHER ATOMS   (A**2):  NULL ;  NULL ;  NULL
REMARK   3   LONG RANGE B REFINED ATOMS     (A**2):  8576 ; 5.155 ;25.385
REMARK   3   LONG RANGE B OTHER ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3 ANISOTROPIC THERMAL FACTOR RESTRAINTS.    COUNT   RMS   WEIGHT
REMARK   3   RIGID-BOND RESTRAINTS          (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; FREE ATOMS         (A**2):  NULL ;  NULL ;  NULL
REMARK   3   SPHERICITY; BONDED ATOMS       (A**2):  NULL ;  NULL ;  NULL
REMARK   3
REMARK   3  NCS RESTRAINTS STATISTICS
REMARK   3   NUMBER OF DIFFERENT NCS GROUPS : NULL
REMARK   3
REMARK   3  TLS DETAILS
REMARK   3   NUMBER OF TLS GROUPS  : 2
REMARK   3
REMARK   3   TLS GROUP : 1
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   A    86        A   722
REMARK   3    ORIGIN FOR THE GROUP (A):  49.9997 -27.2715 -28.2823
REMARK   3    T TENSOR
REMARK   3      T11:   0.0204 T22:   0.1866
REMARK   3      T33:   0.0403 T12:   0.0182
REMARK   3      T13:  -0.0253 T23:  -0.0099
REMARK   3    L TENSOR
REMARK   3      L11:   2.5871 L22:   0.3687
REMARK   3      L33:   0.2687 L12:  -0.0016
REMARK   3      L13:   0.5283 L23:   0.1386
REMARK   3    S TENSOR
REMARK   3      S11:  -0.0940 S12:   0.1717 S13:   0.2402
REMARK   3      S21:   0.0544 S22:   0.0898 S23:  -0.0715
REMARK   3      S31:   0.0278 S32:   0.0367 S33:   0.0042
REMARK   3
REMARK   3   TLS GROUP : 2
REMARK   3    NUMBER OF COMPONENTS GROUP : 1
REMARK   3    COMPONENTS        C SSSEQI   TO  C SSSEQI
REMARK   3    RESIDUE RANGE :   B    86        B   737
REMARK   3    ORIGIN FOR THE GROUP (A):  16.4234 -14.7081  -0.5079
REMARK   3    T TENSOR
REMARK   3      T11:   0.0258 T22:   0.2142
REMARK   3      T33:   0.0043 T12:   0.0303
REMARK   3      T13:   0.0012 T23:   0.0033
REMARK   3    L TENSOR
REMARK   3      L11:   0.8555 L22:   0.9948
REMARK   3      L33:   0.3293 L12:  -0.4198
REMARK   3      L13:   0.0428 L23:  -0.1688
REMARK   3    S TENSOR
REMARK   3      S11:   0.0539 S12:  -0.0768 S13:  -0.0458
REMARK   3      S21:  -0.0105 S22:  -0.0422 S23:   0.0542
REMARK   3      S31:  -0.0115 S32:   0.0222 S33:  -0.0117
REMARK   3
REMARK   3  BULK SOLVENT MODELLING.
REMARK   3   METHOD USED : MASK
REMARK   3   PARAMETERS FOR MASK CALCULATION
REMARK   3   VDW PROBE RADIUS   : 1.20
REMARK   3   ION PROBE RADIUS   : 0.80
REMARK   3   SHRINKAGE RADIUS   : 0.80
REMARK   3
REMARK   3  OTHER REFINEMENT REMARKS: HYDROGENS HAVE BEEN USED IF PRESENT IN
REMARK   3  THE INPUT
REMARK   4
REMARK   4 4WBH COMPLIES WITH FORMAT V. 3.30, 13-JUL-11
REMARK 100
REMARK 100 THIS ENTRY HAS BEEN PROCESSED BY RCSB ON 03-SEP-14.
REMARK 100 THE DEPOSITION ID IS D_1000203471.
REMARK 200
REMARK 200 EXPERIMENTAL DETAILS
REMARK 200  EXPERIMENT TYPE                : X-RAY DIFFRACTION
REMARK 200  DATE OF DATA COLLECTION        : 25-APR-12
REMARK 200  TEMPERATURE           (KELVIN) : 100
REMARK 200  PH                             : 6.0
REMARK 200  NUMBER OF CRYSTALS USED        : NULL
REMARK 200
REMARK 200  SYNCHROTRON              (Y/N) : Y
REMARK 200  RADIATION SOURCE               : DIAMOND
REMARK 200  BEAMLINE                       : I04
REMARK 200  X-RAY GENERATOR MODEL          : NULL
REMARK 200  MONOCHROMATIC OR LAUE    (M/L) : M
REMARK 200  WAVELENGTH OR RANGE        (A) : 0.9795
REMARK 200  MONOCHROMATOR                  : NULL
REMARK 200  OPTICS                         : NULL
REMARK 200
REMARK 200  DETECTOR TYPE                  : CCD
REMARK 200  DETECTOR MANUFACTURER          : ADSC QUANTUM 270
REMARK 200  INTENSITY-INTEGRATION SOFTWARE : XDS
REMARK 200  DATA SCALING SOFTWARE          : SCALA
REMARK 200
REMARK 200  NUMBER OF UNIQUE REFLECTIONS   : 40214
REMARK 200  RESOLUTION RANGE HIGH      (A) : 2.200
REMARK 200  RESOLUTION RANGE LOW       (A) : 67.000
REMARK 200  REJECTION CRITERIA  (SIGMA(I)) : NULL
REMARK 200
REMARK 200 OVERALL.
REMARK 200  COMPLETENESS FOR RANGE     (%) : 99.7
REMARK 200  DATA REDUNDANCY                : 4.000
REMARK 200  R MERGE                    (I) : 0.08000
REMARK 200  R SYM                      (I) : NULL
REMARK 200   FOR THE DATA SET  : 10.1000
REMARK 200
REMARK 200 IN THE HIGHEST RESOLUTION SHELL.
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE HIGH (A) : 2.20
REMARK 200  HIGHEST RESOLUTION SHELL, RANGE LOW  (A) : 2.32
REMARK 200  COMPLETENESS FOR SHELL     (%) : 99.0
REMARK 200  DATA REDUNDANCY IN SHELL       : 4.10
REMARK 200  R MERGE FOR SHELL          (I) : 0.60800
REMARK 200  R SYM FOR SHELL            (I) : NULL
REMARK 200   FOR SHELL         : 1.900
REMARK 200
REMARK 200 DIFFRACTION PROTOCOL: SINGLE WAVELENGTH
REMARK 200 METHOD USED TO DETERMINE THE STRUCTURE: SAD
REMARK 200 SOFTWARE USED: NULL
REMARK 200 STARTING MODEL: NULL
REMARK 200
REMARK 200 REMARK: NULL
REMARK 280
REMARK 280 CRYSTAL
REMARK 280 SOLVENT CONTENT, VS   (%): 32.58
REMARK 280 MATTHEWS COEFFICIENT, VM (ANGSTROMS**3/DA): 1.82
REMARK 280
REMARK 280 CRYSTALLIZATION CONDITIONS: 30% PEG3350, 100MM BISTRIS 6.0, 5%
REMARK 280  GLYCEROL, 200MM AMMONIUM SULFATE, PH 6.0, VAPOR DIFFUSION,
REMARK 280  TEMPERATURE 293K
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY
REMARK 290 SYMMETRY OPERATORS FOR SPACE GROUP: P 21 21 2
REMARK 290
REMARK 290      SYMOP   SYMMETRY
REMARK 290     NNNMMM   OPERATOR
REMARK 290       1555   X,Y,Z
REMARK 290       2555   -X,-Y,Z
REMARK 290       3555   -X+1/2,Y+1/2,-Z
REMARK 290       4555   X+1/2,-Y+1/2,-Z
REMARK 290
REMARK 290     WHERE NNN -> OPERATOR NUMBER
REMARK 290           MMM -> TRANSLATION VECTOR
REMARK 290
REMARK 290 CRYSTALLOGRAPHIC SYMMETRY TRANSFORMATIONS
REMARK 290 THE FOLLOWING TRANSFORMATIONS OPERATE ON THE ATOM/HETATM
REMARK 290 RECORDS IN THIS ENTRY TO PRODUCE CRYSTALLOGRAPHICALLY
REMARK 290 RELATED MOLECULES.
REMARK 290   SMTRY1   1  1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   1  0.000000  1.000000  0.000000        0.00000
REMARK 290   SMTRY3   1  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   2 -1.000000  0.000000  0.000000        0.00000
REMARK 290   SMTRY2   2  0.000000 -1.000000  0.000000        0.00000
REMARK 290   SMTRY3   2  0.000000  0.000000  1.000000        0.00000
REMARK 290   SMTRY1   3 -1.000000  0.000000  0.000000       36.35100
REMARK 290   SMTRY2   3  0.000000  1.000000  0.000000       87.71650
REMARK 290   SMTRY3   3  0.000000  0.000000 -1.000000        0.00000
REMARK 290   SMTRY1   4  1.000000  0.000000  0.000000       36.35100
REMARK 290   SMTRY2   4  0.000000 -1.000000  0.000000       87.71650
REMARK 290   SMTRY3   4  0.000000  0.000000 -1.000000        0.00000
REMARK 290
REMARK 290 REMARK: NULL
REMARK 300
REMARK 300 BIOMOLECULE: 1
REMARK 300 SEE REMARK 350 FOR THE AUTHOR PROVIDED AND/OR PROGRAM
REMARK 300 GENERATED ASSEMBLY INFORMATION FOR THE STRUCTURE IN
REMARK 300 THIS ENTRY. THE REMARK MAY ALSO PROVIDE INFORMATION ON
REMARK 300 BURIED SURFACE AREA.
REMARK 350
REMARK 350 COORDINATES FOR A COMPLETE MULTIMER REPRESENTING THE KNOWN
REMARK 350 BIOLOGICALLY SIGNIFICANT OLIGOMERIZATION STATE OF THE
REMARK 350 MOLECULE CAN BE GENERATED BY APPLYING BIOMT TRANSFORMATIONS
REMARK 350 GIVEN BELOW.  BOTH NON-CRYSTALLOGRAPHIC AND
REMARK 350 CRYSTALLOGRAPHIC OPERATIONS ARE GIVEN.
REMARK 350
REMARK 350 BIOMOLECULE: 1
REMARK 350 AUTHOR DETERMINED BIOLOGICAL UNIT: DIMERIC
REMARK 350 SOFTWARE DETERMINED QUATERNARY STRUCTURE: DIMERIC
REMARK 350 SOFTWARE USED: PISA
REMARK 350 TOTAL BURIED SURFACE AREA: 1790 ANGSTROM**2
REMARK 350 SURFACE AREA OF THE COMPLEX: 30210 ANGSTROM**2
REMARK 350 CHANGE IN SOLVENT FREE ENERGY: -33.0 KCAL/MOL
REMARK 350 APPLY THE FOLLOWING TO CHAINS: A, B
REMARK 350   BIOMT1   1  1.000000  0.000000  0.000000        0.00000
REMARK 350   BIOMT2   1  0.000000  1.000000  0.000000        0.00000
REMARK 350   BIOMT3   1  0.000000  0.000000  1.000000        0.00000
REMARK 465
REMARK 465 MISSING RESIDUES
REMARK 465 THE FOLLOWING RESIDUES WERE NOT LOCATED IN THE
REMARK 465 EXPERIMENT. (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN
REMARK 465 IDENTIFIER; SSSEQ=SEQUENCE NUMBER; I=INSERTION CODE.)
REMARK 465
REMARK 465   M RES C SSSEQI
REMARK 465     GLU A    35
REMARK 465     THR A    36
REMARK 465     GLY A    37
REMARK 465     ARG A    38
REMARK 465     THR A    39
REMARK 465     GLU A    40
REMARK 465     ALA A    41
REMARK 465     ALA A    42
REMARK 465     PRO A    43
REMARK 465     ALA A    44
REMARK 465     ALA A    45
REMARK 465     GLY A    46
REMARK 465     GLN A    47
REMARK 465     PRO A    48
REMARK 465     VAL A    49
REMARK 465     GLU A    50
REMARK 465     SER A    51
REMARK 465     PHE A    52
REMARK 465     PRO A    53
REMARK 465     LEU A    54
REMARK 465     ASP A    55
REMARK 465     PHE A    56
REMARK 465     THR A    57
REMARK 465     ALA A    58
REMARK 465     VAL A    59
REMARK 465     GLU A    60
REMARK 465     GLY A    61
REMARK 465     ASN A    62
REMARK 465     MET A    63
REMARK 465     ASP A    64
REMARK 465     SER A    65
REMARK 465     PHE A    66
REMARK 465     MET A    67
REMARK 465     ALA A    68
REMARK 465     GLN A    69
REMARK 465     VAL A    70
REMARK 465     LYS A    71
REMARK 465     SER A    72
REMARK 465     LEU A    73
REMARK 465     ALA A    74
REMARK 465     GLN A    75
REMARK 465     SER A    76
REMARK 465     LEU A    77
REMARK 465     TYR A    78
REMARK 465     PRO A    79
REMARK 465     CYS A    80
REMARK 465     SER A    81
REMARK 465     ALA A    82
REMARK 465     GLN A    83
REMARK 465     GLN A    84
REMARK 465     LEU A    85
REMARK 465     GLU A   196
REMARK 465     LYS A   197
REMARK 465     ASN A   198
REMARK 465     THR A   352
REMARK 465     GLY A   353
REMARK 465     GLN A   354
REMARK 465     HIS A   419
REMARK 465     ASP A   420
REMARK 465     SER A   421
REMARK 465     HIS A   422
REMARK 465     LYS A   423
REMARK 465     ALA A   424
REMARK 465     SER A   425
REMARK 465     LYS A   426
REMARK 465     THR A   427
REMARK 465     PRO A   428
REMARK 465     LEU A   429
REMARK 465     LYS A   430
REMARK 465     GLY A   431
REMARK 465     VAL A   452
REMARK 465     ARG A   453
REMARK 465     ASP A   454
REMARK 465     GLN A   455
REMARK 465     PHE A   456
REMARK 465     THR A   457
REMARK 465     GLY A   458
REMARK 465     GLN A   459
REMARK 465     GLU A   460
REMARK 465     MET A   461
REMARK 465     ASN A   462
REMARK 465     VAL A   463
REMARK 465     ALA A   464
REMARK 465     GLN A   465
REMARK 465     PHE A   466
REMARK 465     LEU A   467
REMARK 465     MET A   468
REMARK 465     HIS A   469
REMARK 465     MET A   470
REMARK 465     GLY A   471
REMARK 465     PHE A   472
REMARK 465     ASP A   473
REMARK 465     MET A   474
REMARK 465     GLN A   475
REMARK 465     THR A   476
REMARK 465     VAL A   477
REMARK 465     ALA A   478
REMARK 465     GLN A   479
REMARK 465     PRO A   480
REMARK 465     GLN A   481
REMARK 465     GLY A   482
REMARK 465     LEU A   483
REMARK 465     GLU A   484
REMARK 465     PRO A   485
REMARK 465     SER A   486
REMARK 465     GLU A   487
REMARK 465     LEU A   488
REMARK 465     LEU A   489
REMARK 465     GLY A   490
REMARK 465     MET A   491
REMARK 465     LEU A   492
REMARK 465     SER A   493
REMARK 465     ASN A   494
REMARK 465     GLY A   495
REMARK 465     SER A   496
REMARK 465     GLY A   497
REMARK 465     THR A   498
REMARK 465     LYS A   499
REMARK 465     HIS A   500
REMARK 465     HIS A   501
REMARK 465     HIS A   502
REMARK 465     HIS A   503
REMARK 465     HIS A   504
REMARK 465     HIS A   505
REMARK 465     GLU B    35
REMARK 465     THR B    36
REMARK 465     GLY B    37
REMARK 465     ARG B    38
REMARK 465     THR B    39
REMARK 465     GLU B    40
REMARK 465     ALA B    41
REMARK 465     ALA B    42
REMARK 465     PRO B    43
REMARK 465     ALA B    44
REMARK 465     ALA B    45
REMARK 465     GLY B    46
REMARK 465     GLN B    47
REMARK 465     PRO B    48
REMARK 465     VAL B    49
REMARK 465     GLU B    50
REMARK 465     SER B    51
REMARK 465     PHE B    52
REMARK 465     PRO B    53
REMARK 465     LEU B    54
REMARK 465     ASP B    55
REMARK 465     PHE B    56
REMARK 465     THR B    57
REMARK 465     ALA B    58
REMARK 465     VAL B    59
REMARK 465     GLU B    60
REMARK 465     GLY B    61
REMARK 465     ASN B    62
REMARK 465     MET B    63
REMARK 465     ASP B    64
REMARK 465     SER B    65
REMARK 465     PHE B    66
REMARK 465     MET B    67
REMARK 465     ALA B    68
REMARK 465     GLN B    69
REMARK 465     VAL B    70
REMARK 465     LYS B    71
REMARK 465     SER B    72
REMARK 465     LEU B    73
REMARK 465     ALA B    74
REMARK 465     GLN B    75
REMARK 465     SER B    76
REMARK 465     LEU B    77
REMARK 465     TYR B    78
REMARK 465     PRO B    79
REMARK 465     CYS B    80
REMARK 465     SER B    81
REMARK 465     ALA B    82
REMARK 465     GLN B    83
REMARK 465     GLN B    84
REMARK 465     LEU B    85
REMARK 465     GLU B   196
REMARK 465     LYS B   197
REMARK 465     HIS B   419
REMARK 465     ASP B   420
REMARK 465     SER B   421
REMARK 465     HIS B   422
REMARK 465     LYS B   423
REMARK 465     ALA B   424
REMARK 465     SER B   425
REMARK 465     LYS B   426
REMARK 465     THR B   427
REMARK 465     PRO B   428
REMARK 465     LEU B   429
REMARK 465     LYS B   430
REMARK 465     GLY B   431
REMARK 465     VAL B   452
REMARK 465     ARG B   453
REMARK 465     ASP B   454
REMARK 465     GLN B   455
REMARK 465     PHE B   456
REMARK 465     THR B   457
REMARK 465     GLY B   458
REMARK 465     GLN B   459
REMARK 465     GLU B   460
REMARK 465     MET B   461
REMARK 465     ASN B   462
REMARK 465     VAL B   463
REMARK 465     ALA B   464
REMARK 465     GLN B   465
REMARK 465     PHE B   466
REMARK 465     LEU B   467
REMARK 465     MET B   468
REMARK 465     HIS B   469
REMARK 465     MET B   470
REMARK 465     GLY B   471
REMARK 465     PHE B   472
REMARK 465     ASP B   473
REMARK 465     MET B   474
REMARK 465     GLN B   475
REMARK 465     THR B   476
REMARK 465     VAL B   477
REMARK 465     ALA B   478
REMARK 465     GLN B   479
REMARK 465     PRO B   480
REMARK 465     GLN B   481
REMARK 465     GLY B   482
REMARK 465     LEU B   483
REMARK 465     GLU B   484
REMARK 465     PRO B   485
REMARK 465     SER B   486
REMARK 465     GLU B   487
REMARK 465     LEU B   488
REMARK 465     LEU B   489
REMARK 465     GLY B   490
REMARK 465     MET B   491
REMARK 465     LEU B   492
REMARK 465     SER B   493
REMARK 465     ASN B   494
REMARK 465     GLY B   495
REMARK 465     SER B   496
REMARK 465     GLY B   497
REMARK 465     THR B   498
REMARK 465     LYS B   499
REMARK 465     HIS B   500
REMARK 465     HIS B   501
REMARK 465     HIS B   502
REMARK 465     HIS B   503
REMARK 465     HIS B   504
REMARK 465     HIS B   505
REMARK 470
REMARK 470 MISSING ATOM
REMARK 470 THE FOLLOWING RESIDUES HAVE MISSING ATOMS (M=MODEL NUMBER;
REMARK 470 RES=RESIDUE NAME; C=CHAIN IDENTIFIER; SSEQ=SEQUENCE NUMBER;
REMARK 470 I=INSERTION CODE):
REMARK 470   M RES CSSEQI  ATOMS
REMARK 470     GLU A 199    CG   CD   OE1  OE2
REMARK 470     GLN A 357    CG   CD   OE1  NE2
REMARK 470     GLU B 134    CG   CD   OE1  OE2
REMARK 470     GLN B 354    CG   CD   OE1  NE2
REMARK 470     GLN B 357    CG   CD   OE1  NE2
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: CLOSE CONTACTS
REMARK 500
REMARK 500 THE FOLLOWING ATOMS THAT ARE RELATED BY CRYSTALLOGRAPHIC
REMARK 500 SYMMETRY ARE IN CLOSE CONTACT.  AN ATOM LOCATED WITHIN 0.15
REMARK 500 ANGSTROMS OF A SYMMETRY RELATED ATOM IS ASSUMED TO BE ON A
REMARK 500 SPECIAL POSITION AND IS, THEREFORE, LISTED IN REMARK 375
REMARK 500 INSTEAD OF REMARK 500.  ATOMS WITH NON-BLANK ALTERNATE
REMARK 500 LOCATION INDICATORS ARE NOT INCLUDED IN THE CALCULATIONS.
REMARK 500
REMARK 500 DISTANCE CUTOFF:
REMARK 500 2.2 ANGSTROMS FOR CONTACTS NOT INVOLVING HYDROGEN ATOMS
REMARK 500 1.6 ANGSTROMS FOR CONTACTS INVOLVING HYDROGEN ATOMS
REMARK 500
REMARK 500  ATM1  RES C  SSEQI   ATM2  RES C  SSEQI  SSYMOP   DISTANCE
REMARK 500   NH2  ARG B   292     NH2  ARG B   292     2555     1.50
REMARK 500
REMARK 500 REMARK: NULL
REMARK 500
REMARK 500 GEOMETRY AND STEREOCHEMISTRY
REMARK 500 SUBTOPIC: TORSION ANGLES
REMARK 500
REMARK 500 TORSION ANGLES OUTSIDE THE EXPECTED RAMACHANDRAN REGIONS:
REMARK 500 (M=MODEL NUMBER; RES=RESIDUE NAME; C=CHAIN IDENTIFIER;
REMARK 500 SSEQ=SEQUENCE NUMBER; I=INSERTION CODE).
REMARK 500
REMARK 500 STANDARD TABLE:
REMARK 500 FORMAT:(10X,I3,1X,A3,1X,A1,I4,A1,4X,F7.2,3X,F7.2)
REMARK 500
REMARK 500 EXPECTED VALUES: GJ KLEYWEGT AND TA JONES (1996). PHI/PSI-
REMARK 500 CHOLOGY: RAMACHANDRAN REVISITED. STRUCTURE 4, 1395 - 1400
REMARK 500
REMARK 500  M RES CSSEQI        PSI       PHI
REMARK 500    TRP A 128     -140.75     61.72
REMARK 500    TYR A 129      171.86    178.38
REMARK 500    SER A 232     -113.19     54.64
REMARK 500    TYR A 274      -70.49    -61.99
REMARK 500    ASN A 348       15.65     83.09
REMARK 500    GLU A 390      154.24     74.33
REMARK 500    ILE A 391      -37.00   -154.44
REMARK 500    VAL A 434      -34.85   -134.42
REMARK 500    SER A 439       58.07    -98.90
REMARK 500    TRP B 128     -137.75     70.50
REMARK 500    TYR B 129      175.81    175.51
REMARK 500    MET B 143       43.43   -146.42
REMARK 500    SER B 232     -124.61     62.28
REMARK 500    VAL B 280      -70.02   -109.37
REMARK 500    LEU B 351       56.87    -98.88
REMARK 500    GLN B 354     -161.51     59.88
REMARK 500    GLU B 390      159.56     67.94
REMARK 500    ILE B 391      -33.37   -159.90
REMARK 500    HIS B 444        8.97     85.39
REMARK 500
REMARK 500 REMARK: NULL
REMARK 800
REMARK 800 SITE
REMARK 800 SITE_IDENTIFIER: AC1
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CL A 601
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC2
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CL A 602
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC3
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CL A 603
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC4
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue CL B 602
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC5
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for residue SO4 B 603
REMARK 800
REMARK 800 SITE_IDENTIFIER: AC6
REMARK 800 EVIDENCE_CODE: SOFTWARE
REMARK 800 SITE_DESCRIPTION: binding site for Mono-Saccharide NAG B 601 bound
REMARK 800  to ASN B 96
DBREF  4WBH A   38   496  UNP    Q6P988   NOTUM_HUMAN     38    496
DBREF  4WBH B   38   496  UNP    Q6P988   NOTUM_HUMAN     38    496
SEQADV 4WBH GLU A   35  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH THR A   36  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH GLY A   37  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH GLY A  497  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH THR A  498  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH LYS A  499  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS A  500  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS A  501  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS A  502  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS A  503  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS A  504  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS A  505  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH GLU B   35  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH THR B   36  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH GLY B   37  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH GLY B  497  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH THR B  498  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH LYS B  499  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS B  500  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS B  501  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS B  502  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS B  503  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS B  504  UNP  Q6P988              EXPRESSION TAG
SEQADV 4WBH HIS B  505  UNP  Q6P988              EXPRESSION TAG
SEQRES   1 A  471  GLU THR GLY ARG THR GLU ALA ALA PRO ALA ALA GLY GLN
SEQRES   2 A  471  PRO VAL GLU SER PHE PRO LEU ASP PHE THR ALA VAL GLU
SEQRES   3 A  471  GLY ASN MET ASP SER PHE MET ALA GLN VAL LYS SER LEU
SEQRES   4 A  471  ALA GLN SER LEU TYR PRO CYS SER ALA GLN GLN LEU ASN
SEQRES   5 A  471  GLU ASP LEU ARG LEU HIS LEU LEU LEU ASN THR SER VAL
SEQRES   6 A  471  THR CYS ASN ASP GLY SER PRO ALA GLY TYR TYR LEU LYS
SEQRES   7 A  471  GLU SER ARG GLY SER ARG ARG TRP LEU LEU PHE LEU GLU
SEQRES   8 A  471  GLY GLY TRP TYR CYS PHE ASN ARG GLU ASN CYS ASP SER
SEQRES   9 A  471  ARG TYR ASP THR MET ARG ARG LEU MET SER SER ARG ASP
SEQRES  10 A  471  TRP PRO ARG THR ARG THR GLY THR GLY ILE LEU SER SER
SEQRES  11 A  471  GLN PRO GLU GLU ASN PRO TYR TRP TRP ASN ALA ASN MET
SEQRES  12 A  471  VAL PHE ILE PRO TYR CYS SER SER ASP VAL TRP SER GLY
SEQRES  13 A  471  ALA SER SER LYS SER GLU LYS ASN GLU TYR ALA PHE MET
SEQRES  14 A  471  GLY ALA LEU ILE ILE GLN GLU VAL VAL ARG GLU LEU LEU
SEQRES  15 A  471  GLY ARG GLY LEU SER GLY ALA LYS VAL LEU LEU LEU ALA
SEQRES  16 A  471  GLY SER SER ALA GLY GLY THR GLY VAL LEU LEU ASN VAL
SEQRES  17 A  471  ASP ARG VAL ALA GLU GLN LEU GLU LYS LEU GLY TYR PRO
SEQRES  18 A  471  ALA ILE GLN VAL ARG GLY LEU ALA ASP SER GLY TRP PHE
SEQRES  19 A  471  LEU ASP ASN LYS GLN TYR ARG HIS THR ASP CYS VAL ASP
SEQRES  20 A  471  THR ILE THR CYS ALA PRO THR GLU ALA ILE ARG ARG GLY
SEQRES  21 A  471  ILE ARG TYR TRP ASN GLY VAL VAL PRO GLU ARG CYS ARG
SEQRES  22 A  471  ARG GLN PHE GLN GLU GLY GLU GLU TRP ASN CYS PHE PHE
SEQRES  23 A  471  GLY TYR LYS VAL TYR PRO THR LEU ARG CYS PRO VAL PHE
SEQRES  24 A  471  VAL VAL GLN TRP LEU PHE ASP GLU ALA GLN LEU THR VAL
SEQRES  25 A  471  ASP ASN VAL HIS LEU THR GLY GLN PRO VAL GLN GLU GLY
SEQRES  26 A  471  LEU ARG LEU TYR ILE GLN ASN LEU GLY ARG GLU LEU ARG
SEQRES  27 A  471  HIS THR LEU LYS ASP VAL PRO ALA SER PHE ALA PRO ALA
SEQRES  28 A  471  CYS LEU SER HIS GLU ILE ILE ILE ARG SER HIS TRP THR
SEQRES  29 A  471  ASP VAL GLN VAL LYS GLY THR SER LEU PRO ARG ALA LEU
SEQRES  30 A  471  HIS CYS TRP ASP ARG SER LEU HIS ASP SER HIS LYS ALA
SEQRES  31 A  471  SER LYS THR PRO LEU LYS GLY CYS PRO VAL HIS LEU VAL
SEQRES  32 A  471  ASP SER CYS PRO TRP PRO HIS CYS ASN PRO SER CYS PRO
SEQRES  33 A  471  THR VAL ARG ASP GLN PHE THR GLY GLN GLU MET ASN VAL
SEQRES  34 A  471  ALA GLN PHE LEU MET HIS MET GLY PHE ASP MET GLN THR
SEQRES  35 A  471  VAL ALA GLN PRO GLN GLY LEU GLU PRO SER GLU LEU LEU
SEQRES  36 A  471  GLY MET LEU SER ASN GLY SER GLY THR LYS HIS HIS HIS
SEQRES  37 A  471  HIS HIS HIS
SEQRES   1 B  471  GLU THR GLY ARG THR GLU ALA ALA PRO ALA ALA GLY GLN
SEQRES   2 B  471  PRO VAL GLU SER PHE PRO LEU ASP PHE THR ALA VAL GLU
SEQRES   3 B  471  GLY ASN MET ASP SER PHE MET ALA GLN VAL LYS SER LEU
SEQRES   4 B  471  ALA GLN SER LEU TYR PRO CYS SER ALA GLN GLN LEU ASN
SEQRES   5 B  471  GLU ASP LEU ARG LEU HIS LEU LEU LEU ASN THR SER VAL
SEQRES   6 B  471  THR CYS ASN ASP GLY SER PRO ALA GLY TYR TYR LEU LYS
SEQRES   7 B  471  GLU SER ARG GLY SER ARG ARG TRP LEU LEU PHE LEU GLU
SEQRES   8 B  471  GLY GLY TRP TYR CYS PHE ASN ARG GLU ASN CYS ASP SER
SEQRES   9 B  471  ARG TYR ASP THR MET ARG ARG LEU MET SER SER ARG ASP
SEQRES  10 B  471  TRP PRO ARG THR ARG THR GLY THR GLY ILE LEU SER SER
SEQRES  11 B  471  GLN PRO GLU GLU ASN PRO TYR TRP TRP ASN ALA ASN MET
SEQRES  12 B  471  VAL PHE ILE PRO TYR CYS SER SER ASP VAL TRP SER GLY
SEQRES  13 B  471  ALA SER SER LYS SER GLU LYS ASN GLU TYR ALA PHE MET
SEQRES  14 B  471  GLY ALA LEU ILE ILE GLN GLU VAL VAL ARG GLU LEU LEU
SEQRES  15 B  471  GLY ARG GLY LEU SER GLY ALA LYS VAL LEU LEU LEU ALA
SEQRES  16 B  471  GLY SER SER ALA GLY GLY THR GLY VAL LEU LEU ASN VAL
SEQRES  17 B  471  ASP ARG VAL ALA GLU GLN LEU GLU LYS LEU GLY TYR PRO
SEQRES  18 B  471  ALA ILE GLN VAL ARG GLY LEU ALA ASP SER GLY TRP PHE
SEQRES  19 B  471  LEU ASP ASN LYS GLN TYR ARG HIS THR ASP CYS VAL ASP
SEQRES  20 B  471  THR ILE THR CYS ALA PRO THR GLU ALA ILE ARG ARG GLY
SEQRES  21 B  471  ILE ARG TYR TRP ASN GLY VAL VAL PRO GLU ARG CYS ARG
SEQRES  22 B  471  ARG GLN PHE GLN GLU GLY GLU GLU TRP ASN CYS PHE PHE
SEQRES  23 B  471  GLY TYR LYS VAL TYR PRO THR LEU ARG CYS PRO VAL PHE
SEQRES  24 B  471  VAL VAL GLN TRP LEU PHE ASP GLU ALA GLN LEU THR VAL
SEQRES  25 B  471  ASP ASN VAL HIS LEU THR GLY GLN PRO VAL GLN GLU GLY
SEQRES  26 B  471  LEU ARG LEU TYR ILE GLN ASN LEU GLY ARG GLU LEU ARG
SEQRES  27 B  471  HIS THR LEU LYS ASP VAL PRO ALA SER PHE ALA PRO ALA
SEQRES  28 B  471  CYS LEU SER HIS GLU ILE ILE ILE ARG SER HIS TRP THR
SEQRES  29 B  471  ASP VAL GLN VAL LYS GLY THR SER LEU PRO ARG ALA LEU
SEQRES  30 B  471  HIS CYS TRP ASP ARG SER LEU HIS ASP SER HIS LYS ALA
SEQRES  31 B  471  SER LYS THR PRO LEU LYS GLY CYS PRO VAL HIS LEU VAL
SEQRES  32 B  471  ASP SER CYS PRO TRP PRO HIS CYS ASN PRO SER CYS PRO
SEQRES  33 B  471  THR VAL ARG ASP GLN PHE THR GLY GLN GLU MET ASN VAL
SEQRES  34 B  471  ALA GLN PHE LEU MET HIS MET GLY PHE ASP MET GLN THR
SEQRES  35 B  471  VAL ALA GLN PRO GLN GLY LEU GLU PRO SER GLU LEU LEU
SEQRES  36 B  471  GLY MET LEU SER ASN GLY SER GLY THR LYS HIS HIS HIS
SEQRES  37 B  471  HIS HIS HIS
HET     CL  A 601       1
HET     CL  A 602       1
HET     CL  A 603       1
HET    NAG  B 601      14
HET     CL  B 602       1
HET    SO4  B 603       5
HETNAM      CL CHLORIDE ION
HETNAM     NAG N-ACETYL-D-GLUCOSAMINE
HETNAM     SO4 SULFATE ION
FORMUL   3   CL    4(CL 1-)
FORMUL   6  NAG    C8 H15 N O6
FORMUL   8  SO4    O4 S 2-
FORMUL   9  HOH   *59(H2 O)
HELIX    1 AA1 ASN A  132  MET A  143  1                                  12
HELIX    2 AA2 ARG A  144  SER A  148  5                                   5
HELIX    3 AA3 THR A  159  SER A  163  5                                   5
HELIX    4 AA4 MET A  203  GLY A  217  1                                  15
HELIX    5 AA5 ARG A  218  ALA A  223  5                                   6
HELIX    6 AA6 SER A  232  LEU A  252  1                                  21
HELIX    7 AA7 ALA A  286  ASN A  299  1                                  14
HELIX    8 AA8 PRO A  303  ARG A  308  1                                   6
HELIX    9 AA9 GLU A  314  PHE A  319  5                                   6
HELIX   10 AB1 PHE A  320  TYR A  325  1                                   6
HELIX   11 AB2 PRO A  326  LEU A  328  5                                   3
HELIX   12 AB3 GLU A  341  ASP A  347  1                                   7
HELIX   13 AB4 GLN A  357  LEU A  375  1                                  19
HELIX   14 AB5 ARG A  394  ASP A  399  5                                   6
HELIX   15 AB6 LEU A  407  LEU A  418  1                                  12
HELIX   16 AB7 ASN B  132  MET B  143  1                                  12
HELIX   17 AB8 ARG B  144  MET B  147  5                                   4
HELIX   18 AB9 THR B  159  SER B  163  5                                   5
HELIX   19 AC1 MET B  203  GLY B  217  1                                  15
HELIX   20 AC2 ARG B  218  ALA B  223  5                                   6
HELIX   21 AC3 SER B  232  LEU B  252  1                                  21
HELIX   22 AC4 ALA B  286  ASN B  299  1                                  14
HELIX   23 AC5 PRO B  303  ARG B  308  1                                   6
HELIX   24 AC6 GLU B  314  PHE B  319  5                                   6
HELIX   25 AC7 PHE B  320  TYR B  325  1                                   6
HELIX   26 AC8 PRO B  326  LEU B  328  5                                   3
HELIX   27 AC9 GLU B  341  ASP B  347  1                                   7
HELIX   28 AD1 GLN B  357  LEU B  375  1                                  19
HELIX   29 AD2 ARG B  394  ASP B  399  5                                   6
HELIX   30 AD3 LEU B  407  LEU B  418  1                                  12
SHEET    1 AA110 THR A 155  ARG A 156  0
SHEET    2 AA110 LEU A  89  LEU A  93 -1  N  LEU A  89   O  ARG A 156
SHEET    3 AA110 GLY A 108  LYS A 112 -1  O  TYR A 109   N  HIS A  92
SHEET    4 AA110 ASN A 176  ILE A 180 -1  O  PHE A 179   N  TYR A 110
SHEET    5 AA110 ARG A 119  LEU A 124  1  N  PHE A 123   O  ILE A 180
SHEET    6 AA110 VAL A 225  SER A 231  1  O  LEU A 227   N  TRP A 120
SHEET    7 AA110 GLN A 258  ASP A 264  1  O  GLN A 258   N  LEU A 226
SHEET    8 AA110 VAL A 332  VAL A 335  1  O  VAL A 335   N  ALA A 263
SHEET    9 AA110 SER A 381  ALA A 383  1  O  PHE A 382   N  VAL A 334
SHEET   10 AA110 HIS A 435  VAL A 437  1  O  LEU A 436   N  SER A 381
SHEET    1 AA2 2 PHE A 339  ASP A 340  0
SHEET    2 AA2 2 LEU A 387  SER A 388  1  O  SER A 388   N  PHE A 339
SHEET    1 AA3 2 GLN A 401  VAL A 402  0
SHEET    2 AA3 2 THR A 405  SER A 406 -1  O  THR A 405   N  VAL A 402
SHEET    1 AA410 THR B 155  ARG B 156  0
SHEET    2 AA410 LEU B  89  LEU B  93 -1  N  LEU B  89   O  ARG B 156
SHEET    3 AA410 GLY B 108  LYS B 112 -1  O  TYR B 109   N  HIS B  92
SHEET    4 AA410 ASN B 176  ILE B 180 -1  O  PHE B 179   N  TYR B 110
SHEET    5 AA410 ARG B 119  LEU B 124  1  N  PHE B 123   O  VAL B 178
SHEET    6 AA410 VAL B 225  SER B 231  1  O  LEU B 227   N  TRP B 120
SHEET    7 AA410 GLN B 258  ASP B 264  1  O  ARG B 260   N  LEU B 228
SHEET    8 AA410 VAL B 332  VAL B 335  1  O  PHE B 333   N  GLY B 261
SHEET    9 AA410 SER B 381  ALA B 383  1  O  PHE B 382   N  VAL B 334
SHEET   10 AA410 HIS B 435  VAL B 437  1  O  LEU B 436   N  ALA B 383
SHEET    1 AA5 2 PHE B 339  ASP B 340  0
SHEET    2 AA5 2 LEU B 387  SER B 388  1  O  SER B 388   N  PHE B 339
SHEET    1 AA6 2 GLN B 401  VAL B 402  0
SHEET    2 AA6 2 THR B 405  SER B 406 -1  O  THR B 405   N  VAL B 402
SSBOND   1 CYS A  101    CYS A  183                          1555   1555  2.04
SSBOND   2 CYS A  130    CYS A  136                          1555   1555  2.03
SSBOND   3 CYS A  279    CYS A  285                          1555   1555  2.04
SSBOND   4 CYS A  306    CYS A  318                          1555   1555  2.06
SSBOND   5 CYS A  386    CYS A  449                          1555   1555  2.03
SSBOND   6 CYS A  413    CYS A  432                          1555   1555  2.01
SSBOND   7 CYS A  440    CYS A  445                          1555   1555  2.03
SSBOND   8 CYS B  101    CYS B  183                          1555   1555  2.04
SSBOND   9 CYS B  130    CYS B  136                          1555   1555  2.03
SSBOND  10 CYS B  279    CYS B  285                          1555   1555  2.05
SSBOND  11 CYS B  306    CYS B  318                          1555   1555  2.07
SSBOND  12 CYS B  386    CYS B  449                          1555   1555  2.03
SSBOND  13 CYS B  413    CYS B  432                          1555   1555  2.04
SSBOND  14 CYS B  440    CYS B  445                          1555   1555  2.03
LINK         ND2 ASN B  96                 C1  NAG B 601     1555   1555  1.46
SITE     1 AC1  1 ARG A 409
SITE     1 AC2  1 HOH A 722
SITE     1 AC3  3 ARG A  90  HIS A  92  ARG A 218
SITE     1 AC4  1 ARG B 218
SITE     1 AC5  3 SER B 117  ARG B 119  ASN B 174
SITE     1 AC6  3 ASN B  96  GLU B 210  ARG B 213
CRYST1   72.702  175.433   60.764  90.00  90.00  90.00 P 21 21 2     8
ORIGX1      1.000000  0.000000  0.000000        0.00000
ORIGX2      0.000000  1.000000  0.000000        0.00000
ORIGX3      0.000000  0.000000  1.000000        0.00000
SCALE1      0.013755  0.000000  0.000000        0.00000
SCALE2      0.000000  0.005700  0.000000        0.00000
SCALE3      0.000000  0.000000  0.016457        0.00000
TER    2784      THR A 451
TER    5593      THR B 451
MASTER      618    0    6   30   28    0    6    6 5656    2   50   74
END